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Acleistorhinus
''Acleistorhinus'' (ah-kles-toe-RYE-nuss) is an extinct genus of parareptile known from the Early Permian (middle Kungurian stage) of Oklahoma. It is notable for being the earliest known anapsid reptile yet discovered. The morphology of the lower temporal fenestra of the skull of ''Acleistorhinus'' bears a superficial resemblance to that seen in early synapsids, a result of convergent evolution. Only a single species, ''A. pteroticus'', is known, and it is classified in the Family Acleistorhinidae, along with ''Colobomycter'' (also from the Early Permian of Oklahoma). Etymology ''Acleistorhinus'' was first discovered and named by Eleanor Daly in 1969 in the Hennessey Formation of South Grandfield, Tillman county, Oklahoma. The name ''Acleistorhinus'' combines Greek ''rhin'' (ῥῑ́ν), meaning "nose," and akleistos, Greek for “unclosed.“ Description and Paleobiology Skull Although its total body length is unknown, an ''Acleistorhinus'' skull is about 3.5 centimetres lo ...
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Acleistorhinus Four Images
''Acleistorhinus'' (ah-kles-toe-RYE-nuss) is an extinct genus of Parareptilia, parareptile known from the Early Permian (middle Kungurian stage) of Oklahoma. It is notable for being the earliest known Anapsida, anapsid reptile yet discovered. The morphology (biology), morphology of the lower temporal fenestra of the skull of ''Acleistorhinus'' bears a superficial resemblance to that seen in early Synapsida, synapsids, a result of convergent evolution. Only a single species, ''A. pteroticus'', is known, and it is classified in the Family (biology), Family Acleistorhinidae, along with ''Colobomycter'' (also from the Early Permian of Oklahoma). Etymology ''Acleistorhinus'' was first discovered and named by Eleanor Daly in 1969 in the Hennessey Formation of South Grandfield, Tillman county, Oklahoma. The name ''Acleistorhinus'' combines Greek ''rhin'' (ῥῑ́ν), meaning "nose," and akleistos, Greek for “unclosed.“ Description and Paleobiology Skull Although its total body leng ...
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Acleistorhinidae
Acleistorhinidae is an extinct family of Late Carboniferous and Early Permian-aged ( Moscovian to Kungurian stage) parareptiles. Acleistorhinids are most diverse from the Richards Spur locality of the Early Permian of Oklahoma. Richards Spur acleistorhinids include '' Acleistorhinus'', '' Colobomycter'', and possibly '' Delorhynchus'' and '' Feeserpeton''. Other taxa include ''Carbonodraco'' from the Late Carboniferous of Ohio and '' Karutia'' from the Early Permian of Brazil. Acleistorhinidae is commonly considered a subgroup of lanthanosuchoids, related to taxa such as '' Chalcosaurus'', '' Lanthaniscus'' and '' Lanthanosuchus''. However, a re-examination of parareptile phylogeny conducted by Cisneros ''et al.'' (2021) argued that lanthanosuchids were not closely related to acleistorhinids. The phylogenetic analysis conducted by these authors recovered acleistorhinids as the sister group of the clade Procolophonia, while lanthanosuchids were recovered within the procolophon ...
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Colobomycter
''Colobomycter'' is an extinct genus of lanthanosuchoid parareptile known from the Early Permian of Oklahoma. Discovery The type species, ''Colobomycter pholeter'', was first described from fossil remains in 1958, at which time it was believed to represent a synapsid, specifically, a pelycosaur. However, the discovery of new material and reexamination of the holotype led to its reclassification as a member of the Eureptilia. More recent studies indicate that ''Colobomycter'' is properly placed within the amniote clade Parareptilia, as part of the group Lanthanosuchoidea and closely related to the taxon '' Acleistorhinus''. A second species of ''Colobomycter'' was described in 2016, ''Colobomycter vaughni''. No postcranial material is known for ''Colobomycter'', and the skull material referred to the genus has all been recovered from a single locality, the Richards Spur site at the Dolese Brothers Limestone Quarry, 11 kilometers north of Fort Sill, Comanche County, Oklaho ...
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Parareptilia
Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as ''Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by the ...
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Parareptile
Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as ''Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by the ...
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Parareptilia
Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as ''Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by the ...
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Macroleter
''Macroleter'' is an extinct genus of nycteroleterid parareptile which existed in Oklahoma and Russia during the upper Permian period. It was a quite generalized primitive reptile, in many ways resembling their amphibian ancestors. It was first named by paleontologists Tverdochlebova and Ivachnenko in 1984. According to classification by Michel Laurin and Robert R. Reisz, the genus is a parareptile, belonging to the same branch as Millerettidae, Procolophonidae and other generalized anapsid reptiles. The type species is ''Macroleter poezicus'' from Upper Permian of Russia. ''Macroleter'' had an 8 cm skull, and an overall length of 75 cm. It was generally lizard-like in build with a rather flat and broad skull. The teeth were small and pointy, indication it predominantly hunted insects and other small invertebrates. ''Seymouria agilis'' (Olson, 1980) that is known from only one specimen (holotype UCMP 143 277) was originally thought to be a reptile-like amphibian a ...
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Lanthanosuchidae
Lanthanosuchidae is a family of procolophonomorph parareptile Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near th ...s that lived 268–255 million years ago. The group was named in 1946 by Ivan Antonovich Efremov.Ivakhnenko, M.F. (1980). "Lanthanosuchids from the Permian of the East European Platform". ''Paleontologica Journal'' 1980: 80-90. References Procolophonomorphs Permian reptiles Prehistoric reptile families {{paleo-reptile-stub ...
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Millerettidae
Millerettidae is an extinct family of parareptiles from the Middle Permian to the Late Permian period (Capitanian - Changhsingian stages) of South Africa. The millerettids were small insectivores and probably resembled modern lizards in appearance and lifestyle. The following cladogram shows the phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... position of the Millerettidae, from Ruta ''et al.'', 2011. References External linksBerkeley University information. Permian reptiles Prehistoric reptiles of Africa Guadalupian first appearances Lopingian extinctions Prehistoric reptile families Parareptiles {{Permian-reptile-stub ...
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Diapsida
Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are sometimes placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. Although some diapsids have lost either one hole (lizards), or both holes (snakes and turtles), or have a heavily restructured skull (modern birds), they are still classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals are extant: 9,159 birds, and 7,925 snakes, lizards, tuatara, turtles, and crocodiles. Characteristics The name Diapsida means "two arches", and diapsids are tradition ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Westphalian (stage)
The Westphalian is a stage in the regional stratigraphy of northwest Europe with an age between roughly 313 and 304 Ma (million years ago). It is a subdivision of the Carboniferous system or period and the regional Silesian series. The Westphalian is named for the region of Westphalia (German: ''Westfalen'') in western Germany where strata of this age occur. The Coal Measures of England and Wales are also largely of Westphalian age though they also extend into the succeeding Stephanian. The Westphalian age is preceded by the Namurian stage/age (which corresponds to the Millstone Grit Series of Great Britain) and succeeded by the Stephanian stage/age (which corresponds to the uppermost part of the Coal Measures of Great Britain). In the official geologic timescale of the ICS, the Westphalian is placed within the Pennsylvanian epoch (318-299 Ma).F M Gradstein, J G Ogg, A G Smith et al 2004, ‘A Geologic Time Scale 2004’. The (regionally defined) Westphalian stage corres ...
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