Braulioceras
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Braulioceras
Braulioceras is a smooth, orthoconic orthocerid with very close spaced septa (around 8 over a length equal to the diameter of the shell) added to the Sactorthoceratidae in 2007. The type species, ''Braulioceras sanjuanense'' comes from the Middle Ordovician of the San Juan Formation, Argentine Precordillera. ''Braulioceras'' is very similar to ''Sactorthoceras''. The siphuncle The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ... is central, with orthochoanitic septal necks and segments which are slightly expanded between septa, giving it a beaded appearance. The thickness of the connecting rings is similar to that in ellesmerocerids. Endosiphuncular or cameral deposits are unknown. ''Braulioceras'' is distinguished from ''Sactorthoceras'' by its more closely central siphuncle, ...
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Sactorthoceratidae
The Sactorthoceratidae comprise Orthocerataceaen genera with a subcentral suborthochoanitic siphuncle composed of slightly expanded segments and free of organic deposits. The camerae (chambers) of the phragmocone (chambered portion) likewise have organic deposits that are typically retarded or sparse. Discussion The family, Sactorthoceratidae, was established by Rousseau Flower in 1946, based on the genus '' Sactorthoceras''. In 1962 Flower pointed out the problem of defining the Sactorthoceratidae based on ''Sactorthoceras''; there being three species groups within the genus: those with (1) short camerae and rather short tubular septal necks, (2) long camerae and slightly expanded siphuncle segments, (3) typically tubular siphuncle segments that are rapidly contracted at the septal foremina. In defining the Sactorthceratidae, the second group with slightly expanded segments was considered. However the type species ''S. goniaseptum'' belongs to the third group, with contrac ...
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Orthocone
An orthocone is an unusually long straight shell of a nautiloid cephalopod.; During the 18th and 19th centuries, all shells of this type were named ''Orthoceras'', creating a wastebasket taxon, but it is now known that many groups of nautiloids developed or retained this type of shell. An orthocone can be thought of as like a ''nautilus'', but with the shell straight and uncoiled. It was previously believed that these represented the most primitive form of nautiloid, but it is now known that the earliest nautiloids had shells that were slightly curved. An orthoconic form evolved several times among cephalopods, and among nautiloid cephalopods is prevalent among the ellesmerocerids, endocerids, actinocerids, orthoceratoids, and bactritids. Orthocones existed from the Late Cambrian to the Late Triassic, but they were most common in the early Paleozoic. Revivals of the orthocone design later occurred in other cephalopod groups, notably baculitid ammonites in the Cretaceous Peri ...
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Orthocerida
Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician () possibly to the Late Triassic (). A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous (). They were most common however from the Ordovician to the Devonian. Shell form The shell is usually long, and may be straight ("orthoconic") or gently curved. In life, these animals may have been similar to the modern squid, except for the long shell. The internal structure of the shell consists of concavo-convex chambers linked by a centrally-placed tube called a siphuncle. There is a tendency for the chambers to develop cameral deposits, which were used as ballast to balance the long gas-filled shell. Depending on the family, the siphuncle has orthochoanitic (short and straight) or cyrtochoanitic (outwardly curved) septal necks, which protrude from the septa. The shell surface may be (depending on the specie ...
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Septum
In biology, a septum (Latin for ''something that encloses''; plural septa) is a wall, dividing a cavity or structure into smaller ones. A cavity or structure divided in this way may be referred to as septate. Examples Human anatomy * Interatrial septum, the wall of tissue that is a sectional part of the left and right atria of the heart * Interventricular septum, the wall separating the left and right ventricles of the heart * Lingual septum, a vertical layer of fibrous tissue that separates the halves of the tongue. *Nasal septum: the cartilage wall separating the nostrils of the nose * Alveolar septum: the thin wall which separates the alveoli from each other in the lungs * Orbital septum, a palpebral ligament in the upper and lower eyelids * Septum pellucidum or septum lucidum, a thin structure separating two fluid pockets in the brain * Uterine septum, a malformation of the uterus * Vaginal septum, a lateral or transverse partition inside the vagina * Intermuscular sep ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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San Juan Formation, Argentina
The San Juan Formation ( es, Formación San Juan) is a geologic formation in Argentina. The formation comprising limestones, mudstones and marls was deposited in a shallow marine reefal environment and preserves many fossils dating back to the Ordovician period. The formation overlies the La Silla Formation and crops out in the Precordillera of San Juan Province. See also * Geological history of the Precordillera terrane * Mesón Group Mesón Group ( es, Grupo Mesón) is a Cambrian to Early Ordovician sedimentary formation located in the Argentine Northwest and nearby parts of Bolivia. The group members rest unconformably on top of the Ediacaran–Cambrian Puncoviscana Format ... References Further reading * J. L. Benedetto. 2012. ''Gatosella'', a new basal plectambonitoid brachiopod with undercut cardinal process from Middle Ordovician limestones of the Precordillera terrane, Argentina. Journal of Systematic Palaeontology 10(3):435-443 * M. G. Carrera. 2006. The n ...
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Precordillera
Precordillera is a Spanish geographical term for hills and mountains lying before a greater range, foothills. The term is derived from ''cordillera'' (mountain range)—literally "pre-mountain range"—and applied usually to the Andes. Some places usually called precordillera are: *Andean mountains east of the main ranges of Andes in Argentina. It is separated from the much higher Frontal Cordillera to the west by Uspallata Valley in Argentina. Precordillera mountains reach around 3,000 m a.s.l. in Sierras de Villavicencio. *Used all over Chile from north to south as a morphological unit lying just between the Andes and the Intermediate Depression The Central Valley ( es, Valle Central), Intermediate Depression, or Longitudinal Valley is the depression between the Chilean Coastal Range and the Andes Mountains. The Chilean Central Valley extends from the border with Peru to Puerto Montt in .... See also * Geological history of the precordillera terrane * References {{Geo-te ...
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Sactorthoceras
''Sactorthceras'' is an orthoceratoid genus (subclass Nautiloidea s.l.) known from the Middle Ordovician of eastern North America (NY), Norway and Korea and is the type genus of the Sactorthoceratidae. ''Sactorthceras'' was named by Kobayashi in 1934 to include smooth or striated, straight or slightly curved longicones with short camerae and slender, subcentral, suborthochoanitic siphuncles with segments slightly inflated and no cameral deposits. ''Sactorthocras'' has also been recognized in the Silurian Racine Formation of Wisconsin, and from the Ordovician of China and Iran. Related genera include ''Centroonoceras ''Centroonoceras'' is a middle Ordovician cyrtoconic nautiloid cephalopod, otherwise similar to the orthoconic '' Sactorthoceras'' and also included in the Sactorthoceratidae. It was named by Kobayashi, 1934, and has been found in Korea and in ...'' and '' Sigmocycloceras'' -References- *Sweet, Walter C. 1964. Nautiloidea -Orthocerida. Treatise on Inverteb ...
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Siphuncle
The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ''Spirula''. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name. Function The siphuncle is used primarily in emptying water from new chambers as the shell grows. To perform this task, the cephalopod increases the saltiness of the blood in the siphuncle, and the water moves from the more dilute chamber into the blood through osmosis. At the sam ...
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Ellesmerocerida
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician. Morphology The Ellesmerocerida are characterized by shells that are typically small, some even tiny, with close-spaced septa and relatively large ventral siphuncles. In some genera (e.g. ''Paleoceras''), the septa are uniformly spaced. Shells of ellesmerocerids are typically smooth and compressed and vary in form. They may be breviconic (short) or longiconic (elongate), straight (orthoconic) or curved (cyrtoconic). Cyrtoconic forms are usually endogastric, with longitudinally convex ventral margins. The apeces of straight forms typically have an endogastric curvature. Some may have grown to as much as 15 cm. Siphuncle segments are tubular or concave. Septal necks are short. Connecting rings which may appear layered are thick and typically wedge shaped with their maximum width at or near where t ...
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Camera (cephalopod)
Camerae (singular camera) are the spaces or chambers enclosed between two adjacent septa in the phragmocone of a nautiloid or ammonoid cephalopod molluscus. These can be seen in cross-sections of a nautilus shell and in the polished cross-sections of ammonites. In life these chambers are filled with gas, mediated by the siphuncle, and used to control buoyancy. Some Palaeozoic nautiloid genera, especially those with long, straight shells, are distinguished by cameral deposits. These were accumulations of calcium carbonate secreted in the empty chambers of the shell, used for ballast Ballast is material that is used to provide stability to a vehicle or structure. Ballast, other than cargo, may be placed in a vehicle, often a ship or the gondola of a balloon or airship, to provide stability. A compartment within a boat, ship, ... and control of buoyancy. The nature and form of these deposits are very useful in nautiloid classification. References * {{DEFAULTSORT:Camera (A ...
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Nautiloids
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rela ...
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