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Andaluciina
''Andalucia'' is a genus of jakobids. Classification The morphology of ''Andalucia'' broadly resembles that of other jakobids. Molecular data has not always been conclusive, but recent phylogenomic analyses indicate that ''Andalucia'' is a sister group to the other jakobids, in other words more closely related to them than to the Heterolobosea or Euglenozoa (the other two groups in the Discoba). The α-tubulin gene of ''Andalucia'' more closely resembles that of opisthokonts and diplomonads than its closer relatives, the apparent result of horizontal gene transfer. Species As of 2009, the genus contains two species: * '' A. incarcerata'' was until 2006 known as ''Jakoba incarcerata''. It lives in sulfide-rich sediments, especially marine intertidal ones. Currently it is the type species of the genus '' Stygiella'' Pánek, Táborský & Čepička 2015. * '' A. godoyi'' is a soil heterotroph A heterotroph (; ) is an organism that cannot produce its own food, instead taking ...
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Jakobid
Jakobids are an order of free-living, heterotrophic, flagellar eukaryotes in the supergroup Excavata. They are small (less than 15 μm), and can be found in aerobic and anaerobic environments. The order Jakobida, believed to be monophyletic, consists of only twenty species at present, and was classified as a group in 1993. There is ongoing research into the mitochondrial genomes of jakobids, which are unusually large and bacteria-like, evidence that jakobids may be important to the evolutionary history of eukaryotes. Molecular phylogenetic evidence suggests strongly that jakobids are most closely related to Heterolobosea (Percolozoa) and Euglenozoa. Structure and Biology Jakobids have two flagella, inserted in the anterior end of the cell, and, like other members of order Excavata, have a ventral feeding groove and associated cytoskeleton support. The posterior flagella has a dorsal vane and is aligned within the ventral groove, where it generates a current that the cell uses f ...
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Jakobea
Jakobids are an order of free-living, heterotrophic, flagellar eukaryotes in the supergroup Excavata. They are small (less than 15 μm), and can be found in aerobic and anaerobic environments. The order Jakobida, believed to be monophyletic, consists of only twenty species at present, and was classified as a group in 1993. There is ongoing research into the mitochondrial genomes of jakobids, which are unusually large and bacteria-like, evidence that jakobids may be important to the evolutionary history of eukaryotes. Molecular phylogenetic evidence suggests strongly that jakobids are most closely related to Heterolobosea (Percolozoa) and Euglenozoa. Structure and Biology Jakobids have two flagella, inserted in the anterior end of the cell, and, like other members of order Excavata, have a ventral feeding groove and associated cytoskeleton support. The posterior flagella has a dorsal vane and is aligned within the ventral groove, where it generates a current that the cell uses f ...
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Eukaryota
Eukaryotes () are organisms whose Cell (biology), cells have a cell nucleus, nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the Three-domain system, three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard (archaea), Asgard archaea. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass (ecology), biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as Flagellated cell, flagellated phagotrophs. The ...
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Diplomonad
The diplomonads (Greek for "two units") are a group of flagellates, most of which are parasitic. They include ''Giardia duodenalis'', which causes giardiasis in humans. They are placed among the metamonads, and appear to be particularly close relatives of the retortamonads. Most diplomonads are double cells: they have two nuclei, each with four associated flagella, arranged symmetrically about the body's main axis. Like the retortamonads, they lack both mitochondria and Golgi apparatuses. However, they are now known to possess modified mitochondria, in the case of ''G. duodenalis'', called mitosomes. These are not used in ATP synthesis the way mitochondria are, but are involved in the maturation of iron-sulfur proteins. Possible sexual reproduction in ''Giardia'' The common intestinal parasite ''Giardia duodenalis'' (synonyms ''Giardia lamblia'', ''G. intestinalis'') was once considered to be a descendant of a protist lineage that predated the emergence of meiosis and sex ...
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Heterotroph
A heterotroph (; ) is an organism that cannot produce its own food, instead taking nutrition from other sources of organic carbon, mainly plant or animal matter. In the food chain, heterotrophs are primary, secondary and tertiary consumers, but not producers. Living organisms that are heterotrophic include all animals and fungi, some bacteria and protists, and many parasitic plants. The term heterotroph arose in microbiology in 1946 as part of a classification of microorganisms based on their type of nutrition. The term is now used in many fields, such as ecology in describing the food chain. Heterotrophs may be subdivided according to their energy source. If the heterotroph uses chemical energy, it is a chemoheterotroph (e.g., humans and mushrooms). If it uses light for energy, then it is a photoheterotroph (e.g., green non-sulfur bacteria). Heterotrophs represent one of the two mechanisms of nutrition (trophic levels), the other being autotrophs (''auto'' = self, ''troph'' = ...
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Stygiella
''Stygiella'' /ˌstɪ.d͡ʒiˈɛ.lə/ is a genus of free-living marine flagellates belonging to the family Stygiellidae in the jakobids (excavata). The genus currently includes four species, all of which are secondary obligate anaerobes. The species are all unicellular and crescent-shaped.Bernard, C, Simpson, A. G. B. & Patterson, D. J. (2000) Some free-living flagellates (protista) from anoxic habitats, Ophelia, 52:2, 113-142, DOI: 10.1080/00785236.1999.10409422. All members possess hydrogenosomes, a type of acristate mitochondrion-derived organelle (MRO) that produces hydrogen gas as a metabolic product.Leger, M. M., Eme, L., Hug, L. A., & Roger, A. J. (2016). ovel hydrogenosomes in the microaerophilic jakobid ''Stygiella incarcerata'' Molecular Biology and Evolution, 33(9), 2318-2336. Retrieved April 28, 2020. doi: doi.org/10.1093/2Fmolbev/2Fmsw103. Stygiella is a deep-branching lineage within excavates, and hydrogenosome genes sometimes show eubacterium-like mechanisms th ...
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Intertidal Zone
The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species of life, such as seastars, sea urchins, and many species of coral with regional differences in biodiversity. Sometimes it is referred to as the ''littoral zone'' or '' seashore'', although those can be defined as a wider region. The well-known area also includes steep rocky cliffs, sandy beaches, bogs or wetlands (e.g., vast mudflats). The area can be a narrow strip, as in Pacific islands that have only a narrow tidal range, or can include many meters of shoreline where shallow beach slopes interact with high tidal excursion. The peritidal zone is similar but somewhat wider, extending from above the highest tide level to below the lowest. Organisms in the intertidal zone are adapted to an environment of harsh extremes, living in water pr ...
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Sulfide
Sulfide (British English also sulphide) is an inorganic anion of sulfur with the chemical formula S2− or a compound containing one or more S2− ions. Solutions of sulfide salts are corrosive. ''Sulfide'' also refers to chemical compounds large families of inorganic and organic compounds, e.g. lead sulfide and dimethyl sulfide. Hydrogen sulfide (H2S) and bisulfide (SH−) are the conjugate acids of sulfide. Chemical properties The sulfide ion, S2−, does not exist in aqueous alkaline solutions of Na2S. Instead sulfide converts to hydrosulfide: :S2− + H2O → SH− + OH− Upon treatment with an acid, sulfide salts convert to hydrogen sulfide: :S2− + H+ → SH− :SH− + H+ → H2S Oxidation of sulfide is a complicated process. Depending on the conditions, the oxidation can produce elemental sulfur, polysulfides, polythionates, sulfite, or sulfate. Metal sulfides react with halogens, forming sulfur and metal salts. :8 MgS + 8 I2 → S8 + 8 M ...
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Jakoba
''Jakoba'' is a genus in the taxon Excavata, and currently has a single described species, ''Jakoba libera'' described by Patterson in 1990, and named in honour of Dutch botanist (Algology, Myology and Lichenology) Jakoba Ruinen. (Previously described ''Jakoba incarcerata'' has been renamed ''Andalucia incarcerata'', and ''Jakoba bahamensis'' /''Jakoba bahamiensis'' is not formally described.) Appearance and characteristics ''Jakoba'' are small bacterivorous zooflagellates (jakobids) found in marine and hypersaline environments. They are free swimming trophic cells with two flagella and range between five and ten micrometers in length. Cells rotate along their longitudinal axis to allow for swimming in straight lines unless deformation and “squirming” occurs due to compression in debris. During feeding, bacteria are removed from the water column by a current created by the posterior flagellum. This current causes the bacteria to collect in the groove on the right ventra ...
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Andalucia Incarcerata
''Andalucia incarcerata'' is a species of Excavata Excavata is a major supergroup of unicellular organisms belonging to the domain Eukaryota. It was first suggested by Simpson and Patterson in 1999 and introduced by Thomas Cavalier-Smith in 2002 as a formal taxon. It contains a variety of free- .... It used to be known as ''Jakoba incarcerata'' but is currently placed in the genus ''Stygiella'' Pánek, Táborský & Čepička 2015 as ''Stygiella incarcerata'' (Bernard, Simpson & Patterson 2000) Pánek, Táborský & Čepička 2015. References Jakobids {{Excavata-stub ...
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Horizontal Gene Transfer
Horizontal gene transfer (HGT) or lateral gene transfer (LGT) is the movement of genetic material between Unicellular organism, unicellular and/or multicellular organisms other than by the ("vertical") transmission of DNA from parent to offspring (reproduction). HGT is an important factor in the evolution of many organisms. HGT is influencing scientific understanding of higher order evolution while more significantly shifting perspectives on bacterial evolution. Horizontal gene transfer is the primary mechanism for the spread of antibiotic resistance in bacteria, and plays an important role in the evolution of bacteria that can degrade novel compounds such as human-created Bactericide, pesticides and in the evolution, maintenance, and transmission of virulence. It often involves Temperateness (virology), temperate bacteriophages and plasmids. Genes responsible for antibiotic resistance in one species of bacteria can be transferred to another species of bacteria through various m ...
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Tubulin
Tubulin in molecular biology can refer either to the tubulin protein superfamily of globular proteins, or one of the member proteins of that superfamily. α- and β-tubulins polymerize into microtubules, a major component of the eukaryotic cytoskeleton. Microtubules function in many essential cellular processes, including mitosis. Tubulin-binding drugs kill cancerous cells by inhibiting microtubule dynamics, which are required for DNA segregation and therefore cell division. In eukaryotes, there are six members of the tubulin superfamily, although not all are present in all species.Turk E, Wills AA, Kwon T, Sedzinski J, Wallingford JB, Stearns "Zeta-Tubulin Is a Member of a Conserved Tubulin Module and Is a Component of the Centriolar Basal Foot in Multiciliated Cells"Current Biology (2015) 25:2177-2183. Both α and β tubulins have a mass of around 50 kDa and are thus in a similar range compared to actin (with a mass of ~42 kDa). In contrast, tubulin polymers (microtubules) te ...
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