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Archocyrtinae
''Archocyrtus'' is an extinct genus of small-headed flies in the family Acroceridae. The genus is known from compression fossils from the Late Jurassic Karabastau Formation of Kazakhstan. The genus is the only member of the subfamily Archocyrtinae. A study on the holotype specimen of ''Archocyrtus kovalevi'' was published by Khramov & Lukashevich (2019). They reported evidence of an extremely long proboscis, almost twice the length of the body of the insect. They suggested that it was probably a pollinator of extinct seed plants belonging to the order Bennettitales Bennettitales (also known as cycadeoids) is an extinct order of seed plants that first appeared in the Permian period and became extinct in most areas toward the end of the Cretaceous. Bennettitales are among the most common Mesozoic seed plants .... Species Two species belong to the genus ''Archocyrtus'': * †''A. gibbosus'' Ussatchov, 1968 * †''A. kovalevi'' (Nartshuk, 1996) References † Prehistoric D ...
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Acroceridae
The Acroceridae are a small family (biology), family of odd-looking flies. They have a hump-backed appearance with a strikingly small head, generally with a long proboscis for accessing nectar. They are rare and not widely known. The most frequently applied common names are small-headed flies or hunch-back flies. Many are bee or wasp mimicry, mimics. Because they are parasitoids of spiders, they also are sometimes known as spider flies. Description The Acroceridae vary in size from small to fairly large, about the size of large bees, with a wingspan over 25 mm in some species. As a rule, both sexes have tiny heads and a characteristic hump-backed appearance because of the large, rounded thorax. In appearance, they are compact flies without major bristles, but many species have a bee-like hairiness on their bodies, and some are bee or wasp mimicry, mimics. In most species, the eyes are holoptic in both sexes, the heads seemingly composed mainly of the large ommatidia, facete ...
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Journal Of Systematic Palaeontology
The ''Journal of Systematic Palaeontolog'y'' (Print: , online: ) is a peer-reviewed scientific journal of palaeontology published by Taylor & Francis on behalf of the British Natural History Museum. , the editor-in-chief is Paul D. Taylor. The journal covers papers on new or poorly known faunas and floras and new approaches to systematics. It was established in 2003. According to the ''Journal Citation Reports'', the journal has a 2014 impact factor The impact factor (IF) or journal impact factor (JIF) of an academic journal is a scientometric index calculated by Clarivate that reflects the yearly mean number of citations of articles published in the last two years in a given journal, as ... of 3.727, ranking it second out of 49 journals in the category 'Paleontology'. References External links * Paleontology journals Taylor & Francis academic journals Quarterly journals Publications established in 2003 English-language journals {{paleo-journal-stub ...
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Extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Compression Fossil
A compression fossil is a fossil preserved in sedimentary rock that has undergone physical compression. While it is uncommon to find animals preserved as good compression fossils, it is very common to find plants preserved this way. The reason for this is that physical compression of the rock often leads to distortion of the fossil. The best fossils of leaves are found preserved in fine layers of sediment that have been compressed in a direction perpendicular to the plane of the deposited sediment. Since leaves are basically flat, the resulting distortion is minimal. Plant stems and other three-dimensional plant structures do not preserve as well under compression. Typically, only the basic outline and surface features are preserved in compression fossils; internal anatomy is not preserved. These fossils may be studied while still partially entombed in the sedimentary rock matrix where they are preserved, or once lifted out of the matrix by a peel or transfer technique. Compr ...
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Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age. In the past, ''Malm'' was also used to indicate the unit of geological time, but this usage is now discouraged to make a clear distinction between lithostratigraphic and geochronologic/chronostratigraphic units. Subdivisions The Late Jurassic is divided into three ages, which correspond with the three (faunal) stages of Upper Jurassic rock: Paleogeography During the Late Jurassic Epoch, Pangaea broke up into two supercontinents, Laurasia to the north, and Gondwana to the south. The result of this break-up was the spawning of the Atlantic Ocean. However, at this time, the Atlantic Ocean was relatively narrow. Life forms of the epoch This epoch is well known for many famous types of dinosau ...
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Karabastau Formation
The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that has been studied since the early 20th century, alongside the rarer remains of vertebrates, including pterosaurs, salamanders, lizards and crocodiles.Barrett, P.M., Butler, R.J., Edwards, N.P., & Milner, A.R. Pterosaur distribution in time and space: an atlas. p61-107. in Flugsaurier: Pterosaur papers in honour of Peter Wellnhofer. 2008. Hone, D.W.E., and Buffetaut, E. (eds). Zitteliana B, 28. 264p/ref> Lithology and depositional environment The primary lithology consists of 1 mm thick varve laminations of claystone, with a dark part and a light dolomitic part, which probably correspond to a wet and dry season respectively, alongside rare, several cm thick sandstone interbeds. These were deposited within an ancient freshwater paleol ...
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Kazakhstan
Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbekistan to the south, and Turkmenistan to the southwest, with a coastline along the Caspian Sea. Its capital is Astana, known as Nur-Sultan from 2019 to 2022. Almaty, Kazakhstan's largest city, was the country's capital until 1997. Kazakhstan is the world's largest landlocked country, the largest and northernmost Muslim-majority country by land area, and the ninth-largest country in the world. It has a population of 19 million people, and one of the lowest population densities in the world, at fewer than 6 people per square kilometre (15 people per square mile). The country dominates Central Asia economically and politically, generating 60 percent of the region's GDP, primarily through its oil and gas industry; it also has vast mineral ...
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Systematic Entomology
''Systematic Entomology'' is a scientific journal covering the field of systematic entomology, published by the Royal Entomological Society of London. Having begun in 1932 as '' Proceedings of the Royal Entomological Society of London, Series B: Taxonomy'', the title was changed to ''Journal of Entomology, Series B: Taxonomy'' in 1971, starting with volume 40. After volume 44 in 1976, the journal became ''Systematic Entomology'', starting again with volume 1. According to the ''Journal Citation Reports'', the journal has a 2020 impact factor of 3.844. It is indexed in the following bibliographic databases: *''Academic Search'' *''AGRICOLA'' *''Aquatic Sciences and Fisheries Abstracts'' *'' BIOBASE'' *''Biological Abstracts'' *''BIOSIS Previews'' *'' CAB Direct'' *'' CSA Biological Sciences Database'' *'' CSA Environmental Sciences & Pollution Management Database'' *''Current Contents'' *''Embiology'' *'' IBIDS'' *'' InfoTrac'' *''Journal Citation Reports'' *''Science Citation In ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Proboscis
A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular mouthparts used for feeding and sucking. In vertebrates, a proboscis is an elongated nose or snout. Etymology First attested in English in 1609 from Latin , the latinisation of the Ancient Greek (), which comes from () 'forth, forward, before' + (), 'to feed, to nourish'. The plural as derived from the Greek is , but in English the plural form ''proboscises'' occurs frequently. Invertebrates The most common usage is to refer to the tubular feeding and sucking organ of certain invertebrates such as insects (e.g., moths, butterflies, and mosquitoes), worms (including Acanthocephala, proboscis worms) and gastropod molluscs. Acanthocephala The Acanthocephala or thorny-headed worms, or spiny-headed worms are characterized by the presence of an eversible proboscis, armed with spines, which it uses to pierce and ...
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Spermatophyte
A spermatophyte (; ), also known as phanerogam (taxon Phanerogamae) or phaenogam (taxon Phaenogamae), is any plant that produces seeds, hence the alternative name seed plant. Spermatophytes are a subset of the embryophytes or land plants. They include most familiar types of plants, including all flowers and most trees, but exclude some other types of plants such as ferns, mosses, algae. The term ''phanerogams'' or ''phanerogamae'' is derived from the Greek (), meaning "visible", in contrast to the cryptogamae (), together with the suffix (), meaning "to marry". These terms distinguished those plants with hidden sexual organs (cryptogamae) from those with visible sexual organs (phanerogamae). Description The extant spermatophytes form five divisions, the first four of which are traditionally grouped as gymnosperms, plants that have unenclosed, "naked seeds": * Cycadophyta, the cycads, a subtropical and tropical group of plants, * Ginkgophyta, which includes a single living s ...
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Bennettitales
Bennettitales (also known as cycadeoids) is an extinct order of seed plants that first appeared in the Permian period and became extinct in most areas toward the end of the Cretaceous. Bennettitales are among the most common Mesozoic seed plants, and had morphologies including shrub and cycad-like forms. The foliage of bennettitaleans is superficially nearly indistinguishable from that of cycads, but they are distinguished from cycads by their more complex flower-like reproductive organs, at least some of which were likely pollinated by insects. Although certainly gymnosperms (cone-bearing seed plants), the relationships of bennettitaleans to other seed plants is debated. Their general resemblance to cycads is contradicted by numerous more subtle features of their reproductive systems and leaf structure. Some authors have linked bennettitaleans to angiosperms (flowering plants) and gnetophytes (a rare and unusual group of modern gymnosperms), forming a broader group known as Anth ...
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