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Aquificales
The ''Aquificota'' phylum is a diverse collection of bacteria that live in harsh environmental settings. The name ''Aquificota'' was given to this phylum based on an early genus identified within this group, ''Aquifex'' (“water maker”), which is able to produce water by oxidizing hydrogen. They have been found in springs, pools, and oceans. They are autotrophs, and are the primary carbon fixers in their environments. These bacteria are Gram-negative, non-spore-forming rods. They are true bacteria (domain Bacteria) as opposed to the other inhabitants of extreme environments, the Archaea. Taxonomy The ''Aquificota'' currently contain 15 genera and 42 validly published species. The phylum comprises a single class and two orders. Aquificales consists of the families Aquificaceae and Hydrogenothermaceae, while the Desulfurobacteriaceae are the only family within the Desulfobacteriales. ''Thermosulfidibacter takaii'' is not assigned to a family within the phylum based on its phylog ...
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Aquificales
The ''Aquificota'' phylum is a diverse collection of bacteria that live in harsh environmental settings. The name ''Aquificota'' was given to this phylum based on an early genus identified within this group, ''Aquifex'' (“water maker”), which is able to produce water by oxidizing hydrogen. They have been found in springs, pools, and oceans. They are autotrophs, and are the primary carbon fixers in their environments. These bacteria are Gram-negative, non-spore-forming rods. They are true bacteria (domain Bacteria) as opposed to the other inhabitants of extreme environments, the Archaea. Taxonomy The ''Aquificota'' currently contain 15 genera and 42 validly published species. The phylum comprises a single class and two orders. Aquificales consists of the families Aquificaceae and Hydrogenothermaceae, while the Desulfurobacteriaceae are the only family within the Desulfobacteriales. ''Thermosulfidibacter takaii'' is not assigned to a family within the phylum based on its phylog ...
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Thermotogota
The Thermotogota are a phylum of the domain Bacteria. The phylum Thermotogota is composed of Gram-negative staining, anaerobic, and mostly thermophilic and hyperthermophilic bacteria.Gupta, RS (2014) The Phylum Thermotogae. The Prokaryotes 989-1015. Springer Berlin Heidelberg. Characteristics The name of this phylum is derived from the existence of many of these organisms at high temperatures along with the characteristic sheath structure, or "toga", surrounding the cells of these species.Reysenbach, A.-L. (2001) Phylum BII. Thermotogae phy. nov. In: Bergey's Manual of Systematic Bacteriology, pp. 369-387. Eds D. R. Boone, R. W. Castenholz. Springer-Verlag: Berlin. Recently, some Thermotogota existing at moderate temperatures have also been identified. Although Thermotogota species exhibit Gram-negative staining, they are bounded by a single-unit lipid membrane, hence they are monoderm bacteria. Because of the ability of some Thermotogota species to thrive at high temperatures, th ...
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Hydrogenothermaceae
The Hydrogenothermaceae family are bacteria that live in harsh environmental settings. They have been found in hot springs, sulfur pools, and thermal ocean vents. They are true bacteria as opposed to the other inhabitants of extreme environments, the Archaea. An example occurrence of certain extremophiles in this family are organisms of the genus ''Sulfurihydrogenibium'' that are capable of surviving in extremely hot environments such as Hverigerdi, Iceland. Obtaining energy Hydrogenothermaceae families consist of aerobic or microaerophilic bacteria, which generally obtain energy by oxidation of hydrogen or reduced sulfur compounds by molecular oxygen. Phylogeny The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LSPN) and the National Center for Biotechnology Information The National Center for Biotechnology Information (NCBI) is part of the United States National Library of Medicine (NLM), a branch of the National Inst ...
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Thermotogales
The Thermotogota are a phylum of the domain Bacteria. The phylum Thermotogota is composed of Gram-negative staining, anaerobic, and mostly thermophilic and hyperthermophilic bacteria.Gupta, RS (2014) The Phylum Thermotogae. The Prokaryotes 989-1015. Springer Berlin Heidelberg. Characteristics The name of this phylum is derived from the existence of many of these organisms at high temperatures along with the characteristic sheath structure, or "toga", surrounding the cells of these species.Reysenbach, A.-L. (2001) Phylum BII. Thermotogae phy. nov. In: Bergey's Manual of Systematic Bacteriology, pp. 369-387. Eds D. R. Boone, R. W. Castenholz. Springer-Verlag: Berlin. Recently, some Thermotogota existing at moderate temperatures have also been identified. Although Thermotogota species exhibit Gram-negative staining, they are bounded by a single-unit lipid membrane, hence they are monoderm bacteria. Because of the ability of some Thermotogota species to thrive at high temperatures, ...
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Aquifex
''Aquifex'' is a bacterial genus, belonging to phylum Aquificota. There is one species of ''Aquifex'' with a validly published name – '' A. pyrophilus'' – but "'' A. aeolicus''" is sometimes considered as species though it has no standing as a name given it has not been validly or effectively published. ''Aquifex'' spp. are extreme thermophiles, growing best at temperature of 85 °C to 95 °C. They are members of the ''Bacteria'' as opposed to the other inhabitants of extreme environments, the Archaea. ''Aquifex'' spp. are rod-shaped bacteria with a length of 2 to 6 µm, have a diameter of around 0.5 µm and are motile. They are non-sporeforming, Gram negative autotrophs. ''Aquifex'' means ''water-maker'' in Latin, and refers to the fact that its method of respiration creates water. ''Aquifex'' tend to form cell aggregates composed of up to 100 individual cells. ''Aquifex'' spp. are thermophilic and often grow near underwater volcanoes or hot springs. ''A. ae ...
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Conserved Signature Indels
Conserved signature inserts and deletions (CSIs) in protein sequences provide an important category of molecular markers for understanding phylogenetic relationships. CSIs, brought about by rare genetic changes, provide useful phylogenetic markers that are generally of defined size and they are flanked on both sides by conserved regions to ensure their reliability. While indels can be arbitrary inserts or deletions, CSIs are defined as only those protein indels that are present within conserved regions of the protein. The CSIs that are restricted to a particular clade or group of species, generally provide good phylogenetic markers of common evolutionary descent. Due to the rarity and highly specific nature of such changes, it is less likely that they could arise independently by either convergent or parallel evolution (i.e. homoplasy) and therefore are likely to represent synapomorphy. Other confounding factors such as differences in evolutionary rates at different sites or among ...
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Thermophilic
A thermophile is an organism—a type of extremophile—that thrives at relatively high temperatures, between . Many thermophiles are archaea, though they can be bacteria or fungi. Thermophilic eubacteria are suggested to have been among the earliest bacteria. Thermophiles are found in various geothermally heated regions of the Earth, such as hot springs like those in Yellowstone National Park (see image) and deep sea hydrothermal vents, as well as decaying plant matter, such as peat bogs and compost. Thermophiles can survive at high temperatures, whereas other bacteria or archaea would be damaged and sometimes killed if exposed to the same temperatures. The enzymes in thermophiles function at high temperatures. Some of these enzymes are used in molecular biology, for example the ''Taq'' polymerase used in PCR. "Thermophile" is derived from the el, θερμότητα (''thermotita''), meaning heat, and el, φίλια (''philia''), love. Classification Thermophiles can be c ...
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Horizontal Gene Transfer
Horizontal gene transfer (HGT) or lateral gene transfer (LGT) is the movement of genetic material between Unicellular organism, unicellular and/or multicellular organisms other than by the ("vertical") transmission of DNA from parent to offspring (reproduction). HGT is an important factor in the evolution of many organisms. HGT is influencing scientific understanding of higher order evolution while more significantly shifting perspectives on bacterial evolution. Horizontal gene transfer is the primary mechanism for the spread of antibiotic resistance in bacteria, and plays an important role in the evolution of bacteria that can degrade novel compounds such as human-created Bactericide, pesticides and in the evolution, maintenance, and transmission of virulence. It often involves Temperateness (virology), temperate bacteriophages and plasmids. Genes responsible for antibiotic resistance in one species of bacteria can be transferred to another species of bacteria through various m ...
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Cavalier-Smith
Thomas (Tom) Cavalier-Smith, FRS, FRSC, NERC Professorial Fellow (21 October 1942 – 19 March 2021), was a professor of evolutionary biology in the Department of Zoology, at the University of Oxford. His research has led to discovery of a number of unicellular organisms (protists) and advocated for a variety of major taxonomic groups, such as the Chromista, Chromalveolata, Opisthokonta, Rhizaria, and Excavata. He was known for his systems of classification of all organisms. Life and career Cavalier-Smith was born on 21 October 1942 in London. His parents were Mary Maude (née Bratt) and Alan Hailes Spencer Cavalier Smith. He was educated at Norwich School, Gonville and Caius College, Cambridge (MA) and King's College London (PhD). He was under the supervision of Sir John Randall for his PhD thesis between 1964 and 1967; his thesis was entitled "''Organelle Development in'' Chlamydomonas reinhardii". From 1967 to 1969, Cavalier-Smith was a guest investigato ...
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SecA
The SecA protein is a cell membrane associated subunit of the eubacterial Sec or Type II secretory pathway, a system which is responsible for the secretion of proteins through the cell membrane. Within this system the SecA ATPase forms a translocase complex with the SecYEG channel, thereby driving the movement of the protein substrate across the membrane. Structure SecA is a complex protein whose structure consists of six characterized domains that can explain SecA's capabilities to bind substrates and to move them. The following five domains seem to be present in all SecA proteins that have been structurally analyzed so far. DEAD motor domain This amino acid domain is subdivided into the two nucleotide binding folds 1 and 2 (NBF1 and NBF2) where ATP is bound and hydrolyzed. The chemical energy from the phosphodiester bonds results in a conformational change which is transferred to other domains (especially the HWD and the PPXD domains) which consequently mechanically mov ...
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Campylobacterota
Campylobacterota are a phylum of bacteria. All species of this phylum are Gram-negative. The Campylobacterota consist of few known genera, mainly the curved to spirilloid ''Wolinella'' spp., ''Helicobacter'' spp., and '' Campylobacter'' spp. Most of the known species inhabit the digestive tracts of animals and serve as symbionts (''Wolinella'' spp. in cattle) or pathogens (''Helicobacter'' spp. in the stomach, ''Campylobacter'' spp. in the duodenum). Many Campylobacterota are motile with flagella. Numerous environmental sequences and isolates of Campylobacterota have also been recovered from hydrothermal vents and cold seep habitats. Examples of isolates include ''Sulfurimonas autotrophica'', ''Sulfurimonas paralvinellae'', ''Sulfurovum lithotrophicum'' and ''Nautilia profundicola''. A member of the phylum Campylobacterota occurs as an endosymbiont in the large gills of the deepwater sea snail ''Alviniconcha hessleri''. The Campylobacterota found at deep-sea hydrothermal vents ...
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Hyperthermophilic
A hyperthermophile is an organism that thrives in extremely hot environments—from 60 °C (140 °F) upwards. An optimal temperature for the existence of hyperthermophiles is often above 80 °C (176 °F). Hyperthermophiles are often within the domain Archaea, although some bacteria are also able to tolerate extreme temperatures. Some of these bacteria are able to live at temperatures greater than 100 °C, deep in the ocean where high pressures increase the boiling point of water. Many hyperthermophiles are also able to withstand other environmental extremes, such as high acidity or high radiation levels. Hyperthermophiles are a subset of extremophiles. Their existence may support the possibility of extraterrestrial life, showing that life can thrive in environmental extremes. History Hyperthermophiles isolated from hot springs in Yellowstone National Park were first reported by Thomas D. Brock in 1965. Since then, more than 70 species have been established. The most extreme hyper ...
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