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Boletus Impolitus
''Hemileccinum impolitum'' is a basidiomycete fungus of the family Boletaceae, native to Europe. It is commonly referred to as the iodine bolete, because its fruit bodies tend to emit an iodine odour when cut, more detectable in the stem base or overripe specimens. Like other members of the family, ''H. impolitum'' has tubes and pores instead of gills in the hymenial surface of its fruit bodies. It is widely distributed in temperate and southern Europe, where it grows in mycorrhizal symbiosis with broad-leaved trees, particularly oak (''Quercus''). Taxonomy and phylogeny The iodine bolete was first described by Elias Magnus Fries, an eminent mycologist of the 19th century, who placed the fungus in genus ''Boletus''. The Latin epithet ''impolitum'' (meaning "rough"), likely refers to the cap of the species, which is initially felty and covered in a finely filamentous coating when viewed under a magnifying glass. The species' taxonomic position had long remained uncertain and va ...
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Fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''t ...
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Hymenium
The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in others some cells develop into sterile cells called cystidia (basidiomycetes) or paraphyses (ascomycetes). Cystidia are often important for microscopic identification. The subhymenium consists of the supportive hyphae from which the cells of the hymenium grow, beneath which is the hymenophoral trama, the hyphae that make up the mass of the hymenophore. The position of the hymenium is traditionally the first characteristic used in the classification and identification of mushrooms. Below are some examples of the diverse types which exist among the macroscopic Basidiomycota and Ascomycota. * In agarics, the hymenium is on the vertical faces of the gills. * In boletes and polypores, it is in a spongy mass of downward-pointing tubes. * In puffballs, ...
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Boletus Impolitus 2009 G1
''Boletus'' is a genus of mushroom-producing fungi, comprising over 100 species. The genus ''Boletus'' was originally broadly defined and described by Carl Linnaeus in 1753, essentially containing all fungi with hymenial pores instead of gills. Since then, other genera have been defined gradually, such as ''Tylopilus'' by Petter Adolf Karsten in 1881, and old names such as ''Leccinum'' have been resurrected or redefined. Some mushrooms listed in older books as members of the genus have now been placed in separate genera. These include such as ''Boletus scaber'', now ''Leccinum scabrum'', ''Tylopilus felleus'', ''Chalciporus piperatus'' and ''Suillus luteus''. Most boletes have been found to be ectomycorrhizal fungi, which mean that they form a mutualistic relationship with the roots system of certain kinds of plants. More recently, ''Boletus'' has been found to be massively polyphyletic, with only a small percentage of the over 300 species that have been assigned to ''Boletus' ...
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Monophyly
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have taken ...
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Corneroboletus
''Corneroboletus'' is a fungal genus in the family Boletaceae. It was circumscribed in 2012 to contain the species formerly known as ''Boletus indecorus''. ''Corneroboletus indecorus'' is found in southeastern Asia, where it grows on the ground in clusters in beech Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engle ... forests. The genus name honors botanist E.J.H. Corner. References Boletaceae Fungi of Asia Monotypic Boletales genera {{Boletales-stub ...
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Lineage (evolution)
An evolutionary lineage is a temporal series of populations, organisms, cells, or genes connected by a continuous line of descent from ancestor to descendant.The Oxford English Dictionary defines biological lineage as "a sequence of species each of which is considered to have evolved from its predecessorOED definition of lineage/ref> Lineages are subsets of the evolutionary tree of life. Lineages are often determined by the techniques of molecular systematics. Phylogenetic representation of lineages 299x299px, A rooted tree of life into three ancient monophyletic lineages: bacteria, archaea, and eukaryotes">archaea.html" ;"title="bacteria, archaea">bacteria, archaea, and eukaryotes based on rRNA genes Lineages are typically visualized as subsets of a phylogenetic tree. A lineage is a single line of descent or linear chain within the tree, while a clade is a (usually branched) monophyletic group, containing a single ancestor and all its descendants. Phylogenetic trees are typica ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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28S Ribosomal RNA
28S ribosomal RNA is the structural ribosomal RNA (rRNA) for the large subunit (LSU) of eukaryotic cytoplasmic ribosomes, and thus one of the basic components of all eukaryotic cells. It has a size of 25S in plants and 28S in mammals, hence the alias of 25S–28S rRNA. Combined with 5.8S rRNA to the 5' side, it is the eukaryotic nuclear homologue of the prokaryotic 23S and mitochondrial 16S ribosomal RNAs. Use in phylogeny The genes coding for 28S rRNA are referred to as 28S rDNA. The comparison of the sequences from these genes are sometimes used in molecular analysis to construct phylogenetic trees, for example in protists, fungi, insects, arachnids, tardigrades, and vertebrates. Structure The 28S rRNA is typically 4000–5000 nt long. Some eukaryotes cleave 28S rRNA into two parts before assembling both into the ribosome, a phenomenon termed the "hidden break". Databases Several databases provide alignments and annotations of LSU rRNA sequences for comparati ...
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Magnifying Glass
A magnifying glass is a convex lens that is used to produce a magnified image of an object. The lens is usually mounted in a frame with a handle. A magnifying glass can be used to focus light, such as to concentrate the sun's radiation to create a hot spot at the focus for fire starting. A sheet magnifier consists of many very narrow concentric ring-shaped lenses, such that the combination acts as a single lens but is much thinner. This arrangement is known as a Fresnel lens. The magnifying glass is an icon of detective fiction, particularly that of Sherlock Holmes. History "The evidence indicates that the use of lenses was widespread throughout the Middle East and the Mediterranean basin over several millennia". The earliest explicit written evidence of a magnifying device is a joke in Aristophanes's ''The Clouds'' from 424 BC, where magnifying lenses to ignite tinder were sold in a pharmacy, and Pliny the Elder's "lens", a glass globe filled with water, used to cauteri ...
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Pileus (mycology)
The pileus is the technical name for the cap, or cap-like part, of a basidiocarp or ascocarp (fungal fruiting body) that supports a spore-bearing surface, the hymenium.Moore-Landecker, E: "Fundamentals of the Fungi", page 560. Prentice Hall, 1972. The hymenium (hymenophore) may consist of lamellae, tubes, or teeth, on the underside of the pileus. A pileus is characteristic of agarics, boletes, some polypores, tooth fungi, and some ascomycetes. Classification Pilei can be formed in various shapes, and the shapes can change over the course of the developmental cycle of a fungus. The most familiar pileus shape is hemispherical or ''convex.'' Convex pilei often continue to expand as they mature until they become flat. Many well-known species have a convex pileus, including the button mushroom, various ''Amanita'' species and boletes. Some, such as the parasol mushroom, have distinct bosses or umbos and are described as ''umbonate''. An umbo is a knobby protrusion at the center of th ...
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Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the Roman Republic it became the dominant language in the Italian region and subsequently throughout the Roman Empire. Even after the fall of Western Rome, Latin remained the common language of international communication, science, scholarship and academia in Europe until well into the 18th century, when other regional vernaculars (including its own descendants, the Romance languages) supplanted it in common academic and political usage, and it eventually became a dead language in the modern linguistic definition. Latin is a highly inflected language, with three distinct genders (masculine, feminine, and neuter), six or seven noun cases (nominative, accusative, genitive, dative, ablative, and vocative), five declensions, four verb conjuga ...
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Quercus
An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably '' Lithocarpus'' (stone oaks), as well as in those of unrelated species such as ''Grevillea robusta'' (silky oaks) and the Casuarinaceae (she-oaks). The genus ''Quercus'' is native to the Northern Hemisphere, and includes deciduous and evergreen species extending from cool temperate to tropical latitudes in the Americas, Asia, Europe, and North Africa. North America has the largest number of oak species, with approximately 160 species in Mexico of which 109 are endemic and about 90 in the United States. The second greatest area of oak diversity is China, with approximately 100 species. Description Oaks have spirally arranged leaves, with lobate margins in many species; some have serrated leaves or entire leaves with smooth margins. ...
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