Tryporyza
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''Scirpophaga'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Crambidae The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies includ ...
described by
Georg Friedrich Treitschke Georg Friedrich Treitschke (; 29 August 1776 – 4 June 1842) was a German libretto, librettist, translator and lepidopterist. He was born in Leipzig and died in Vienna. In 1800 he came to the Vienna Hofoper. From 1809 to 1814 he was principal ...
in 1832. Asian species include significant
rice Rice is the seed of the grass species ''Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima ''Oryza glaberrima'', commonly known as African rice, is one of the two domesticated rice species. It was first domesticated and grown i ...
stemborer pests.


Taxonomy

The genus ''Scirpophaga'' was first introduced by Treitschke in 1832 as a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus; including as single species ''Scirpophaga phantasmatella'' (which he misspelled as ''S. phantasmella'', and which is now known as ''S. praelata''). During most of history this genus has been completely confused, with most specimens being wrongly identified and most taxa being based on a
type series In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes t ...
containing numerous species. Males and females of the same species were often recognised as two independent species. Almost two centuries after the first species was described, in 1960 the Australian
entomologist Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arach ...
Ian Francis Bell Common was the first to examine the
genitalia A sex organ (or reproductive organ) is any part of an animal or plant that is involved in sexual reproduction. The reproductive organs together constitute the reproductive system. In animals, the testis in the male, and the ovary in the female, a ...
(for centuries the standard method by which one determines species in Lepidoptera) of the Australian specimens in this group, recombining and splitting the then defined genera into a number of new genera. He created the new genus ''Tryporyza'', in which he incorporated two species: ''Chilo incertulas'' and ''Tipanaea innotata'' (of which DNA research in 2019 has shown should be synonymised with ''
Scirpophaga nivella ''Scirpophaga nivella'' is a moth in the family Crambidae. It was described by Johan Christian Fabricius in 1794. It is found in southern Asia from the Indian Subcontinent in the west to southern China in the east, south to New Guinea and Austra ...
''). In 1980 P. Wang also classified ''Scirpophaga nivella'' within the genus ''Tryporyza'', only to have Angoon Lewvanich, after an exhaustive study of the genitals of over 6000 specimens from throughout the range of the group, to retire the genus ''Tryporyza'' as a synonym of ''Scirpophaga'' in the following year (1981). Treitschke gives as
etymology Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological chan ...
for the
generic epithet Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
the word ''
Scirpus ''Scirpus'' is a genus of grass-like species in the sedge family Cyperaceae many with the common names club-rush, wood club-rush or bulrush (see also bulrush for other plant genera so-named). They mostly inhabit wetlands and damp locations. Taxo ...
'', a type of plant commonly known as a "rush", and the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
word φαγεῖν (transliterated ''phageîn''), which means "to eat".


Description

Palpi Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ...
porrect (extending forward) extending from once to twice the length of head, slightly hairy, and with downcurved third joint. Maxillary palp rather short and dilated with scales. A slight rounded frontal projection can be seen. Antennae of male minutely serrated, and ciliated. Patagia of male with spreading upturned hair. Tibia with outer spurs about half the length of inner. Abdomen long, where in female expanding at extremity and with very large anal tuft. Wings long and narrow. Forewings with vein 3 from before angle of cell. Veins 4 and 5 from angle and vein 7 straight and well separated from veins 8 and 9. Veins 10 and 11 free, or vein 11 becoming coincident with vein 12. Hindwings with vein 3 from near angle of cell. Veins 6 and 7 from upper angle.


Species

*''praelata''
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
**''
Scirpophaga gilviberbis ''Scirpophaga gilviberbis'' is a moth in the family Crambidae. It was described by Philipp Christoph Zeller in 1863. It is found on the Comoros and in the Democratic Republic of the Congo, Kenya, South Africa, Zambia, India, Indonesia (Java, Sula ...
'' Zeller, 1863 **'' Scirpophaga humilis'' Wang, Li & Chen, 1986 **''
Scirpophaga imparellus ''Scirpophaga imparellus'' is a moth in the family Crambidae. It was described by Edward Meyrick in 1878. It is found in Australia, where it has been recorded from Queensland, New South Wales and Victoria Victoria most commonly refers to: * V ...
'' (Meyrick, 1878) **''
Scirpophaga melanoclista ''Scirpophaga melanoclista'' is a moth in the family Crambidae. It was described by Edward Meyrick in 1935. It is found in Angola, the Democratic Republic of the Congo, Ghana, Madagascar, Senegal, Sierra Leone and Zambia. References

Moth ...
'' Meyrick, 1935 **''
Scirpophaga nivella ''Scirpophaga nivella'' is a moth in the family Crambidae. It was described by Johan Christian Fabricius in 1794. It is found in southern Asia from the Indian Subcontinent in the west to southern China in the east, south to New Guinea and Austra ...
'' (Fabricius, 1794) **''
Scirpophaga parvalis ''Scirpophaga parvalis'' is a moth in the family Crambidae. It was described by Alfred Ernest Wileman in 1911. It is found in China (Heilongjiang, Beijing, Hebei, Shandong, Henan, Shaanxi, Gansu Jiangsu, Anhui, Hubei, Fujian), Japan and Korea. ...
'' (Wileman, 1911) **''
Scirpophaga percna ''Scirpophaga percna'' is a moth in the family Crambidae. It was described by Ian Francis Bell Common in 1960. It is found on Sulawesi, on New Guinea and northern Australia, where it has been recorded from the Northern Territory and Queensland. ...
'' Common, 1960 **''
Scirpophaga praelata ''Scirpophaga praelata'' is a species of moth of the family Crambidae. It is found in most of Europe (except Ireland, Great Britain, Portugal, the Benelux, Germany, Fennoscandia, Estonia and Latvia), Russia, Turkey, Iran, Syria, Lebanon, North ...
'' (Scopoli, 1763) **''
Scirpophaga phaedima ''Scirpophaga phaedima'' is a moth in the family Crambidae. It was described by Ian Francis Bell Common in 1960. It is found in northern Australia. The wingspan The wingspan (or just span) of a bird or an airplane is the distance from one ...
'' Common, 1960 **''
Scirpophaga xantharrenes ''Scirpophaga xantharrenes'' is a moth in the family Crambidae. It was described by Ian Francis Bell Common in 1960. It is found in Australia, where it has been recorded from Queensland ) , nickname = Sunshine State , image_map = Queensland ...
'' Common, 1960 **''
Scirpophaga xanthopygata ''Scirpophaga xanthopygata'' is a species of moth in the family Crambidae. It is found in Russia, Ukraine, Korea, eastern China, Japan, Vietnam and Taiwan. The wingspan The wingspan (or just span) of a bird or an airplane is the distance fr ...
'' Schawerda, 1922 *''excerptalis'' species group **'' Scirpophaga adunctella'' Chen, Song & Wu, 2006 **'' Scirpophaga bradleyi'' Lewvanich, 1981 **'' Scirpophaga brunnealis'' (Hampson, 1919) **'' Scirpophaga excerptalis'' (Walker, 1863) **'' Scirpophaga flavidorsalis'' (Hampson, 1919) **'' Scirpophaga khasis'' Lewvanich, 1981 **''
Scirpophaga linguatella ''Scirpophaga linguatella'' is a moth in the family Crambidae. It was described by Fu-Qiang Chen, Shi-Mei Song and Chun-Sheng Wu in 2006. It is found in Yunnan, China China, officially the People's Republic of China (PRC), is a country i ...
'' Chen, Song & Wu, 2006 **''
Scirpophaga magnella ''Scirpophaga magnella'' is a moth in the family Crambidae. It was described by Joseph de Joannis in 1930. It is found in China (Zhejiang, Hunan, Guangdong, Hainan, Hong Kong, Guangxi, Sichuan, Yunnan, Xizang), Iran, Afghanistan, Pakistan, India, ...
'' de Joannis, 1930 **'' Scirpophaga ochritinctalis'' (Hampson, 1919) **'' Scirpophaga xanthogastrella'' (Walker, 1863) **'' Scirpophaga tongyaii'' Lewvanich, 1981 *''occidentella'' species group **''
Scirpophaga fusciflua ''Scirpophaga fusciflua'' is a moth in the family Crambidae. It was described by George Hampson in 1893. It is found in Taiwan, Afghanistan, India, Nepal, Thailand and Sri Lanka. The wingspan is 16–22 mm for males and 21–27 mm for ...
'' Hampson, 1893 **'' Scirpophaga goliath'' Marion & Viette, 1953 **'' Scirpophaga marginepunctellus'' (de Joannis, 1927) **''
Scirpophaga occidentella ''Scirpophaga occidentella'' is a moth in the family Crambidae. It was described by Francis Walker (entomologist), Francis Walker in 1863. It is found in Angola, the Democratic Republic of the Congo, Ivory Coast, Madagascar, Malawi, Mozambique, N ...
'' (Walker, 1863) **''
Scirpophaga ochroleuca ''Scirpophaga ochroleuca'' is a moth in the family Crambidae. It was described by Edward Meyrick in 1882. It is found on New Guinea and in Australia, where it has been recorded from Queensland. The wingspan The wingspan (or just span) of a ...
'' Meyrick, 1882 **'' Scirpophaga serenus'' (Meyrick, 1935) **''
Scirpophaga subumbrosa ''Scirpophaga subumbrosa'' is a moth in the family Crambidae. It was described by Edward Meyrick in 1933. It is found in the Democratic Republic of the Congo (West Kasai, Katanga), Ethiopia, Ghana, Madagascar, Malawi, Mozambique, Nigeria, Senegal ...
'' Meyrick, 1933 **'' Scirpophaga virginia'' Schultze, 1908 *''lineata'' species group **'' Scirpophaga lineata'' (Butler, 1879) **''
Scirpophaga auristrigellus ''Scirpophaga auristrigellus'' is a moth in the family Crambidae. It was described by George Hampson in 1896. It is found in Guangxi, China, India and Bhutan. The wingspan is 20–26 mm. The forewings are pale yellow, but yellow between the ...
'' (Hampson, 1896) **'' Scirpophaga aurivena'' (Hampson, 1903) *''incertulas'' species group **''
Scirpophaga incertulas ''Scirpophaga incertulas'', the yellow stem borer or rice yellow stem borer, is a species of moth of the family Crambidae. It was described by Francis Walker in 1863. It is found in Afghanistan, Nepal, north-eastern India, Sri Lanka, Banglade ...
'' (Walker, 1863) **''
Scirpophaga innotata ''Scirpophaga innotata'', the rice white stemborer, is a species of moth of the family Crambidae. The species was described by Francis Walker in 1863. It is found in Indonesia, Pakistan, the Philippines and the tropical north of Australia. Th ...
'' (Walker, 1863) *''gotoi'' species group **'' Scirpophaga gotoi'' Lewvanich, 1981 *''whalleyi'' species group **'' Scirpophaga whalleyi'' Lewvanich, 1981 *unknown species group **''
Scirpophaga bipunctatus ''Scirpophaga bipunctatus'' is a moth in the family Crambidae. It was described by Rothschild in 1926. It is found in Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السود ...
'' (Rothschild in Sjöstedt, 1926) **''
Scirpophaga fulvilinealis ''Scirpophaga fulvilinealis'' is a moth in the family Crambidae. It was described by George Hampson in 1900. It is found in Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a ...
'' Hampson, 1900 **''
Scirpophaga kumatai ''Scirpophaga kumatai'' is a moth in the family Crambidae. It was described by Angoon Lewvanich in 1981. It is found in Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, ...
'' Lewvanich, 1981 **'' Scirpophaga micraurea'' Sasaki, 1994 **'' Scirpophaga nepalensis'' Lewvanich, 1981 **'' Scirpophaga terrella'' Hampson, 1896


References

* Chen, F.Q., Song, S. & Weng, C.S. (2006). "A review of the genus ''Scirpophaga'' Treitschke, 1832 in China (Lepidoptera: Pyralidae)" ''Zootaxa'' 1236: 1-22. * Fabricius (1794). ''Entomologia Systematica emendata et Aucta'' 3(2): 296. * * Schawerda (1922). ''Z. öst. Ent. ver.'' 7: 11. * Scopoli (1763). ''Ent. carniolica, exhibens Ins. Carnioliae indigena'': 198. * Treitschke (1832). ''Die Schmetterlinge von Europa'' 9(1): 55. * Wang P, Li, C. & Chen, X. (1986) ''Acta Entomologica Sinica'' 29 (2): 208–210. * Wileman (1911). ''Transactions of the Entomological Society of London'': 355. {{Taxonbar, from=Q7434043 Schoenobiinae Crambidae genera Taxa named by Georg Friedrich Treitschke