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Asplenium Pinnatifidum
''Asplenium pinnatifidum'', commonly known as the lobed spleenwort or pinnatifid spleenwort, is a small fern found principally in the Appalachian Mountains and the Shawnee Hills, growing in rock crevices in moderately acid to subacid strata. Originally identified as a variety of walking fern ('' Asplenium rhizophyllum''), it was classified as a separate species by Thomas Nuttall in 1818. It is believed to have originated by chromosome doubling in a hybrid between walking fern and mountain spleenwort ('' Asplenium montanum''), producing a fertile tetraploid, a phenomenon known as alloploidy; however, the hypothesized parental hybrid has never been located. It is intermediate in morphology between the parent species: while its leaf blades are long and tapering like that of walking fern, the influence of mountain spleenwort means that the blades are lobed, rather than whole. ''A. pinnatifidum'' can itself form sterile hybrids with several other spleenworts. Description ''Asple ...
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Schist
Schist ( ) is a medium-grained metamorphic rock showing pronounced schistosity. This means that the rock is composed of mineral grains easily seen with a low-power hand lens, oriented in such a way that the rock is easily split into thin flakes or plates. This texture (geology), texture reflects a high content of platy minerals, such as micas, talc, chlorite group, chlorite, or graphite. These are often interleaved with more granular minerals, such as feldspar or quartz. Schist typically forms during regional metamorphism accompanying the process of mountain building (orogeny) and usually reflects a medium Metamorphism#Metamorphic grades, grade of metamorphism. Schist can form from many different kinds of rocks, including sedimentary rocks such as mudstones and igneous rocks such as tuffs. Schist metamorphosed from mudstone is particularly common and is often very rich in mica (a ''mica schist''). Where the type of the original rock (the protolith) is discernible, the schist is us ...
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Stipe (botany)
In botany, a stipe is a stalk that supports some other structure. The precise meaning is different depending on which taxonomic group is being described. file:Helicteres-Yucatán-Flowers.jpg, The long stipe of a '' Helicteres'' flower. file:Helicteres-Yucatán-Fruits.jpg, remains as each flower forms a fruit. In the case of ferns, the stipe is only the petiole from the rootstock to the beginning of the leaf tissue, or lamina. The continuation of the structure within the lamina is then termed a rachis. In flowering plants, the term is often used in reference to a stalk that sometimes supports a flower's ovary. In orchids, the stipe or caudicle is the stalk-like support of the pollinia. It is a non-viscid band or strap connecting the pollinia with the viscidium (the viscid part of the rostellum or beak). A stipe is also a structure found in organisms that are studied by botanists but that are no longer classified as plants. It may be the stem-like part of the thallus of a mus ...
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Asplenium Tutwilerae
''Asplenium tutwilerae'' (Tutwiler's spleenwort or Scott's fertile spleenwort) is a rare epipetric fern found only in Hale County, Alabama, United States. ''A. tutwilerae'' is a fertile allotetraploid, formed by the chromosomal doubling of a specimen of the sterile diploid ''A. × ebenoides'', a hybrid of ''A. platyneuron'' and ''A. rhizophyllum''. Except for its spores, which are fertile rather than malformed, ''A. tutwilerae'' is essentially identical to ''A. × ebenoides'' and was described as part of that species until 2007. It is named in honor of Julia Tutwiler, who discovered the only known wild population at Havana Glen in 1873. Description ''Asplenium tutwilerae'' is a small, compact, evergreen, rock-inhabiting fern that grows in individual clumps. It displays a slight frond dimorphism, with the larger, fertile leaf blades more or less upright, while the smaller, usually sterile blades are tightly pressed against the ground. ...
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Asplenium × Ebenoides
''Asplenium'' × ''ebenoides'' (Scott's spleenwort, dragon tail fern or walking spleenwort) is a hybrid fern native to eastern North America, part of the "Appalachian ''Asplenium'' complex" of related hybrids. The sterile offspring of the walking fern ('' A. rhizophyllum'') and the ebony spleenwort ('' A. platyneuron''), ''A.'' × ''ebenoides'' is intermediate in morphology between its two parents, combining the long, narrow blade of ''A. rhizophyllum'' with a dark stem and lobes or pinnae similar to those of ''A. platyneuron''. While ''A.'' × ''ebenoides'' is generally sterile, fertile specimens with double the number of chromosomes are known from Havana Glen, Alabama. These fertile allotetraploids were reclassified as a separate species named '' A. tutwilerae'' in 2007, retaining the name ''A.'' × ''ebenoides'' for the sterile diploids only. The hybrid nature of ''A.'' × ''ebenoides'' was suspected at t ...
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Asplenium Dalhousiae
''Asplenium'' is a genus of about 700 species of ferns, often treated as the only genus in the family Aspleniaceae, though other authors consider ''Hymenasplenium'' separate, based on molecular phylogenetic analysis of DNA sequences, a different chromosome count, and structural differences in the rhizomes. The type species for the genus is ''Asplenium marinum''. The most common vernacular name is spleenworts, applied to the more "typical" species. '' A. nidus'' and several similar species are called bird's-nest ferns, the ''Camptosorus'' group is known as walking ferns, and distinct names are applied to some other particularly well-known species. Taxonomy and genetics Many groups of species have been separated from ''Asplenium'' as segregate genera. These include ''Camptosorus'', ''Ceterach'', ''Phyllitis'', and ''Tarachia'', but these species can form hybrids with other ''Asplenium'' species and because of this are usually included in a more broadly defined ''Asplenium''. ...
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Giant City State Park
Giant City State Park is an Illinois state park on in Jackson and Union Counties, Illinois, United States. Illinois acquired more than in 1927, and dedicated the park as Giant City State Park. A lodge and visitor center welcome state park guests. Giant City State Park is a haven for nature lovers. It is a popular destination in Southern Illinois, and visitors to the park enjoy a number of activities such as hiking, horseback riding, picnicking, and rock climbing. Some of the park's most popular hiking trails include Giant City Nature Trail, Red Cedar Trail, and Trillium Trail. The park is served by the River to River Trail. The park's lodge and original six cabins were constructed by the 696th Company of the Civilian Conservation Corps from 1934 to 1935. The CCC used locally quarried sandstone and locally harvested lumber to build the lodge. In 1936, the lodge and cabins were dedicated by Illinois Governor Henry Horner. The CCC later added a dining room and kitchen as well a ...
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Allotetraploid
Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains one or more chromosomes and comes from each of two parents, resulting in pairs of homologous chromosomes between sets. However, some organisms are polyploid. Polyploidy is especially common in plants. Most eukaryotes have diploid somatic cells, but produce haploid gametes (eggs and sperm) by meiosis. A monoploid has only one set of chromosomes, and the term is usually only applied to cells or organisms that are normally diploid. Males of bees and other Hymenoptera, for example, are monoploid. Unlike animals, plants and multicellular algae have life cycles with two alternating multicellular generations. The gametophyte generation is haploid, and produces gametes by mitosis, the sporophyte generation is diploid and produces spores by meiosis ...
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Sporophyte
A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote produced when a haploid egg cell is fertilized by a haploid sperm and each sporophyte cell therefore has a double set of chromosomes, one set from each parent. All land plants, and most multicellular algae, have life cycles in which a multicellular diploid sporophyte phase alternates with a multicellular haploid gametophyte phase. In the seed plants, the largest groups of which are the gymnosperms and flowering plants (angiosperms), the sporophyte phase is more prominent than the gametophyte, and is the familiar green plant with its roots, stem, leaves and cones or flowers. In flowering plants the gametophytes are very reduced in size, and are represented by the germinated pollen and the embryo sac. The sporophyte produces spores (hence t ...
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Indusium
A sorus (pl. sori) is a cluster of sporangia (structures producing and containing spores) in ferns and fungi. A coenosorus (plural coenosori) is a compound sorus composed of multiple, fused sori. Etymology This New Latin word is from Ancient Greek σωρός (''sōrós'' 'stack, pile, heap'). Structure In lichens and other fungi, the sorus is surrounded by an external layer. In some red algae, it may take the form of depression into the thallus. In ferns, the sori form a yellowish or brownish mass on the edge or underside of a fertile frond. In some species, they are protected during development by a scale or film of tissue called the indusium, which forms an umbrella-like cover. Lifecycle significance Sori occur on the sporophyte generation, the sporangia within producing haploid meio spores. As the sporangia mature, the indusium shrivels so that spore release is unimpeded. The sporangia then burst and release the spores. As an aid to identification The shape, arran ...
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Sorus
A sorus (pl. sori) is a cluster of sporangia (structures producing and containing spores) in ferns and fungi. A coenosorus (plural coenosori) is a compound sorus composed of multiple, fused sori. Etymology This New Latin word is from Ancient Greek σωρός (''sōrós'' 'stack, pile, heap'). Structure In lichens and other fungi, the sorus is surrounded by an external layer. In some red algae, it may take the form of depression into the thallus. In ferns, the sori form a yellowish or brownish mass on the edge or underside of a fertile frond. In some species, they are protected during development by a scale or film of tissue called the indusium, which forms an umbrella-like cover. Lifecycle significance Sori occur on the sporophyte generation, the sporangia within producing haploid meiospores. As the sporangia mature, the indusium shrivels so that spore release is unimpeded. The sporangia then burst and release the spores. As an aid to identification The shape, arrangemen ...
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Asplenium Pinnatifidum Sori
''Asplenium'' is a genus of about 700 species of ferns, often treated as the only genus in the family Aspleniaceae, though other authors consider ''Hymenasplenium'' separate, based on molecular phylogenetic analysis of DNA sequences, a different chromosome count, and structural differences in the rhizomes. The type species for the genus is ''Asplenium marinum''. The most common vernacular name is spleenworts, applied to the more "typical" species. '' A. nidus'' and several similar species are called bird's-nest ferns, the ''Camptosorus'' group is known as walking ferns, and distinct names are applied to some other particularly well-known species. Taxonomy and genetics Many groups of species have been separated from ''Asplenium'' as segregate genera. These include ''Camptosorus'', ''Ceterach'', ''Phyllitis'', and ''Tarachia'', but these species can form hybrids with other ''Asplenium'' species and because of this are usually included in a more broadly defined ''Asplenium''. ...
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Anastomosing
An anastomosis (, plural anastomoses) is a connection or opening between two things (especially cavities or passages) that are normally diverging or branching, such as between blood vessels, leaf veins, or streams. Such a connection may be normal (such as the foramen ovale in a fetus's heart) or abnormal (such as the patent foramen ovale in an adult's heart); it may be acquired (such as an arteriovenous fistula) or innate (such as the arteriovenous shunt of a metarteriole); and it may be natural (such as the aforementioned examples) or artificial (such as a surgical anastomosis). The reestablishment of an anastomosis that had become blocked is called a reanastomosis. Anastomoses that are abnormal, whether congenital or acquired, are often called fistulas. The term is used in medicine, biology, mycology, geology, and geography. Etymology Anastomosis: medical or Modern Latin, from Greek ἀναστόμωσις, anastomosis, "outlet, opening", Gr ana- "up, on, upon", stoma "mouth", ...
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