Argyroxiphium
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Argyroxiphium
''Argyroxiphium'' is a small genus of plants in the family Asteraceae. Its members are known by the common names silversword or greensword due to their long, narrow leaves and the silvery hairs on some species. The silverswords belong to a larger Adaptive radiation, radiation of over 50 species, including the physically different genera ''Dubautia'' and ''Wilkesia''. This grouping is often referred to as the silversword alliance.[B. Baldwin. Age and Rate of Diversification of the Hawaiian Silversword Alliance (Compositae). 1998.] Botanist P. H. Raven referred to this radiation as "the best example of adaptive radiation in plants". Description Species in ''Argyroxiphium'' are Perennial plant, perennial, Rosette (botany), rosette-forming shrubs. They may consist of a single large rosette or a collection of several rosettes. In all ''Argyroxiphium'' species, long, narrow leaves contain interstitial gels hypothesized to function as water storage. For some species, leaves are cove ...
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Haleakalā Silversword
''Argyroxiphium sandwicense'' subsp. ''macrocephalum'', the east Maui silversword or Haleakala silversword, is a rare plant, part of the family (biology), family Asteraceae. The silversword in general is referred to as ''āhinahina'' in Hawaiian language, Hawaiian (literally, "very gray"). Distribution The Haleakala silversword is found on the island of Maui at elevations above on the dormant Haleakalā volcanoon the summit depression, the rim summits, and surrounding slopesin Haleakalā National Park. The Haleakala silversword has been a threatened species as defined by the U.S. Fish and Wildlife Service, since May 15, 1992. Prior to that time, excessive grazing by cattle and goats, and vandalism inflicted by people in the 1920s, had caused its near extinction. Since strict monitoring and governmental protection took effect, the species' recovery is considered a successful conservation story, although threats remain. Description The Haleakala silversword, ''Argyroxiphium sand ...
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Silversword Alliance
The silversword alliance, also known as the tarweeds, refers to an adaptive radiation of around 30 species in the composite or sunflower family, Asteraceae. The group is endemic to Hawaii, and is derived from a single immigrant to the islands. For radiating from a common ancestor at an estimated 5.2±0.8 Ma, the clade is extremely diverse, composed of trees, shrubs, subshrubs, mat-plants, cushion plants, rosette plants, and lianas. The silversword alliance is named for its most famous and visually striking members, the silverswords. The species of the clade break down into three genera: ''Wilkesia, Argyroxiphium,'' and ''Dubautia.'' There are three species of silverswords and two greenswords in the genus ''Argyroxiphium'', confined to the islands of Maui and Hawaii, and two species of ''Wilkesia'' (''iliau'') on Kauai. The bulk of the species are placed in the genus ''Dubautia'', which is widespread on all the main islands. The genus ''Dubautia'' contains a wide variety of form ...
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Adaptive Radiation
In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic interactions or opens new environmental niches. Starting with a single ancestor, this process results in the speciation and phenotypic adaptation of an array of species exhibiting different morphological and physiological traits. The prototypical example of adaptive radiation is finch speciation on the Galapagos ("Darwin's finches"), but examples are known from around the world. Characteristics Four features can be used to identify an adaptive radiation: #A common ancestry of component species: specifically a ''recent'' ancestry. Note that this is not the same as a monophyly in which ''all'' descendants of a common ancestor are included. #A phenotype-environment correlation: a ''significant'' association between environments and the morp ...
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Madieae
Madieae is a tribe of flowering plants in the family Asteraceae. It is sometimes considered a subtribe of Heliantheae. Notable species include the tarweeds of the Western United States as well as the silverswords of Hawaii. Subtribes and genera Madieae subtribes and genera recognized by the Global Compositae Database as of April 2022: *Subtribe Arnicinae **''Arnica'' *Subtribe Baeriinae **'' Amblyopappus'' **'' Baeriopsis'' **'' Constancea'' **''Eriophyllum'' **''Lasthenia'' **'' Monolopia'' **'' Pseudobahia'' **'' Syntrichopappus'' *Subtribe Hulseinae **'' Eatonella'' **'' Hulsea'' *Subtribe Madiinae **''Achyrachaena'' **'' Adenothamnus'' **'' Anisocarpus'' **''Argyroxiphium'' **'' Blepharipappus'' **'' Blepharizonia'' **'' Calycadenia'' **'' Carlquistia'' **'' Centromadia'' **''Deinandra'' **''Dubautia'' **''Harmonia'' **'' Hemizonella'' **'' Hemizonia'' **'' Holocarpha'' **'' Holozonia'' **'' Jensia'' **''Kyhosia'' **'' Lagophylla'' **'' L ...
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Monocarpic
Monocarpic plants are those that flower and set seeds only once, and then die. The term is derived from Greek (''mono'', "single" + ''karpos'', "fruit" or "grain"), and was first used by Alphonse de Candolle. Other terms with the same meaning are ''hapaxanth'' and ''semelparous''. The antonym is polycarpic, a plant that flowers and sets seeds many times during its lifetime; the antonym of semelparous is ''iteroparous''. Plants which flower en masse (gregariously) before dying are known as plietesials. The term ''hapaxanth'' is most often in conjunction with describing some of the taxa of Arecaceae (palms) and some species of bamboo, but rarely used otherwise; its antonym is ''pleonanth''. This was first used by Alexander Braun. Monocarpic plants are not necessarily annuals, because some monocarpic plants can live a number of years before they will flower. In some monocarpic plants, flowering signals senescence, while in others the production of fruits and seeds causes changes wi ...
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Hawaiian Tropical Rainforests
The Hawaiian tropical rainforests are a tropical moist broadleaf forest ecoregion in the Hawaiian Islands. They cover an area of in the windward lowlands and montane regions of the islands. Coastal mesic forests are found at elevations from sea level to . Mixed mesic forests occur at elevations of , while wet forests are found from . Moist bogs and shrublands exist on montane plateaus and depressions. For the 28 million years of existence of the Hawaiian Islands, they have been isolated from the rest of the world by vast stretches of the Pacific Ocean, and this isolation has resulted in the evolution of an incredible diversity of endemic species, including fungi, mosses, snails, birds, and other wildlife. In the lush, moist forests high in the mountains, trees are draped with vines, orchids, ferns, and mosses. This ecoregion includes one of the world's wettest places, the slopes of Mount Waialeale, which average of rainfall per year. Coastal mesic forests Coastal mesic ...
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Wilkesia
''Wilkesia'' is a genus of Hawaiian plants in the tribe Madieae within the family Asteraceae. It contains two perennials, both of which are endemic to the island of Kauai in Hawaii. ''Wilkesia'' is a component of the silversword alliance and is named after Captain Charles Wilkes. ;Species * ''Wilkesia gymnoxiphium'' A.Gray, 1852 * ''Wilkesia hobdyi'' H.St.John, 1971 ;formerly included ''Wilkesia grayana - Argyroxiphium grayanum ''Argyroxiphium grayanum'', commonly known as the greensword, is a species of flowering plant in the family Asteraceae, and a member of the silversword alliance, a group of over 50 species which are diverse in morphology and habitat but are gene ...'' References External links * Madieae Endemic flora of Hawaii Biota of Kauai Asteraceae genera Taxa named by Asa Gray {{Madieae-stub ...
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Pollinator
A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the major pollinators of most plants, and insect pollinators include all families of bees and most families of aculeate wasps; ants; many families of flies; many lepidopterans (both butterflies and moths); and many families of beetles. Vertebrates, mainly bats and birds, but also some non-bat mammals (monkeys, lemurs, possums, rodents) and some lizards pollinate certain plants. Among the pollinating birds are hummingbirds, honeyeaters and sunbirds with long beaks; they pollinate a number of deep-throated flowers. Humans may also carry out artificial pollination. A pollinator is different from a pollenizer, a plant that is a source of pollen for the pollination process. Background Plants fall into pollination syndromes that reflect the type o ...
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Hawaii (island)
Hawaii ( ; haw, Hawaii ) is the List of islands of the United States by area, largest island in the United States, located in the U.S. state, state of Hawaii. It is the southeasternmost of the Hawaiian Islands, a chain of High island, volcanic islands in the Pacific Ocean, North Pacific Ocean. With an area of , it has 63% of the Hawaiian archipelago's combined landmass. However, it has only 13% of Hawaiʻi's population. The island of Hawaiʻi is the third largest island in Polynesia, behind the two main List of islands of New Zealand, islands of New Zealand. The island is often referred to as the Island of Hawaii or Hawaii Island to distinguish it from the state. It is also referred to as the Big Island. Administratively, the island is coextensive with Hawaii County, Hawaii, Hawaii County. As of the 2020 United States census, 2020 census, the population was 200,629. The county seat and largest town is Hilo, Hawaii, Hilo. There are no Municipal corporation, incorporated cities i ...
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Maui
The island of Maui (; Hawaiian: ) is the second-largest of the islands of the state of Hawaii at 727.2 square miles (1,883 km2) and is the 17th largest island in the United States. Maui is the largest of Maui County's four islands, which also includes Molokai, Lānai, and unpopulated Kahoolawe. In 2020, Maui had a population of 168,307, the third-highest of the Hawaiian Islands, behind that of Oahu and Hawaii Island. Kahului is the largest census-designated place (CDP) on the island with a population of 26,337 , and is the commercial and financial hub of the island. Wailuku is the seat of Maui County and is the third-largest CDP . Other significant places include Kīhei (including Wailea and Makena in the Kihei Town CDP, the island's second-most-populated CDP), Lāhainā (including Kāanapali and Kapalua in the Lāhainā Town CDP), Makawao, Pukalani, Pāia, Kula, Haikū, and Hāna. Etymology Native Hawaiian tradition gives the origin of the island's name in th ...
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Hawaii
Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only state geographically located within the tropics. Hawaii comprises nearly the entire Hawaiian archipelago, 137 volcanic islands spanning that are physiographically and ethnologically part of the Polynesian subregion of Oceania. The state's ocean coastline is consequently the fourth-longest in the U.S., at about . The eight main islands, from northwest to southeast, are Niihau, Kauai, Oahu, Molokai, Lānai, Kahoolawe, Maui, and Hawaii—the last of these, after which the state is named, is often called the "Big Island" or "Hawaii Island" to avoid confusion with the state or archipelago. The uninhabited Northwestern Hawaiian Islands make up most of the Papahānaumokuākea Marine National Monument, the United States' largest protected ...
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Endemism
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can be also be referred to as an ''endemism'' or in scientific literature as an ''endemite''. For example '' Cytisus aeolicus'' is an endemite of the Italian flora. '' Adzharia renschi'' was once believed to be an endemite of the Caucasus, but it was later discovered to be a non-indigenous species from South America belonging to a different genus. The extreme opposite of an endemic species is one with a cosmopolitan distribution, having a global or widespread range. A rare alternative term for a species that is endemic is "precinctive", which applies to ...
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