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Boveri–Sutton Chromosome Theory
The Boveri–Sutton chromosome theory (also known as the chromosome theory of inheritance or the Sutton–Boveri theory) is a fundamental unifying theory of genetics which identifies chromosomes as the carriers of genetic material.1902: Theodor Boveri (1862-1915) and Walter Sutton (1877-1916) propose that chromosomes bear hereditary factors in accordance with Mendelian laws
Genetics and Genomics Timeline. Genome News Network an online publication of the J. Craig Venter Institute.

Holmgren Lab Northwestern Uni ...
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Theodor Boveri Walter Sutton
Theodor is a masculine given name. It is a German form of Theodore. It is also a variant of Teodor. List of people with the given name Theodor * Theodor Adorno, (1903–1969), German philosopher * Theodor Aman, Romanian painter * Theodor Blueger, Latvian professional ice hockey forward for the Pittsburgh Penguins of the National Hockey League (NHL) * Theodor Burghele, Romanian surgeon, President of the Romanian Academy * Theodor Busse, German general during World War I and World War II * Theodor Cazaban, Romanian writer * Theodor Fischer (fencer), German Olympic épée and foil fencer * Theodor Fontane, (1819–1898), German writer * Theodor Geisel, American writer and cartoonist, known by the pseudonym Dr. Seuss * Theodor W. Hänsch (born 1940), German physicist * Theodor Herzl, (1860–1904), Austrian-Hungary Jewish journalist and the founder of modern political Zionism * Theodor Heuss, (1884–1963), German politician and publicist * Theodor Innitzer, Austrian Catholic car ...
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Embryonic Development
An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm cell. The resulting fusion of these two cells produces a single-celled zygote that undergoes many cell divisions that produce cells known as blastomeres. The blastomeres are arranged as a solid ball that when reaching a certain size, called a morula, takes in fluid to create a cavity called a blastocoel. The structure is then termed a blastula, or a blastocyst in mammals. The mammalian blastocyst hatches before implantating into the endometrial lining of the womb. Once implanted the embryo will continue its development through the next stages of gastrulation, neurulation, and organogenesis. Gastrulation is the formation of the three germ layers that will form all of the different parts of the body. Neurulation forms the nervous syst ...
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Biology Theories
Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary information encoded in genes, which can be transmitted to future generations. Another major theme is evolution, which explains the unity and diversity of life. Energy processing is also important to life as it allows organisms to move, grow, and reproduce. Finally, all organisms are able to regulate their own internal environments. Biologists are able to study life at multiple levels of organization, from the molecular biology of a cell to the anatomy and physiology of plants and animals, and evolution of populations.Based on definition from: Hence, there are multiple subdisciplines within biology, each defined by the nature of their research questions and the tools that they use. Like other scientists, biologists use the scientific meth ...
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Drosophila Melanogaster
''Drosophila melanogaster'' is a species of fly (the taxonomic order Diptera) in the family Drosophilidae. The species is often referred to as the fruit fly or lesser fruit fly, or less commonly the "vinegar fly" or "pomace fly". Starting with Charles W. Woodworth's 1901 proposal of the use of this species as a model organism, ''D. melanogaster'' continues to be widely used for biological research in genetics, physiology, microbial pathogenesis, and life history evolution. As of 2017, five Nobel Prizes have been awarded to drosophilists for their work using the insect. ''D. melanogaster'' is typically used in research owing to its rapid life cycle, relatively simple genetics with only four pairs of chromosomes, and large number of offspring per generation. It was originally an African species, with all non-African lineages having a common origin. Its geographic range includes all continents, including islands. ''D. melanogaster'' is a common pest in homes, restaurants, and othe ...
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Genetic Linkage
Genetic linkage is the tendency of DNA sequences that are close together on a chromosome to be inherited together during the meiosis phase of sexual reproduction. Two genetic markers that are physically near to each other are unlikely to be separated onto different chromatids during chromosomal crossover, and are therefore said to be more ''linked'' than markers that are far apart. In other words, the nearer two genes are on a chromosome, the lower the chance of recombination between them, and the more likely they are to be inherited together. Markers on different chromosomes are perfectly ''unlinked'', although the penetrance of potentially deleterious alleles may be influenced by the presence of other alleles, and these other alleles may be located on other chromosomes than that on which a particular potentially deleterious allele is located. Genetic linkage is the most prominent exception to Gregor Mendel's Law of Independent Assortment. The first experiment to demonstrate li ...
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Thomas Hunt Morgan
Thomas Hunt Morgan (September 25, 1866 – December 4, 1945) was an American evolutionary biologist, geneticist, embryologist, and science author who won the Nobel Prize in Physiology or Medicine in 1933 for discoveries elucidating the role that the chromosome plays in heredity. Morgan received his Ph.D. from Johns Hopkins University in zoology in 1890 and researched embryology during his tenure at Bryn Mawr. Following the rediscovery of Mendelian inheritance in 1900, Morgan began to study the genetic characteristics of the fruit fly ''Drosophila melanogaster''. In his famous Fly Room at Columbia University's Schermerhorn Hall, Morgan demonstrated that genes are carried on chromosomes and are the mechanical basis of heredity. These discoveries formed the basis of the modern science of genetics. During his distinguished career, Morgan wrote 22 books and 370 scientific papers. As a result of his work, ''Drosophila'' became a major model organism in contemporary genetics. The ...
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Independent Assortment
Mendelian inheritance (also known as Mendelism) is a type of biological inheritance following the principles originally proposed by Gregor Mendel in 1865 and 1866, re-discovered in 1900 by Hugo de Vries and Carl Correns, and later popularized by William Bateson. These principles were initially controversial. When Mendel's theories were integrated with the Boveri–Sutton chromosome theory of inheritance by Thomas Hunt Morgan in 1915, they became the core of classical genetics. Ronald Fisher combined these ideas with the theory of natural selection in his 1930 book ''The Genetical Theory of Natural Selection'', putting evolution onto a mathematical footing and forming the basis for population genetics within the modern evolutionary synthesis. History The principles of Mendelian inheritance were named for and first derived by Gregor Johann Mendel, a nineteenth-century Moravian monk who formulated his ideas after conducting simple hybridisation experiments with pea plants (' ...
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Eleanor Carothers
Estrella Eleanor Carothers (4 December 1882 – 1957), known primarily as Eleanor Carothers, was an American zoologist, geneticist, and cytologist known for her work with grasshoppers. She discovered important physical evidence for the concept of independent assortment, vital to modern understanding of genetics. Early life and education Carothers was born on December 4, 1883, in Newton, Kansas to Z. W. Carothers and Mary Bates. She studied at the Nickerson Normal College and went on to earn her Bachelor of Arts degree and her Master's degree from the University of Kansas, in 1911 and 1912 respectively. Research career She began her career as a Pepper Fellow at Pennsylvania State University, where she stayed from 1913 to 1914. That year, she was appointed an assistant professor of zoology there, and graduated with her Ph.D. in 1916. Carothers stayed on as assistant professor until 1936, though she also had an overlapping appointment as an independent investigator for the Mari ...
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Edmund Beecher Wilson
Edmund Beecher Wilson (October 19, 1856 – March 3, 1939) was a pioneering American zoologist and geneticist. He wrote one of the most influential textbooks in modern biology, ''The Cell''. Career Wilson was born in Geneva, Illinois, the son of Isaac G. Wilson, a judge, and his wife, Carioline Clark. He graduated from Yale University in 1878. He earned his Ph.D. at Johns Hopkins in 1881. He was a lecturer at Williams College in 1883–84 and at the Massachusetts Institute of Technology in 1884–85. He served as professor of biology at Bryn Mawr College from 1885 to 1891. In 1888, he was elected as a member to the American Philosophical Society. He spent the balance of his career at Columbia University where he was successively adjunct professor of biology (1891–94), professor of invertebrate zoology (1894–1897), and professor of zoology (from 1897). Wilson is credited as America's first cell biologist. In 1898 he used the similarity in embryos to describe phylogene ...
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Grasshopper
Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshoppers are typically ground-dwelling insects with powerful hind legs which allow them to escape from threats by leaping vigorously. As hemimetabolous insects, they do not undergo complete metamorphosis; they hatch from an egg into a nymph or "hopper" which undergoes five moults, becoming more similar to the adult insect at each developmental stage. The grasshopper hears through the tympanal organ which can be found in the first segment of the abdomen attached to the thorax; while its sense of vision is in the compound eyes, the change in light intensity is perceived in the simple eyes (ocelli). At high population densities and under certain environmental conditions, some grasshopper species can change color and behavior and form swarms. Under ...
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Sea Urchin
Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of sea urchins are round and spiny, ranging in diameter from . Sea urchins move slowly, crawling with tube feet, and also propel themselves with their spines. Although algae are the primary diet, sea urchins also eat slow-moving (sessile) animals. Predators that eat sea urchins include a wide variety of fish, starfish, crabs, marine mammals. Sea urchins are also used as food especially in Japan. Adult sea urchins have fivefold symmetry, but their pluteus larvae feature bilateral (mirror) symmetry, indicating that the sea urchin belongs to the Bilateria group of animal phyla, which also comprises the chordates and the arthropods, the annelids and the molluscs, and are found in every ocean and in every climate, from the tropics to the pol ...
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Genetics
Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar working in the 19th century in Brno, was the first to study genetics scientifically. Mendel studied "trait inheritance", patterns in the way traits are handed down from parents to offspring over time. He observed that organisms (pea plants) inherit traits by way of discrete "units of inheritance". This term, still used today, is a somewhat ambiguous definition of what is referred to as a gene. Trait inheritance and molecular inheritance mechanisms of genes are still primary principles of genetics in the 21st century, but modern genetics has expanded to study the function and behavior of genes. Gene structure and function, variation, and distribution are studied within the context of the cell, the organism (e.g. dominance), and within the ...
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