Atlantogenata
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Atlantogenata
Atlantogenata is a proposed clade of mammals containing the cohorts or superorders Xenarthra and Afrotheria. These groups originated and radiated in the South American and African continents, respectively, presumably in the Cretaceous. Together with Boreoeutheria, they make up Eutheria. The monophyly of this grouping was supported by some genetic evidence. Alternative hypotheses are that Boreoeutheria and Afrotheria combine to form Epitheria (as generally supported by anatomical and other physiological evidence) or that Boreoeutheria and Xenarthra combine to form Exafroplacentalia or Notolegia. Updated analysis of transposable element A transposable element (TE, transposon, or jumping gene) is a nucleic acid sequence in DNA that can change its position within a genome, sometimes creating or reversing mutations and altering the cell's genetic identity and genome size. Transp ... insertions around the time of divergence strongly supports the fourth hypothesis of a near-conco ...
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Epitheria
Epitherians comprise all the placental mammals except the Xenarthra. They are primarily characterized by having a stirrup-shaped stapes in the middle ear, which allows for passage of a blood vessel. This is in contrast to the column-shaped stapes found in marsupials, monotremes, and xenarthrans. They are also characterized by having a shorter fibula relative to the tibia. Epitheria like Xenarthra and Afrotheria originated after the K-Pg boundary , with the placental diversification occurring within the first hundred thousand years after the K-Pg event and the first modern placental orders began appearing 2–3 million years later. Epitheres are one of the most successful groups of animals. The monophyly of Epitheria has been challenged by molecular phylogenetic studies. While preliminary analysis of a set of retroposons shared by both Afrotheria, and Boreoeutheria ( presence/absence data) supported the Epitheria clade, more extensive analysis of such transposable element insert ...
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Exafroplacentalia
Exafroplacentalia or Notolegia is a clade of placental mammals proposed in 2001 on the basis of molecular research. Exafroplacentalia places Xenarthra as a sister group to the Boreoeutheria (comprising Laurasiatheria and Euarchontoglires), thus making Afrotheria a primitive group of placental mammals (the group name roughly means "those which are not African placentals"). Classification However, this classification makes the autapomorphy (character shared only among Exafroplacentalia) dubious: it is hard to classify a group by the absence of a feature (in this case "not coming from Africa").Nishihara, H., Maruyama, S. & Okada, N. 2009. Retroposon analysis and recent geological data suggest near-simultaneous divergence of the three superorders of placental mammals. PNAS 106: 5235-40. Hence, several alternative hypotheses can be considered. Alternative hypotheses One alternative hypothesis is the Epitheria hypothesis: Another alternative hypothesis is the Atlantogenata hypo ...
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Notolegia
Exafroplacentalia or Notolegia is a clade of placental mammals proposed in 2001 on the basis of molecular research. Exafroplacentalia places Xenarthra as a sister group to the Boreoeutheria (comprising Laurasiatheria and Euarchontoglires), thus making Afrotheria a primitive group of placental mammals (the group name roughly means "those which are not African placentals"). Classification However, this classification makes the autapomorphy (character shared only among Exafroplacentalia) dubious: it is hard to classify a group by the absence of a feature (in this case "not coming from Africa").Nishihara, H., Maruyama, S. & Okada, N. 2009. Retroposon analysis and recent geological data suggest near-simultaneous divergence of the three superorders of placental mammals. PNAS 106: 5235-40. Hence, several alternative hypotheses can be considered. Alternative hypotheses One alternative hypothesis is the Epitheria hypothesis: Another alternative hypothesis is the Atlantogenata hypo ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Eutheria
Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic traits of the feet, ankles, jaws and teeth. All extant eutherians lack epipubic bones, which are present in all other living mammals (marsupials and monotremes). This allows for expansion of the abdomen during pregnancy. The oldest-known eutherian species is '' Juramaia sinensis'', dated at from the early Late Jurassic ( Oxfordian) of China. Eutheria was named in 1872 by Theodore Gill; in 1880 Thomas Henry Huxley defined it to encompass a more broadly defined group than Placentalia. Characteristics Distinguishing features are: *an enlarged malleolus ("little hammer") at the bottom of the tibia, the larger of the two shin bones *the joint between the first metatarsal bone and the entocuneiform bone (the innermost of the three cuneiform ...
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Xenarthra
Xenarthra (; from Ancient Greek ξένος, xénos, "foreign, alien" + ἄρθρον, árthron, "joint") is a major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Extinct xenarthrans include the glyptodonts, pampatheres and ground sloths. Xenarthrans originated in South America during the late Paleocene about 60 million years ago. They evolved and diversified extensively in South America during the continent's long period of isolation in the early to mid Cenozoic Era. They spread to the Antilles by the early Miocene and, starting about 3 Mya, spread to Central and North America as part of the Great American Interchange. Nearly all of the formerly abundant megafaunal xenarthrans became extinct at the end of the Pleistocene. Xenarthrans share several characteristics not present in other placental mammals, which suggest their ancestors were subterranean diggers for insects. The name Xenarthra de ...
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Pilosa
The order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes the anteaters and sloths (which includes the extinct ground sloths). The name comes from the Latin word for "hairy". Origins and taxonomy The biogeographic origins of the Pilosa are still unclear, but they can be traced back in South America as far as the early Paleogene (about 60 million years ago, only a short time after the end of the Mesozoic Era). The presence of these animals in Central America and their former presence in North America is a result of the Great American Interchange. A number of sloths were also formerly present on the Antilles, which they reached from South America by some combination of rafting or floating with the prevailing currents. Together with the armadillos, which are in the order Cingulata, pilosans are part of the larger superorder Xenarthra, a defining characteristic of which is the presence of xenarthrals (extra formations between lumbar verte ...
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Boreoeutheria
Boreoeutheria (, "northern true beasts") is a magnorder of placental mammals that groups together superorders Euarchontoglires and Laurasiatheria. With a few exceptionsExceptional clades whose males lack the usual boreoeutherian scrotum are moles, hedgehogs, pangolins, some pinnipeds, rhinoceroses, tapirs, hippopotamuses, and cetaceans. male animals in the clade have a scrotum, an ancestral feature of the clade. The sub-clade Scrotifera was named after this feature. Etymology The name of this magnorder comes from Ancient Greek words: * () meaning ' north wind' or 'the North', * () meaning 'good', 'right', or 'true', * and () meaning 'beast'. Boreoeutherian ancestor The majority of earliest known fossils belonging to this group date to about 66 million years ago, shortly after the K-Pg extinction event, though molecular data suggest they may have originated earlier, during the Cretaceous period. This is further supported with fossils of '' Altacreodus magnus'' and two ...
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Laurasiatheria
Laurasiatheria ("laurasian beasts") is a superorder of placental mammals that groups together true insectivores ( eulipotyphlans), bats ( chiropterans), carnivorans, pangolins ( pholidotes), even-toed ungulates (artiodactyls), odd-toed ungulates (perissodactyls), and all their extinct relatives. From systematics and phylogenetic perspectives, it is subdivided into order Eulipotyphla and clade Scrotifera. It is a sister group to Euarchontoglires with which it forms the magnorder Boreoeutheria. Laurasiatheria was discovered on the basis of the similar gene sequences shared by the mammals belonging to it; no anatomical features have yet been found that unite the group. The Laurasiatheria clade is based on DNA sequence analyses and retrotransposon presence/absence data. The superorder originated on the northern supercontinent of Laurasia, after it split from Gondwana when Pangaea broke up. Its last common ancestor is supposed to have diversified ca. 76 to 90 million years ago. Ety ...
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Three-toed Sloth
The three-toed or three-fingered sloths are arboreal neotropical mammals . They are the only members of the genus ''Bradypus'' and the family Bradypodidae. The four living species of three-toed sloths are the brown-throated sloth, the maned sloth, the pale-throated sloth, and the pygmy three-toed sloth. In complete contrast to past morphological studies, which tended to place ''Bradypus'' as the sister group to all other folivorans, molecular studies place them nested within the sloth superfamily Megatherioidea, making them the only surviving members of that radiation. Extant species Evolution A study of mitochondrial cytochrome b and 16S rRNA sequences suggests that '' B. torquatus'' diverged from '' B. variegatus'' and '' B. tridactylus'' about 12 million years ago, while the latter two split 5 to 6 million years ago. The diversification of ''B. variegatus'' lineages was estimated to have started 4 to 5 million years ago. Relation to the two-toed sloth Both types of sloth t ...
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Sloth
Sloths are a group of Neotropical xenarthran mammals constituting the suborder Folivora, including the extant arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of movement, tree sloths spend most of their lives hanging upside down in the trees of the tropical rainforests of South America and Central America. Sloths are considered to be most closely related to anteaters, together making up the xenarthran order Pilosa. There are six extant sloth species in two genera – '' Bradypus'' (three–toed sloths) and '' Choloepus'' (two–toed sloths). Despite this traditional naming, all sloths have three toes on each rear limb-- although two-toed sloths have only two digits on each forelimb. The two groups of sloths are from different, distantly related families, and are thought to have evolved their morphology via parallel evolution from terrestrial ancestors. Besides the extant species, many species of ground sloths ranging up to the size of ele ...
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Myrmecophagidae
The Myrmecophagidae are a family of anteaters, the name being derived from the Ancient Greek words for 'ant' and 'eat' (''myrmeco-'' and '). Two genera and three species are in the family, consisting of the giant anteater, and the tamanduas. The fossil ''Eurotamandua'' from the Messel Pit in Germany may be an early anteater, but its status is currently debated. Characteristics Myrmecophagids are medium to large animals, with distinctively elongated snouts and long, narrow tongues. They have powerful claws on their toes, enabling them to rip open termite mounds and ant nests to eat the insects inside. They have no teeth, but produce a large amount of sticky saliva to trap the insects, as well as backward-pointing spines on their tongues. Ants and termites are almost their only food in the wild, and their primary source of water, although they sometimes also drink free-standing water, and occasionally eat fruits. Distribution Myrmecophagids are found in Central and South Americ ...
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