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Austriadraconidae
Caviramidae is a group of basal pterosaurs. It was erected by paleontologist Matthew G. Baron in 2020. It was defined as the least inclusive clade that includes ''Arcticodactylus cromptonellus'' and '' Caviramus schesaplanensis''. The members of Caviramidae are also considered to belong to either the family Eudimorphodontidae (''Arcticodactylus'' and '' Carniadactylus''), the family Raeticodactylidae (''Caviramus'' and ''Raeticodactylus''), or just basal eopterosaurians ('' Austriadraco'' and ''Seazzadactylus''). However, Baron, the author of Caviramidae, concluded in his analyses that both Eopterosauria and Eudimorphodontidae are not monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ... groups, therefore he created the family Caviramidae to contain most of the eudim ...
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Arcticodactylus
''Arcticodactylus'' is a genus of basal pterosaur living during the Late Triassic in the area of present Greenland. Its only species was previously attributed to ''Eudimorphodon'', and its closest relatives may have been ''Eudimorphodon'' or ''Austriadraco''. History of discovery In 1989, William Amaral on the McKnight Bjerg in the east of Greenland discovered a rich fossil site. It was excavated in 1991 and 1992. Part of the material was a small skeleton of a pterosaur. In 2001, Farish Jenkins, Neil Shubin, Stephen Gatesy and Kevin Padian named and described it as a new species of ''Eudimorphodon'': ''Eudimorphodon cromptonellus''. The specific name honors Professor Alfred Walter Crompton. The suffix ''~ellus'', in Latin indicating a diminutive, alluded to the small size of the specimen. The holotype, MGUH VP 3393, was found in the Carlsberg Fjord Beds of the Ørsted Dal Member of the Fleming Fjord Formation dating from the Norian – Rhaetian. It consists of a partial skel ...
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Austriadraco
''Austriadraco'' is a genus of pterosaur living during the Late Triassic in the area of present Austria. Its only species—''Austriadraco dallavecchiai''—was previously attributed to ''Eudimorphodon'', and its closest relatives may have been ''Eudimorphodon'' or ''Arcticodactylus''. Discovery In June 1994, near Seefeld in Austrian Tirol, at a 1600 metres high mountain trail to the ''Reither Spitze'', in the vicinity of the ''Reither Joch-Alm'', Bernd Lammerer discovered a pterosaur skeleton. The remains have been secured as five stone plates, removed on several occasions. In 2003, Peter Wellnhofer identified the fossil as a specimen of ''Eudimorphodon'', a cf. ''E. ranzii''. As it was 10 to 25% shorter than the latter's holotype, Wellnhofer considered it a juvenile. The same year Fabio Marco Dalla Vecchia doubted the comparability to ''E. ranzii'' and suggested that it represent a separate ''Eudimorphodon'' species. In 2009, Dalla Vecchia concluded that the specimen was neith ...
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Arcticodactylus Cromptonellus
''Arcticodactylus'' is a genus of basal pterosaur living during the Late Triassic in the area of present Greenland. Its only species was previously attributed to ''Eudimorphodon'', and its closest relatives may have been ''Eudimorphodon'' or ''Austriadraco''. History of discovery In 1989, William Amaral on the McKnight Bjerg in the east of Greenland discovered a rich fossil site. It was excavated in 1991 and 1992. Part of the material was a small skeleton of a pterosaur. In 2001, Farish Jenkins, Neil Shubin, Stephen Gatesy and Kevin Padian named and described it as a new species of ''Eudimorphodon'': ''Eudimorphodon cromptonellus''. The specific name honors Professor Alfred Walter Crompton. The suffix ''~ellus'', in Latin indicating a diminutive, alluded to the small size of the specimen. The holotype, MGUH VP 3393, was found in the Carlsberg Fjord Beds of the Ørsted Dal Member of the Fleming Fjord Formation dating from the Norian – Rhaetian. It consists of a partial skel ...
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Caviramus Schesaplanensis
''Caviramus'' is a genus of caviramid pterosaur from the Late Triassic (early Rhaetian-age) lower Kössen Formation of the Northern Calcareous Alps of Switzerland. The genus was in 2006 named by Nadia Fröbisch and Jörg Fröbisch. The type species is ''Caviramus schesaplanensis''. The genus name is derived from Latin ''cavus'', "hollow" and ''ramus'', "branch". The specific name refers to Mount Schesaplana. Description The genus is based on holotype PIMUZ A/III 1225, three non-contiguous fragments of a ramus (lower jaw) of the mandible with multicuspate teeth. Two teeth are preserved, one with three cusps, and one with four; despite this difference the authors consider them as essentially isodont. The number of teeth is estimated at a minimum of twelve and a maximum of seventeen. A row of large oval foramina runs parallel to the tooth row; foramina in the form of small holes in the anterior part of the lower jaw suggest some sort of soft-tissue structure, or a keratin cove ...
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Carniadactylus
''Carniadactylus'' is a genus of pterosaur which existed in Europe during the Late Triassic period (late Carnian or early Norian, about 228 million years agoBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref>). The genus contains a single species, ''Carniadactylus rosenfeldi''. Description ''Carniadactylus'' was similar in appearance and anatomy to its close relative ''Eudimorphodon'', though it was significantly smaller. Like ''Eudimorphodon'', it is notable for its complex multi-cusped teeth. Despite their similarities, the size difference between these two pterosaurs likely meant that they occupied different niches and relied on different food sources. This is supported by studies of their teeth. While similar in construction, the teeth of ''Carniadactylus'' show little to no wear, unlike the larger, fish-eating ''Eudimorphodon'', which may have been able to chew its food. The sma ...
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Caviramus
''Caviramus'' is a genus of caviramid pterosaur from the Late Triassic (early Rhaetian-age) lower Kössen Formation of the Northern Calcareous Alps of Switzerland. The genus was in 2006 named by Nadia Fröbisch and Jörg Fröbisch. The type species is ''Caviramus schesaplanensis''. The genus name is derived from Latin ''cavus'', "hollow" and ''ramus'', "branch". The specific name refers to Mount Schesaplana. Description The genus is based on holotype PIMUZ A/III 1225, three non-contiguous fragments of a ramus (lower jaw) of the mandible with multicuspate teeth. Two teeth are preserved, one with three cusps, and one with four; despite this difference the authors consider them as essentially isodont. The number of teeth is estimated at a minimum of twelve and a maximum of seventeen. A row of large oval foramina runs parallel to the tooth row; foramina in the form of small holes in the anterior part of the lower jaw suggest some sort of soft-tissue structure, or a keratin cove ...
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Raeticodactylus
''Raeticodactylus'' is a genus of non- pterodactyloid pterosaur from the late Norian-early Rhaetian-age Upper Triassic lower Kössen Formation of the central Austroalpine of Grisons, Switzerland. It is known from holotype BNM 14524, a single disarticulated partial skeleton including an almost complete skull, found in August 2005. This genus was named and described in 2008 by its discoverer Rico Stecher; the type species is ''Raeticodactylus filisurensis''. The specific name refers to Filisur. Online First Description ''Raeticodactylus'' had a tall thin bony crest running along the midline of the front of the upper jaw, and a keel on the lower jaw; however, it does not seem to be closely related to '' Austriadactylus'', the only other crested Triassic pterosaur named by the time ''Raeticodactylus'' was described. The teeth at the front of the upper jaw, in the premaxillae, were fanglike, whereas the teeth in the upper cheeks (the maxillae) had three, four, or five cusps, simila ...
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Eopterosauria
Eopterosauria is a group of basal pterosaurs from the Triassic, which form their own clade. The term was first used in Andres ''et al.'' (2014) to include ''Preondactylus'', ''Austriadactylus'', ''Peteinosaurus'' and Eudimorphodontidae. Inside the group were two other new clades, Preondactylia, which included ''Preondactylus'' and ''Austriadactylus'', and Eudimorphodontoidea, to include Eudimorphodontidae and Raeticodactylidae. Eopterosauria was defined as "the least inclusive clade containing ''Preondactylus buffarinii'' and ''Eudimorphodon ranzii''". The specimen BSP 1994, previously assigned to ''Eudimorphodon'', was named the separate taxon ''Austriadraco'' in 2015, and assigned to the new family Austriadraconidae, but further classification was not described. The following phylogenetic analysis follows the topology of Andres ''et al.'' (2014). In a 2020 study of early pterosaur interrelationships carried out by Matthew G. Baron, no evidence was found to support the existen ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Seazzadactylus
''Seazzadactylus'' is a basal pterosaur genus that during the late Triassic lived in the area of present Italy. Discovery In 1997, amateur paleontologist Umberto Venier discovered the skeleton of a pterosaur in a boulder laying in the bed of the Seazza brook, just before it joins the Tagliamento river, near Preone in the Dolomites. Venier brought the find to the ''Museo Friulano di Storia Naturale'' at Udine. After partial preparation paleontologist Fabio Marco Dalla Vecchia announced the discovery in the scientific literature in 2000. In 2003, Dalla Vecchia referred the specimen to ''Eudimorphodon''. In 2009 however, further preparation made him conclude that it was a species new to science, not identical to either ''Eudimorphodon'' or ''Carniadactylus''. In 2019, Dalla Vecchia named and described the type species ''Seazzadactylus venieri''. The generic name combines that of the Seazza with the Greek , 'finger'. The specific name (zoology), specific name honours Venier as disco ...
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2020 In Paleontology
Plants Sponges Cnidarians New taxa Research * Revision of tabulate-like fossils from before the latest Middle Ordovician is published by Elias, Lee & Pratt (2020), who reject the interpretation of these fossils as true tabulate corals. * Drake, Whitelegge & Jacobs (2020) report the first recovery, sequencing, and identification of fossil biomineral proteins from a Pleistocene fossil invertebrate (the stony coral ''Orbicella annularis''). Arthropods Bryozoans Brachiopods New taxa Research * A study on the mode of life of Paleozoic strophomenatans is published by Stanley (2020), who argues that nearly all strophomenatans lived infaunally. * A study on the paleobiogeography of Early−Middle Devonian (Pragian−Eifelian) brachiopods from West Gondwana, aiming to determine any potential controls that may have driven bioregionalization, is published by Penn-Clarke & Harper (2020). * A study on the phylogenetic relationships and ecomorphologic diversification of Mesozoi ...
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