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Bothriolepidoidei
Bothriolepidoidei is a suborder of antiarch placoderm fishes. The group is considered paraphyletic. Taxonomy The cladogram is taken from Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. {{Clade, style={{Clade , label1= Bothriolepidoidei , 1={{Clade , 1={{Clade , 1='' Microbrachius'' , 2='' Wudinolepis'' , 2={{Clade , 1={{Clade , 1=''Dianolepis'' , 2=''Jiangxilepis'' , 3='' Kirgisolepis'' , 4='' Tenizolepis'' , 2={{Clade , 1=''Monarolepis'' , 2={{Clade , 1='' Bothriolepis'' , 2=''Grossilepis Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coi ...'' , 3='' Vietnamaspis'' G ...
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Bothriolepidoidei
Bothriolepidoidei is a suborder of antiarch placoderm fishes. The group is considered paraphyletic. Taxonomy The cladogram is taken from Bothriolepid antiarchs (Vertebrata, Placodermi) from the Devonian of the north-western part of the East European Platform. {{Clade, style={{Clade , label1= Bothriolepidoidei , 1={{Clade , 1={{Clade , 1='' Microbrachius'' , 2='' Wudinolepis'' , 2={{Clade , 1={{Clade , 1=''Dianolepis'' , 2=''Jiangxilepis'' , 3='' Kirgisolepis'' , 4='' Tenizolepis'' , 2={{Clade , 1=''Monarolepis'' , 2={{Clade , 1='' Bothriolepis'' , 2=''Grossilepis Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coi ...'' , 3='' Vietnamaspis'' G ...
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Antiarchi
Antiarchi ("opposite anus") is an order (biology), order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coined by Edward Drinker Cope, who, when examining some fossils that he thought were armored tunicates related to ''Chelysoma'', mistakenly thought that the orbital fenestra (i.e., the hole in the headshield for the eyes, nose and pineal foramen) was the opening for the mouth, or oral siphon, and that the opening for the anal siphon was on the other side of the body, as opposed to having both oral and anal siphons together at one end. The front portions of their bodies were heavily armored, to the point of literally resembling a box with eyes, with the sometimes scaled, sometimes naked rear portions often becoming wiktionary:sinuous, sinuous, particularly with later forms. The pair of pectoral fins were modified into a pair of ...
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Dianolepis
Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coined by Edward Drinker Cope, who, when examining some fossils that he thought were armored tunicates related to '' Chelysoma'', mistakenly thought that the orbital fenestra (i.e., the hole in the headshield for the eyes, nose and pineal foramen) was the opening for the mouth, or oral siphon, and that the opening for the anal siphon was on the other side of the body, as opposed to having both oral and anal siphons together at one end. The front portions of their bodies were heavily armored, to the point of literally resembling a box with eyes, with the sometimes scaled, sometimes naked rear portions often becoming sinuous, particularly with later forms. The pair of pectoral fins were modified into a pair of caliper-like, or arthropod-like ...
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Jiangxilepis
Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coined by Edward Drinker Cope, who, when examining some fossils that he thought were armored tunicates related to '' Chelysoma'', mistakenly thought that the orbital fenestra (i.e., the hole in the headshield for the eyes, nose and pineal foramen) was the opening for the mouth, or oral siphon, and that the opening for the anal siphon was on the other side of the body, as opposed to having both oral and anal siphons together at one end. The front portions of their bodies were heavily armored, to the point of literally resembling a box with eyes, with the sometimes scaled, sometimes naked rear portions often becoming sinuous, particularly with later forms. The pair of pectoral fins were modified into a pair of caliper-like, or arthropod-like ...
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Kirgisolepis
Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coined by Edward Drinker Cope, who, when examining some fossils that he thought were armored tunicates related to '' Chelysoma'', mistakenly thought that the orbital fenestra (i.e., the hole in the headshield for the eyes, nose and pineal foramen) was the opening for the mouth, or oral siphon, and that the opening for the anal siphon was on the other side of the body, as opposed to having both oral and anal siphons together at one end. The front portions of their bodies were heavily armored, to the point of literally resembling a box with eyes, with the sometimes scaled, sometimes naked rear portions often becoming sinuous, particularly with later forms. The pair of pectoral fins were modified into a pair of caliper-like, or arthropod-like ...
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Monarolepis
Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coined by Edward Drinker Cope, who, when examining some fossils that he thought were armored tunicates related to '' Chelysoma'', mistakenly thought that the orbital fenestra (i.e., the hole in the headshield for the eyes, nose and pineal foramen) was the opening for the mouth, or oral siphon, and that the opening for the anal siphon was on the other side of the body, as opposed to having both oral and anal siphons together at one end. The front portions of their bodies were heavily armored, to the point of literally resembling a box with eyes, with the sometimes scaled, sometimes naked rear portions often becoming sinuous, particularly with later forms. The pair of pectoral fins were modified into a pair of caliper-like, or arthropod-like ...
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Grossilepis
Antiarchi ("opposite anus") is an order of heavily armored placoderms. The antiarchs form the second-most successful group of placoderms after the arthrodires in terms of numbers of species and range of environments. The order's name was coined by Edward Drinker Cope, who, when examining some fossils that he thought were armored tunicates related to '' Chelysoma'', mistakenly thought that the orbital fenestra (i.e., the hole in the headshield for the eyes, nose and pineal foramen) was the opening for the mouth, or oral siphon, and that the opening for the anal siphon was on the other side of the body, as opposed to having both oral and anal siphons together at one end. The front portions of their bodies were heavily armored, to the point of literally resembling a box with eyes, with the sometimes scaled, sometimes naked rear portions often becoming sinuous, particularly with later forms. The pair of pectoral fins were modified into a pair of caliper-like, or arthropod-like ...
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Emsian
The Emsian is one of three faunal stages in the Early Devonian Epoch. It lasted from 407.6 ± 2.6 million years ago to 393.3 ± 1.2 million years ago. It was preceded by the Pragian Stage and followed by the Eifelian Stage. It is named after the Ems river in Germany. The GSSP is located in the Zinzil'ban Gorge in the Kitab State Geological Reserve of Uzbekistan, above the contact with the Madmon Formation. In North America the Emsian Stage is represented by Sawkill or Sawkillian time. Biological events During this period, earliest known agoniatitid ammonoid fossils began appearing within this stage after first appearing in previous stage and began to evolutionarily radiate within this stage, in which a new ammonoid order Goniatitida rises in the end of Zlichovian stage (Siberian representation; corresponds to early Eifelian and after the end of Early Devonian, before 391.9 mya). Later agoniatitid ammonoids would die out in the Taghanic event in the upper middle Givetian ...
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Famennian
The Famennian is the latter of two faunal stages in the Late Devonian Epoch. The most recent estimate for its duration estimates that it lasted from around 371.1 million years ago to 359.3 million years ago. An earlier 2012 estimate, still used by the International Commission on Stratigraphy, estimated that it lasted from million years ago to million years ago. It was preceded by the Frasnian stage and followed by the Tournaisian stage. Major events In the seas, a novel major group of ammonoid cephalopods called clymeniids appeared, underwent tremendous diversification and spread worldwide, then just as suddenly went extinct. The beginning of the Famennian is marked by the final stages of a major extinction event, the Kellwasser Event, which is the largest component of the Late Devonian Mass extinction. The end of the Famennian experiences a smaller but still quite severe extinction event, the Hangenberg Event. A brief episode of glaciation, possibly linked to the Hangenber ...
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Paraphyly
In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia ( reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor, including all extant reptiles as well as the extinct synapsids, ...
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Microbrachius
''Microbrachius'' is an extinct genus of tiny, advanced antiarch placoderms closely related to the bothriolepids. Complete articulated specimens show that the armored section of the body had an average length of 2-4 cm. Species of ''Microbrachius'' are characterized by having large heads with short thoracic armor. Specimens of ''Microbrachius'' have been found in Scotland, Belarus, Estonia, and China. Specimens range in age from the Lower Devonian Late Emsian Stage (393.3 Ma) to the Middle Devonian Upper Givetian Stage (382.7 Ma). Although two of the four described species are known from Late Emsian-aged strata of Early Devonian China, the type and youngest species, ''M. dicki'', is known from upper Givetian freshwater strata of Scotland. The various species have short thoracic armor, and large heads. There are patterns of small, but noticeable tubercles on the armor, with the arrangement varying from species to species. Anatomy The bony armor of placoderms surrounds the he ...
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Wudinolepis
Microbrachiidae is an extinct family of tiny, advanced antiarch placoderms closely related to the bothriolepidids. Although most species are known from Late Emsian-aged strata of Early Devonian China, the type and youngest species, ''Microbrachius dicki'', is known from upper Givetian freshwater strata of Scotland and Estonia. When ''Microbrachius'' was regarded as an Antiarch '' incertae sedis'', the other genera were thought to be very small bothriolepids. Genera ''Hohsienolepis'' Fossils of this genus are found in the Xindu Formation portion of the Late Emsian-aged Chuandong Assembly, in Pingle, Guangxi Province, China. Compared to other microbrachiid genera, ''Hohsienolepis'' has a comparatively small head, and an elongated thorax. The armor plates are covered in rows of small tubercles, in a pattern similar to that of ''M. sinensis''. ''Microbrachium'' A genus of very small antiarchs, originally described from ''M. dicki'', of Upper Givetian strata of Scotland and Es ...
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