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Carniadactylus
''Carniadactylus'' is a genus of pterosaur which existed in Europe during the Late Triassic period (late Carnian or early Norian, about 228 million years agoBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref>). The genus contains a single species, ''Carniadactylus rosenfeldi''. Description ''Carniadactylus'' was similar in appearance and anatomy to its close relative ''Eudimorphodon'', though it was significantly smaller. Like ''Eudimorphodon'', it is notable for its complex multi-cusped teeth. Despite their similarities, the size difference between these two pterosaurs likely meant that they occupied different niches and relied on different food sources. This is supported by studies of their teeth. While similar in construction, the teeth of ''Carniadactylus'' show little to no wear, unlike the larger, fish-eating ''Eudimorphodon'', which may have been able to chew its food. The sma ...
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Bergamodactylus
''Bergamodactylus'' is a putative genus of basal pterosaur which lived during the Late Triassic (early Norian) in the area of present-day Bergamo province in Italy. Its only species is ''Bergamodactylus wildi''. It was named in 2015 based on a pterosaur specimen which had previously been regarded as a juvenile '' Eudimorphodon'' or as identical to '' Carniadactylus''. Some Triassic pterosaur specialists consider the distinguishing features of ''Bergamodactylus'' to be invalid or insufficient to distinguish it from ''Carniadactylus'', and thus retain the specimen in that genus. History In 1978, Rupert Wild described a small pterosaur specimen in the collection of the '' Museo di Paleontologia dell´Università di Milano'', found near Cene, Lombardy. He referred to it as the "Milan Exemplar" and identified it as a juvenile of '' Eudimorphodon ranzii''. Wild noted considerable differences with the latter's type specimen but these were explained as reflecting the young age of th ...
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Caviramus
''Caviramus'' is a genus of caviramid pterosaur from the Late Triassic (early Rhaetian-age) lower Kössen Formation of the Northern Calcareous Alps of Switzerland. The genus was in 2006 named by Nadia Fröbisch and Jörg Fröbisch. The type species is ''Caviramus schesaplanensis''. The genus name is derived from Latin ''cavus'', "hollow" and ''ramus'', "branch". The specific name refers to Mount Schesaplana. Description The genus is based on holotype PIMUZ A/III 1225, three non-contiguous fragments of a ramus (lower jaw) of the mandible with multicuspate teeth. Two teeth are preserved, one with three cusps, and one with four; despite this difference the authors consider them as essentially isodont. The number of teeth is estimated at a minimum of twelve and a maximum of seventeen. A row of large oval foramina runs parallel to the tooth row; foramina in the form of small holes in the anterior part of the lower jaw suggest some sort of soft-tissue structure, or a keratin cove ...
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Eudimorphodon
''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (mid to late Norian stage),Wellnhofer, P. (1991). "Summary of Triassic Pterosaurs." ''The Illustrated Encyclopedia of Pterosaurs.'' London, UK: Salamander Books Limited. p. 67. . making ''Eudimorphodon'' one of the oldest pterosaurs known.Cranfield, I. ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 280–281. It had a wingspan of about and at the end of its long bony tail may have been a diamond-shaped flap like in the later ''Rhamphorhynchus''. If so, the flap may have helped it steer while maneuvering in the air. ''Eudimorphodon'' is known from several skeletons, including juvenile specimens. Discovery and species ''Eudimorphodon'' currently includes one species, the type sp ...
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Preondactylus Buffarinii
''Preondactylus'' is a genus of long-tailed pterosaurs from the Late Triassic (late Norian)Barrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref> that inhabited what is now Italy. It contains a single known species, ''Preondactylus buffarinii'', which was discovered by Nando Buffarini in 1982 at the Forni Dolostone near Udine in the Preone valley of the Italian Alps."Preondactylus." In: Cranfield, Ingrid (ed.). ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 284-285. Discovery When Buffarini first discovered ''Preondactylus'', the thin slab of bituminous, dolomitic limestone containing the fossil was accidentally broken into pieces while being extracted. After reassembly the rock was cleaned with water by him and his wife and the marl and in it the bone was washed away and lost. All that was left was a negative imprint on ...
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Peteinosaurus
''Peteinosaurus'' ( ; meaning "winged lizard") was a prehistoric genus of pterosaur. It lived in the late Triassic period in the late Norian age (about 221 to 210 million years ago), and at a wingspan of around , was one of the smallest and earliest pterosaurs, although other estimates suggest a wingspan of up to . Discovery Three fossils have been found near Cene, Italy. The first fossil, the holotype MCSNB 2886, is fragmentary and disarticulated. The second, the articulated paratype MCSNB 3359, lacks any diagnostic features of ''Peteinosaurus'' and thus might be a different species. This paratype has a long tail (20 cm) made more stiff by long extensions of the vertebrae; this feature is common among pterosaurs of the Triassic. The third example is MCSNB 3496, another fragmentary skeleton. All specimens are those of subadults and of none has the skull been preserved. Like most pterosaurs, ''Peteinosaurus'' had bones that were strong but very light. ''Peteinosaurus'' is t ...
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Raeticodactylidae
Raeticodactylidae is a family of eudimorphodontoid eopterosaurian pterosaurs that lived in Switzerland during the Late Triassic. The family includes ''Caviramus'', and the type genus ''Raeticodactylus'', which are both known from the Kössen Formation, around 205 mya. Raeticodactylidae was first used in 2014 by Andres ''et al.'', as a group of all pterosaurs closer to ''Raeticodactylus'' than ''Eudimorphodon''. The following phylogenetic analysis follows the topology of Andres ''et al.'' (2014). In 2020 however, a study upheld by Matthew G. Baron about early pterosaur interrelationships found no evidence to support the existence of the clade Eopterosauria (the clade of which raeticodactylids might also belong to) as an early diverging clade within the Pterosauria, therefore, he sunk both ''Caviramus'' and ''Raeticodactylus'' within a clade he called Caviramidae Caviramidae is a group of basal pterosaurs. It was erected by paleontologist Matthew G. Baron in 2020. It was defin ...
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Austriadactylus Cristatus
''Austriadactylus'' is a genus of "rhamphorhynchoid" pterosaur. The fossil remains were unearthed in Late Triassic (middle Norian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref>) rocks of Austria. The genus was named in 2002 by Fabio Marco Dalla Vecchia e.a.. The type species is ''Austriadactylus cristatus''. The genus name is derived from Latin ''Austria'' and Greek ''daktylos'', "finger", in reference to the wing finger of pterosaurs. The specific epithet means "crested" in Latin, a reference to the skull crest. The genus is based on holotype SMNS 56342, a crushed partial skeleton on a slab, found in an abandoned mine near Ankerschlag in Tyrol, in the Norian Seefelder Beds. The counterslab has been lost and with it some of the bone. The fossil consists of the skull, lower jaws, some vertebrae, parts of the limbs and pelvic girdle, and the first part of the tail. The elo ...
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Peteinosaurus Zambellii
''Peteinosaurus'' ( ; meaning "winged lizard") was a prehistoric genus of pterosaur. It lived in the late Triassic period in the late Norian age (about 221 to 210 million years ago), and at a wingspan of around , was one of the smallest and earliest pterosaurs, although other estimates suggest a wingspan of up to . Discovery Three fossils have been found near Cene, Italy. The first fossil, the holotype MCSNB 2886, is fragmentary and disarticulated. The second, the articulated paratype MCSNB 3359, lacks any diagnostic features of ''Peteinosaurus'' and thus might be a different species. This paratype has a long tail (20 cm) made more stiff by long extensions of the vertebrae; this feature is common among pterosaurs of the Triassic. The third example is MCSNB 3496, another fragmentary skeleton. All specimens are those of subadults and of none has the skull been preserved. Like most pterosaurs, ''Peteinosaurus'' had bones that were strong but very light. ''Peteinosaurus'' is t ...
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Carnian
The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by the Norian. Its boundaries are not characterized by major extinctions or biotic turnovers, but a climatic event (known as the Carnian pluvial episode characterized by substantial rainfall) occurred during the Carnian and seems to be associated with important extinctions or biotic radiations. Stratigraphic definitions The Carnian was named in 1869 by Mojsisovics. It is unclear if it was named after the Carnic Alps or after the Austrian region of Carinthia (''Kärnten'' in German) or after the Carnia historical region in northwestern Italy. The name, however, was first used referring to a part of the Hallstatt Limestone cropping out in Austria. The base of the Carnian Stage is defined as the place in the stratigraphic record where t ...
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Eudimorphodontoidea
Eopterosauria is a group of basal pterosaurs from the Triassic, which form their own clade. The term was first used in Andres ''et al.'' (2014) to include ''Preondactylus'', ''Austriadactylus'', ''Peteinosaurus'' and Eudimorphodontidae. Inside the group were two other new clades, Preondactylia, which included ''Preondactylus'' and ''Austriadactylus'', and Eudimorphodontoidea, to include Eudimorphodontidae and Raeticodactylidae. Eopterosauria was defined as "the least inclusive clade containing ''Preondactylus buffarinii'' and ''Eudimorphodon ranzii''". The specimen BSP 1994, previously assigned to ''Eudimorphodon'', was named the separate taxon ''Austriadraco'' in 2015, and assigned to the new family Austriadraconidae, but further classification was not described. The following phylogenetic analysis follows the topology of Andres ''et al.'' (2014). In a 2020 study of early pterosaur interrelationships carried out by Matthew G. Baron, no evidence was found to support the existenc ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Eopterosauria
Eopterosauria is a group of basal pterosaurs from the Triassic, which form their own clade. The term was first used in Andres ''et al.'' (2014) to include ''Preondactylus'', ''Austriadactylus'', ''Peteinosaurus'' and Eudimorphodontidae. Inside the group were two other new clades, Preondactylia, which included ''Preondactylus'' and ''Austriadactylus'', and Eudimorphodontoidea, to include Eudimorphodontidae and Raeticodactylidae. Eopterosauria was defined as "the least inclusive clade containing ''Preondactylus buffarinii'' and ''Eudimorphodon ranzii''". The specimen BSP 1994, previously assigned to ''Eudimorphodon'', was named the separate taxon ''Austriadraco'' in 2015, and assigned to the new family Austriadraconidae, but further classification was not described. The following phylogenetic analysis follows the topology of Andres ''et al.'' (2014). In a 2020 study of early pterosaur interrelationships carried out by Matthew G. Baron, no evidence was found to support the existen ...
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