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Arachniotus Ruber
''Arachniotus ruber'' is a species of fungus belonging to the genus ''Arachniotus'' in the family '' Gymnoascaceae''. This fungus is a mesophile that reproduces both sexually and asexually. So far, there have been no reports of the fungus being pathogenic. History and taxonomy ''Arachniotus ruber'' was first described by Philippe Édouard Léon Van Tieghem in 1877 and it was thought to belong to the genus, ''Gymnoascus'', due to similarities he observed in the gametangial initials. It was not until 1893 that Schroeter transferred the species to his recently established genus, ''Arachniotus'', due to it possessing many of the corresponding traits. At the time, Schroeter did not designate a type species for the genus, which resulted in some debate until a lectotype, ''A. candidus'', was eventually identified by Clements and Shear in 1931. With this change, ''A. ruber'' was excluded from the ''Arachniotus'' genus because it did not display type characteristics within its ascospore ...
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Fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''t ...
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Ascomycota
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defining feature of this fungal group is the " ascus" (), a microscopic sexual structure in which nonmotile spores, called ascospores, are formed. However, some species of the Ascomycota are asexual, meaning that they do not have a sexual cycle and thus do not form asci or ascospores. Familiar examples of sac fungi include morels, truffles, brewers' and bakers' yeast, dead man's fingers, and cup fungi. The fungal symbionts in the majority of lichens (loosely termed "ascolichens") such as ''Cladonia'' belong to the Ascomycota. Ascomycota is a monophyletic group (it contains all descendants of one common ancestor). Previously placed in the Deuteromycota along with asexual species from other fungal taxa, asexual (or anamorphic) ascomyce ...
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Eurotiomycetes
Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. It contains most of the fungi previously known morphologically as "Plectomycetes".


Systematics and phylogeny


Internal relationships

The class Eurotiomycetes was circumscription (taxonomy), circumscribed in 1997 by Sweden, Swedish mycologists Ove Erik Eriksson and Katarina Winka. At that time ...

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Onygenales
The Onygenales are an order of fungi in the class Eurotiomycetes and division Ascomycota. The order's last common ancestor is estimated to have lived 150 million years ago. Onygenales can consume and break down keratin, the main component of the outer layer of skin. They are primarily found on animals, droppings, and areas frequented by animals. Many are dimorphic, and can change from mold to yeast form depending on their environment. Many onygenalean fungi are pathogens. One species, ''Trichophyton rubrum'', is the primary cause of athlete's foot. This order also includes Coccidioides implicated in Valley fever. The Onygenales are important as emerging human pathogens because of the rising rates of immunosuppression due to live-organ transplant, HIV/AIDS, and autoimmune disorders such as lupus erythematosus Lupus erythematosus is a collection of autoimmune diseases in which the human immune system becomes hyperactive and attacks healthy tissues. Symptoms of these disease ...
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Gymnoascaceae
The Gymnoascaceae are a family of fungi in the Ascomycota, class Eurotiomycetes Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichenized class, with more tha .... References Onygenales Ascomycota families {{Eurotiomycetes-stub ...
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Arachniotus
''Arachniotus'' is a genus of fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ... within the family Gymnoascaceae. Species *'' Arachniotus aurantiacus'' *'' Arachniotus candidus'' *'' Arachniotus ellipticosporus'' *'' Arachniotus flavoluteus'' *'' Arachniotus hebridensis'' *'' Arachniotus hyalotrichus'' *'' Arachniotus indicus'' *'' Arachniotus insolitus'' *'' Arachniotus lanatus'' *'' Arachniotus littoralis'' *'' Arachniotus punctatus'' *'' Arachniotus ruber'' *'' Arachniotus terrestris'' *'' Arachniotus trisporus'' *'' Arachniotus verruculosus'' References External links * Eurotiomycetes genera Onygenales Taxa named by Joseph Schröter Taxa described in 1893 {{Eurotiomycetes-stub ...
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Mesophile
A mesophile is an organism that grows best in moderate temperature, neither too hot nor too cold, with an optimum growth range from . The optimum growth temperature for these organisms is 37°C. The term is mainly applied to microorganisms. Organisms that prefer extreme environments are known as extremophiles. Mesophiles have diverse classifications, belonging to two domains: Bacteria, Archaea, and to kingdom Fungi of domain Eucarya. Mesophiles belonging to the domain Bacteria can either be gram-positive or gram-negative. Oxygen requirements for mesophiles can be aerobic or anaerobic. There are three basic shapes of mesophiles: coccus, bacillus, and spiral. Habitat The habitats of mesophiles can include cheese and yogurt. They are often included during fermentation of beer and wine making. Since normal human body temperature is 37 °C, the majority of human pathogens are mesophiles, as are most of the organisms comprising the human microbiome. Mesophiles vs. extremophiles Meso ...
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Philippe Édouard Léon Van Tieghem
Philippe Édouard Léon Van Tieghem (; 19 April 1839 – 28 April 1914) was a French botanist born in Baillleul in the département of Nord. He was one of the best known French botanists of the latter nineteenth century. Life Van Tieghem's father was a textile merchant who died of yellow fever in Martinique before he was born, and his mother shortly thereafter. One of five children, he obtained his ''baccalauréat'' in 1856, and continued his studies at the École Normale Supérieure, where after receiving agrégation, he worked in the laboratory of Louis Pasteur (1822–1895). Here he performed research involving the cultivation of mushrooms. He is credited with creation of the eponymous "Van Tieghem cell", a device mounted on a microscope slide that allows for observing the development of a fungus' mycelium. In 1864 he earned his doctorate in physical sciences with a thesis titled ''Recherches sur la fermentation de l'urée et de l'acide hippurique'', and two years later ob ...
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Gymnoascus
''Gymnoascus'' is a genus of fungi within the Gymnoascaceae family. The genus, widely distributed in northern temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ... areas, contains eight species. References External links * Eurotiomycetes genera Onygenales Taxa described in 1872 {{Eurotiomycetes-stub ...
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Gametangium
A gametangium (plural: gametangia) is an organ or cell in which gametes are produced that is found in many multicellular protists, algae, fungi, and the gametophytes of plants. In contrast to gametogenesis in animals, a gametangium is a haploid structure and formation of gametes does not involve meiosis. Types of gametangia Depending on the type of gamete produced in a gametangium, several types can be distinguished. Female Female gametangia are most commonly called archegonia. They produce egg cells and are the sites for fertilization. Archegonia are common in algae and primitive plants as well as gymnosperms. In flowering plants, they are replaced by the embryo sac inside the ovule. Male The male gametangia are most commonly called antheridia. They produce sperm cells that they release for fertilization. Antheridia producing non-motile sperm (spermatia) are called spermatangia. Some antheridia do not release their sperm. For example, the oomycete antheridium is a syncytium w ...
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Ascospore
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Ascocarp
An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of flecks ...
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