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Azhdarchid Map
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs ''Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as ''Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteran ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Aralazhdarcho
''Aralazhdarcho'' is a genus of azhdarchid pterosaur from the Santonian to the early Campanian stages of the Late Cretaceous period of Bostobe Svita in Kazakhstan. The type and only known species is ''Aralazhdarcho bostobensis''. Etymology ''Aralazhdarcho'' was named in 2007 by Alexander Averianov. In 2004, the holotype had already been described. (''Paleontol. J.'' 38 (4), 426–436) The genus name, ''Aralazhdarcho'', is derived from the Aral Sea and the related genus ''Azhdarcho'', while the specific name, ''bostobensis'', refers to the Bostobe Formation. Description ''Aralazhdarcho'' is based on holotype ZIN PH, no. 9/43, consisting of the anterior end of a neck vertebra, probably the fifth or sixth. Several paratypes have also been referred: a jugal, a toothless lower jaw fragment, centra from vertebrae, the distal end of a scapula, the proximal end of a second phalanx of the left wing finger and the proximal end of a left femur, of which, however, the head has broken off ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Persian Mythology
Persian mythology or Iranian mythology (Persian:اساطیرشناسی ایرانی) is the body of the myths originally told by ancient Persians and other Iranian peoples, and a genre of Ancient Persian folklore. These stories concern the origin and nature of the world, the lives and activities of deities, heroes, and mythological creatures, and the origins and significance of the ancient Persians' own cult and ritual practices. Modern scholars study the myths to shed light on the religious and political institutions of not only modern-day Iran but the Greater Iran, which includes regions of West Asia, Central Asia, South Asia and Transcaucasia where Iranian culture has had significant influence. Historically, these were regions long ruled by dynasties of various Iranian empires, that incorporated considerable aspects of Persian culture through extensive contact with them, or where sufficient Iranian peoples settled to still maintain communities who patronize their respective cul ...
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Azhdaha
Azhdaha, Azhdahak, Ezhdeha ( fa, اژدها) or Azhdar (اژدر) is a mythical creature in Iranian mythology, roughly equivalent to the dragon. They are gigantic snake-like creatures living in the air, in the sea, or on the earth. ''Sad dar-e nathr'' and ''sad dar-e Bondahesh'', ''Shahnameh'' and ''Garshaspnameh'' are among the principal texts that contain information about the creature. In Persian literature, Azhdahas are depicted as a giant snake or lizard with wings. According to tradition, they have a huge body, a fierce face, their mouths are wide with many teeth, and their eyes are bright. Azhdahās are actually normal snakes, according to ''Ajāyeb ul-Makhlooghāt'', a book by Mohammad b. Mahmoud b. Ahmad-e Tusi (wrote in 1160 AD), "when a snake lives 100 years and its length becomes 30 Gazes (a traditional measurement unit approximate to a meter), it is called an azhdahā". He also wrote that "because of their harassment to other creatures, God eventually will throw them ...
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Persian Language
Persian (), also known by its endonym Farsi (, ', ), is a Western Iranian language belonging to the Iranian branch of the Indo-Iranian subdivision of the Indo-European languages. Persian is a pluricentric language predominantly spoken and used officially within Iran, Afghanistan, and Tajikistan in three mutually intelligible standard varieties, namely Iranian Persian (officially known as ''Persian''), Dari Persian (officially known as ''Dari'' since 1964) and Tajiki Persian (officially known as ''Tajik'' since 1999).Siddikzoda, S. "Tajik Language: Farsi or not Farsi?" in ''Media Insight Central Asia #27'', August 2002. It is also spoken natively in the Tajik variety by a significant population within Uzbekistan, as well as within other regions with a Persianate history in the cultural sphere of Greater Iran. It is written officially within Iran and Afghanistan in the Persian alphabet, a derivation of the Arabic script, and within Tajikistan in the Tajik alphabet, a der ...
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Zhejiangopterus
''Zhejiangopterus'' is a genus of azhdarchid pterosaur known from one species, which lived in China during the late Cretaceous Period. The genus was named in 1994 by Chinese paleontologists Cai Zhengquan and Wei Feng. The type species is ''Zhejiangopterus linhaiensis''. The genus name refers to Zhejiang Province and a Latinized Greek ''pteron'', "wing". The specific name refers to the city of Linhai. Discovery In 1986 a young chalkstone quarry worker named Xu Chengfa, found a large fossil near the village of Aolicun in Linhai. Xu by letter informed the director of the Zhejiang Museum of Natural History at Hangzhou, Ming Hua, who from it understood the remains were those of an unknown pterosaur. Therefore, he sent a team consisting of the describers and Wu Weitang to investigate. They secured the fossil, instructing the local population to be alert for possible further finds. Xu himself managed to find three more specimens before being killed in an accident in 1988; another ...
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Wellnhopterus
''Wellnhopterus'' () is an Azhdarchidae, azhdarchid pterosaur recovered from the Late Cretaceous Javelina Formation in Texas that was previously identified as a thalassodrominae, thalassodromine. It consists of a set of upper and lower jaws (TMM 42489-2), as well as some cervical vertebrae (TMM 42489-2.3 to 42489-2.8) and a fragmentary long bone (TMM 42489-2.9). In July 2021, the jaws were given the genus name "''Javelinadactylus''" (meaning "Javelina Finger"), with the type species, type and only species as "''J. sagebieli''"; however, this article has now been Retractions in academic publishing, retracted. In a paper published in December 2021, the complete holotype was independently named ''Wellnhopterus'', with the only species being ''W. brevirostris''. As of 2022, this is the formal name of this pterosaur. Discovery and naming In March 1986, jaws and neck vertebrae of a large pterosaur were discovered in Brewster County by preparator Robert Rainey. In 1991, Peter Wellnhofer ...
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Thanatosdrakon
''Thanatosdrakon'' () (meaning "dragon of death") is a genus of quetzalcoatline azhdarchid pterosaur from the Late Cretaceous (Upper Coniacian–Lower Santonian) Plottier Formation of the Neuquén Basin in western Argentina (Andes mountain range). The genus name is derived from the Greek words ''thanatos'' (=death) and ''drakon'' (=dragon), while the specific name is a Quechuan word meaning "flying serpent" and refers to the Incan deity Amaru. The type and only species is ''Thanatosdrakon amaru'', known from two specimens consisting of several well-preserved axial and appendicular bones including material previously undescribed in giant azhdarchids (e.g. complete notarium, dorsosacral vertebrae and caudal vertebra). ''Thanatosdrakon'' is one of the oldest known members of the Quetzalcoatlinae. ''T. amaru'' lived from about 90 to 86 million years ago. Description ''Thanatosdrakon'' is known from two well-preserved specimens first described in 2018. The holotype, UNCUYO-LD 307 ...
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Quetzalcoatlus
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas ...
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Phosphatodraco
''Phosphatodraco'' is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species ''Phosphatodraco mauritanicus'' in 2003; the genus name means " dragon from the phosphates", and the specific name refers to the region of Mauretania. ''Phosphatodraco'' was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur '' Tethydraco'' represent wing elements of ''Phosphatodraco''. Due to the fragmentary nature of the holotype cervical vertebrae, there has been co ...
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Hatzegopteryx
''Hatzegopteryx'' ("Hațeg basin wing") is a genus of azhdarchid pterosaur found in the late Maastrichtian deposits of the Densuş Ciula Formation, an outcropping in Transylvania, Romania. It is known only from the type species, ''Hatzegopteryx thambema'', named by Buffetaut ''et al.'' in 2002 based on parts of the skull and humerus. Additional specimens, including a neck vertebra, were later placed in the genus, representing a range of sizes. The largest of these remains indicate it was among the biggest pterosaurs, with an estimated wingspan of . Unusually among giant azhdarchids, ''Hatzegopteryx'' had a very wide skull bearing large muscular attachments; bones with a spongy internal texture instead of hollow; and a short, robust, and heavily muscled neck measuring long, which was about half the length of other azhdarchids with comparable wingspans, and was capable of withstanding strong bending forces. ''Hatzegopteryx'' inhabited Hațeg Island, an island situated in the Cret ...
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