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Alalcomeneaus
''Alalcomenaeus'' is one of the most widespread and longest-surviving arthropod genera of the Early and Middle Cambrian. Known from over 300 specimens in the Burgess Shale and the Chengjiang biota. it is a member of the group Megacheira. Morphology ''Alalcomenaeus'' had three median eyes; two stalked, more lateral eyes; a triflagellate great appendage; and two more head appendages posterior to that. Like its body appendages, these were biramous—their inner branch was spiny, segmented, flexible and leg-like, while the outer portion had a large surface area and resembled a flap. ''Alalcomenaeus'' reached about 6 cm in length, although many smaller specimens are known. Its head was covered with a shield, and its eleven body segments were also covered with an exoskeleton. Its body terminated with a paddle-like telson ("tail") which probably helped to propel the organism; this ended with long flat spikes in the plane of the tail fin. Ecology The organism probably swam, waftin ...
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Royal Ontario Museum
The Royal Ontario Museum (ROM) is a museum of art, world culture and natural history in Toronto, Ontario, Canada. It is one of the largest museums in North America and the largest in Canada. It attracts more than one million visitors every year, making the ROM the most-visited museum in Canada. The museum is north of Queen's Park, in the University of Toronto district, with its main entrance on Bloor Street West. Museum subway station is named after the ROM and, since a 2008 renovation, is decorated to resemble the institution's collection at the platform level. Established on April 16, 1912, and opened on March 19, 1914, the museum has maintained close relations with the University of Toronto throughout its history, often sharing expertise and resources. The museum was under the direct control and management of the University of Toronto until 1968, when it became an independent Crown agency of the Government of Ontario. Today, the museum is Canada's largest field-research in ...
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Sanctacaris
''Sanctacaris'' is a Middle Cambrian arthropod from the Burgess Shale of British Columbia. It was most famously regarded as a stem-group chelicerate, a group which includes horseshoe crabs, spiders and scorpions, although subsequent phylogenetic studies have not always supported this conclusion. Its chelicerate affinities regain support in later observations, alongside the reassignment of ''Habelia optata'' as a sanctacaridid-related basal chelicerate. It has been placed as a member of the extinct family Sanctacarididae alongside ''Wisangocaris'' and '' Utahcaris''. ''Sanctacaris'' specimens range from 46 to 93 mm in length. The head bears five pairs of grasping appendages (corresponding to chelicerate's pedipalps and walking legs) and a sixth pair of posterior appendages (correspond to horseshoe crab's chilaria). The grasping appendages each bear an antenna-like exopods. There are 11 body segments, with the former 10 each bearing a pair of biramus appendage with flap-like ex ...
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Wheeler Shale
The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna (including ''Naraoia'', ''Wiwaxia'' and ''Hallucigenia'') and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten. Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America. At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thi ...
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Cambrian Genus Extinctions
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biolo ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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Prehistoric Arthropod Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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Phyllopod Bed
The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess Shale fossil ''Lagerstätte''. It was quarried by Charles Walcott from 1911–1917 (and later named Walcott Quarry), and was the source of 95% of the fossils he collected during this time; tens of thousands of soft-bodied fossils representing over 150 genera have been recovered from the Phyllopod bed alone. Stratigraphy and location The phyllopod bed is a 2.31 m thick layer of the 7 m thick Greater Phyllopod Bed, found in the Walcott Quarry on Fossil Ridge, between Wapta Mountain and Mount Field, at an elevation of around , around north of the railway town of Field, British Columbia, in the Canadian Rocky Mountains. It is adjacent to Mount Burgess, where Walcott first discovered the Burgess Shale formation. Walcott divided the bed into twelve units based on the rock type and fossil content. Certain fossil beds provide reference levels and can b ...
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Stratigraphic Range
Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostratigraphy (lithologic stratigraphy), biostratigraphy (biologic stratigraphy), and chronostratigraphy (stratigraphy by age). Historical development Catholic priest Nicholas Steno established the theoretical basis for stratigraphy when he introduced the law of superposition, the principle of original horizontality and the principle of lateral continuity in a 1669 work on the fossilization of organic remains in layers of sediment. The first practical large-scale application of stratigraphy was by William Smith in the 1790s and early 19th century. Known as the "Father of English geology", Smith recognized the significance of strata or rock layering and the importance of fossil markers for correlating strata; he created the first geologic ...
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Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its west by Nevada. Utah also touches a corner of New Mexico in the southeast. Of the fifty U.S. states, Utah is the 13th-largest by area; with a population over three million, it is the 30th-most-populous and 11th-least-densely populated. Urban development is mostly concentrated in two areas: the Wasatch Front in the north-central part of the state, which is home to roughly two-thirds of the population and includes the capital city, Salt Lake City; and Washington County in the southwest, with more than 180,000 residents. Most of the western half of Utah lies in the Great Basin. Utah has been inhabited for thousands of years by various indigenous groups such as the ancient Puebloans, Navajo and Ute. The Spanish were the first Europe ...
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Chengjiang County
Chengjiang (; earlier Tchinkiang) is a city located in Yuxi, Yunnan Province, China, just north of Fuxian Lake. Administrative divisions Chengjiang City has 2 subdistricts and 4 townships. ;2 subdistricts * Fenglu () * Longjie () ;4 towns Chengjiang Fossil Site In evolutionary biology, and especially paleontology, Chengjiang is noted for soft-tissue fossil finds, of the Maotianshan Shales, dated less than 518 million years ago during the Cambrian explosion, which "are as spectacular as the Burgess Shale fauna, and significantly older." Also These fossils are considered one of the most important fossil finds of the 20th century. They contain an exquisite degree of detail, cover a diverse range of fauna, and are significant in attempts to understand the evolution of life on Earth. In 2012, the Changjiang Fossil Site became a UNESCO World Heritage Site. The fossils were first discovered by Henri Mansuy and Jaques Deprat who described them in 1912, the year after Charles Walc ...
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Arachnomorph
Arachnomorpha is a proposed subdivision or clade of Arthropoda, comprising the group formed by the trilobites and their close relatives (Artiopoda), Megacheira (which may be paraphyletic) and chelicerates. Under this proposed classification scheme, Arachnomorpha is considered the sister group to Mandibulata (including insects, crustaceans and myriapods). The arachnomorph concept has been challenged by suggestions that the artiopods are more closely related to mandibulates, forming the clade Antennulata instead. There is no consensus as to assigning Arachnomorpha a formal Linnean rank. Classification Arachnomorpha Lameere, 1890 Arachnata Paulus, 1979, = Palaeopoda Packard, 1903*†Trilobita Walch, 1771 *†Megacheira Hou & Bergstrom, 1997 *†Strabopida Hou & Bergström, 1997 *†Aglaspida Walcott, 1911 *† Cheloniellida Broili, 1932 *Chelicerata Heymons 1901 **Family † Sanctacarididae Legg & Pates, 2017 **Class Pycnogonida Latreille, 1810 **Clade Euchelicerata Weygoldt ...
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Sidneyia
''Sidneyia'' is an extinct arthropod known from fossils found from the Early Cambrian-age Maotianshan Shales to the Mid Cambrian Burgess Shale formation of British Columbia. 144 specimens of ''Sidneyia'' are known from the Greater Phyllopod bed, where they comprise 0.27% of the community. General description ''Sidneyia'' ranged from in length and is one of the largest arthropods found at the site. It is thought to have been a benthic carnivore and scavenger that walked along the sea floor in search of hard-shelled prey. Gut contents have revealed that ''Sidneyia'' fed largely on small trilobites, as well as on brachiopods, hyoliths and small arthropods. The gut was narrow, but widens posteriorly to form a pocket where digestion presumably took place. The retention of feces likely indicates infrequent feeding Its exquisitely preserved gnathobases resemble those of ''Limulus'', and were probably used to crush prey. ''Sidneyia'' was discovered in 1910 during the first day of ...
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