Acroceridae
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Acroceridae
The Acroceridae are a small family (biology), family of odd-looking flies. They have a hump-backed appearance with a strikingly small head, generally with a long proboscis for accessing nectar. They are rare and not widely known. The most frequently applied common names are small-headed flies or hunch-back flies. Many are bee or wasp mimicry, mimics. Because they are parasitoids of spiders, they also are sometimes known as spider flies. Description The Acroceridae vary in size from small to fairly large, about the size of large bees, with a wingspan over 25 mm in some species. As a rule, both sexes have tiny heads and a characteristic hump-backed appearance because of the large, rounded thorax. In appearance, they are compact flies without major bristles, but many species have a bee-like hairiness on their bodies, and some are bee or wasp mimicry, mimics. In most species, the eyes are holoptic in both sexes, the heads seemingly composed mainly of the large ommatidia, facete ...
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Acroceridae Psilodera Small-headed Fly IMG 1338
The Acroceridae are a small family of odd-looking flies. They have a hump-backed appearance with a strikingly small head, generally with a long proboscis for accessing nectar. They are rare and not widely known. The most frequently applied common names are small-headed flies or hunch-back flies. Many are bee or wasp mimics. Because they are parasitoids of spiders, they also are sometimes known as spider flies. Description The Acroceridae vary in size from small to fairly large, about the size of large bees, with a wingspan over 25 mm in some species. As a rule, both sexes have tiny heads and a characteristic hump-backed appearance because of the large, rounded thorax. In appearance, they are compact flies without major bristles, but many species have a bee-like hairiness on their bodies, and some are bee or wasp mimics. In most species, the eyes are holoptic in both sexes, the heads seemingly composed mainly of the large faceted eyes. This is in contrast to many insects in ...
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Acrocerinae
Acrocerinae is a subfamily of Acroceridae. They are small distinctive flies whose larvae are endoparasites of spiders. Adult hunchback-flies visit flowers to feed on nectar. Traditionally the subfamily included the genera now placed in Cyrtinae and Ogcodinae Ogcodinae is a subfamily of the Acroceridae (small-headed flies). Their larvae are endoparasites of araneomorph spiders in the subgroup Entelegynae The Entelegynae or entelegynes are a subgroup of araneomorph spiders, the largest of the two mai ..., but the subfamily in this sense was found to be polyphyletic and was split up in 2019. Systematics The subfamily includes two extant genera and one extinct: * '' Acrocera'' Meigen, 1803 * '' Carvalhoa'' Koçak & Kemal, 2013 * †'' Schlingeromyia'' Grimaldi & Hauser ''in'' Grimaldi, Arillo, Cumming & Hauser, 2011 The extinct fly genus †'' Burmacyrtus'' Grimaldi & Hauser ''in'' Grimaldi, Arillo, Cumming & Hauser, 2011 was originally placed in this subfamily as well, but acco ...
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Cyrtinae
Cyrtinae is a subfamily of the Acroceridae (small-headed flies). Their larvae are endoparasites of araneomorph spiders in the subgroup Entelegynae. Genera The subfamily includes twelve extant genera and two extinct: * ''Asopsebius'' Nartshuk, 1982 * †''Cyrtinella'' Gillung & Winterton, 2017 * ''Cyrtus'' Pierre André Latreille, Latreille, 1796 * ''Hadrogaster'' Schlinger, 1972 * ''Holops'' Rodolfo Amando Philippi, Philippi, 1865 * ''Meruia'' Curtis Williams Sabrosky, Sabrosky, 1950 * ''Nipponcyrtus'' Schlinger, 1972 * ''Opsebius'' Achille Costa, Costa, 1856 * ''Paracyrtus'' Schlinger, 1972 * ''Sabroskya'' Schlinger, 1960 * ''Subcyrtus'' Enrico Brunetti, Brunetti, 1926 * ''Turbopsebius'' Schlinger, 1972 * †''Villalites'' Willi Hennig, Hennig, 1966 * ''Villalus'' Cole, 1918 References

Acroceridae Brachycera subfamilies Endoparasites {{Nemestrinoidea-stub ...
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Panopinae
Panopinae is a subfamily of small-headed flies (Acroceridae). Their larvae are endoparasites of spiders in the infraorder Mygalomorphae. Genera The subfamily includes 24 extant genera: * '' Apelleia'' Bellardi, 1862 * '' Apsona'' Westwood, 1876 * '' Archipialea'' Schlinger, 1973 * '' Arrhynchus'' Philippi, 1871 * '' Astomella'' Latreille, 1809 * '' Astomelloides'' Schlinger, 1959 * '' Camposella'' Cole, 1919 * '' Coquena'' Schlinger ''in'' Schlinger, Gillung & Borkent, 2013 * '' Corononcodes'' Speiser, 1920 * '' Eulonchus'' Gerstaecker, 1856 * '' Exetasis'' Walker, 1852 * '' Lasia'' Wiedemann, 1824 * '' Lasioides'' Gil Collado, 1928 * '' Leucopsina'' Westwood, 1876 * '' Mesophysa'' Macquart, 1838 * '' Ocnaea'' Erichson, 1840 * '' Panops'' Lamarck, 1804 * '' Physegastrella'' Brunetti, 1926 * '' Pialea'' Erichson, 1840 * '' Psilodera'' Gray ''in'' Griffith & Pidgeon, 1832 * '' Pterodontia'' Gray ''in'' Griffith Griffith may refer to: People * Griffith (name) * Griffi ...
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Philopotinae
Philopotinae is a subfamily of small-headed flies. Their larvae are endoparasites of araneomorph spiders in the subgroup Entelegynae. Genera The subfamily includes twelve extant genera and four extinct: * '' Africaterphis'' Schlinger, 1968 * †'' Archaeterphis'' Hauser & Winterton, 2007 * '' Dimacrocolus'' Schlinger, 1961 * †'' Eulonchiella'' Meunier, 1912 * '' Helle'' Osten Sacken, 1896 * †'' Hoffeinsomyia'' Gillung & Winterton, 2017 * '' Megalybus'' Philippi, 1865 * '' Neophilopota'' Schlinger ''in'' Schlinger, Gillung & Borkent, 2013 * '' Oligoneura'' Bigot, 1878 * '' Parahelle'' Schlinger, 1961 * '' Philopota'' Wiedemann, 1830 * †'' Prophilopota'' Hennig, 1966 * ''Quasi Quasi (phonetics 'kwa - zee') is an American indie rock band formed in Portland, Oregon in 1993 by former spouses Sam Coomes ( vocals, guitar, rocksichord, various keyboards, bass) and Janet Weiss (vocals and drums). Joanna Bolme performe ...'' Gillung & Winterton, 2011 * '' Schlingeriell ...
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Pterodontia
''Pterodontia'' is a genus of small-headed flies (insects in the family Acroceridae). There are at least 20 described species in ''Pterodontia''. Description ''Pterodontia'' have eyes that are covered in hairs. Their antennae are attached below the middle of the head, and are small, short and inconspicuous. Their mouthparts are small and nearly imperceptible. The tibia have small, atypical spurs. The males have a tooth-like projection on the costal edge of the wing. Species These 19 species belong to the genus ''Pterodontia'': * '' Pterodontia aerivaga'' Seguy, 1962 * '' Pterodontia analis'' Macquart, 1846 * '' Pterodontia andina'' Brèthes, 1910 * '' Pterodontia davisi'' Paramonov, 1957 * '' Pterodontia dimidiata'' Westwood, 1876 * '' Pterodontia ezoensis'' Ouchi, 1942 * '' Pterodontia flavipes'' Gray, 1832 * '' Pterodontia flavonigra'' Carrera, 1947 * '' Pterodontia johnsoni'' Cole, 1919 * '' Pterodontia kashmirensis'' Lichtwardt, 1909 * '' Pterodontia longisquama'' Sabro ...
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Archocyrtinae
''Archocyrtus'' is an extinct genus of small-headed flies in the family Acroceridae. The genus is known from compression fossils from the Late Jurassic Karabastau Formation of Kazakhstan. The genus is the only member of the subfamily Archocyrtinae. A study on the holotype specimen of ''Archocyrtus kovalevi'' was published by Khramov & Lukashevich (2019). They reported evidence of an extremely long proboscis, almost twice the length of the body of the insect. They suggested that it was probably a pollinator of extinct seed plants belonging to the order Bennettitales Bennettitales (also known as cycadeoids) is an extinct order of seed plants that first appeared in the Permian period and became extinct in most areas toward the end of the Cretaceous. Bennettitales are among the most common Mesozoic seed plants .... Species Two species belong to the genus ''Archocyrtus'': * †''A. gibbosus'' Ussatchov, 1968 * †''A. kovalevi'' (Nartshuk, 1996) References † Prehistoric D ...
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Archocyrtus
''Archocyrtus'' is an extinct genus of small-headed flies in the family Acroceridae. The genus is known from compression fossils from the Late Jurassic Karabastau Formation of Kazakhstan. The genus is the only member of the subfamily Archocyrtinae. A study on the holotype specimen of ''Archocyrtus kovalevi'' was published by Khramov & Lukashevich (2019). They reported evidence of an extremely long proboscis, almost twice the length of the body of the insect. They suggested that it was probably a pollinator of extinct seed plants belonging to the order Bennettitales Bennettitales (also known as cycadeoids) is an extinct order of seed plants that first appeared in the Permian period and became extinct in most areas toward the end of the Cretaceous. Bennettitales are among the most common Mesozoic seed plants .... Species Two species belong to the genus ''Archocyrtus'': * †''A. gibbosus'' Ussatchov, 1968 * †''A. kovalevi'' (Nartshuk, 1996) References † Prehistoric D ...
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Ogcodinae
Ogcodinae is a subfamily of the Acroceridae (small-headed flies). Their larvae are endoparasites of araneomorph spiders in the subgroup Entelegynae The Entelegynae or entelegynes are a subgroup of araneomorph spiders, the largest of the two main groups into which the araneomorphs were traditionally divided. Females have a genital plate ( epigynum) and a "flow through" fertilization system; m .... Genera The subfamily includes one extant genus and one extinct: * †'' Glaesoncodes'' Hennig, 1968 * '' Ogcodes'' Latreille, 1796 References Acroceridae Brachycera subfamilies Endoparasites {{Nemestrinoidea-stub ...
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Hypermetamorphosis
Hypermetamorphosis, or heteromorphosis,P.J. Gullan & P.S. Cranston. 2010. The Insects: An Outline of Entomology, 4th Edition. Wiley-Blackwell. is a term used in entomology that refers to a class of variants of holometabolism, that is to say, complete insect metamorphosis, but where some larval instars are distinct from each other. Description Hypermetamorphosis, as the term normally is used in entomology, refers to a class of variants of holometabolism. In hypermetamorphosis some larval instars are functionally and morphologically distinct from each other. The general case in holometabolous insects such as flies, moths, or wasps, is that all larval stages look similar, growing larger as the insect matures. In hypermetamorphic insects however, at least one instar, usually the first, differs markedly from the rest. In many hypermetamorphic species, the first instars are numerous, tiny, very mobile larvae that must find their way to a food source. The general term for a mobi ...
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Holoptic
Holoptic refers to one of the ways in which the arthropod eye develops, particularly the eyes of various species of insects. Unlike dichoptic and cycloptic eyes, holoptic eyes meet along the median dorsal line of the head, in many species nearly covering the exterior of the head. Holoptic eyes are typical of several Dipteran males, in particular some Syrphidae, Tabanidae, Pipunculidae, and Acroceridae. Some other insect orders that include species with holoptic males and some in which the females are holoptic as well, include the Coleoptera, Anisoptera, and Archaeognatha The Archaeognatha are an order of apterygotes, known by various common names such as jumping bristletails. Among extant insect taxa they are some of the most evolutionarily primitive; they appeared in the Middle Devonian period at about the sa .... References * {{Insect-anatomy-stub Insect anatomy Eye ...
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Nemestrinidae
Nemestrinidae, or tangle-veined flies is a family of flies in the superfamily Nemestrinoidea, closely related to Acroceridae. The family is small but distributed worldwide, with about 300 species in 34 genera. Larvae are endoparasitoids of either grasshoppers (Trichopsideinae) or scarab beetles (Hirmoneurinae). Some are considered important in the control of grasshopper populations. Adults are often observed on flowers. Genera * '' Atriadops'' * '' Ceyloniola'' * '' Cyclopsidea'' * '' Fallenia'' * '' Hirmoneura'' * '' Hyrmophlaeba'' Rondani, 1864 * '' Moegistorhynchus'' * '' Nemestrinus'' * '' Neohirmoneura'' * '' Neorhynchocephalus'' Lichtwardt, 1909 * '' Nycterimorpha'' * '' Nycterimyia'' * '' Prosoeca'' * '' Stenobasipteron'' * '' Stenopteromyia'' * '' Trichophthalma'' * '' Trichopsidea'' Westwood, 1839 Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net Fossil history Fossils of Nemestrinidae are known from several localities of various ages in Russi ...
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