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Tree squirrels are the members of the squirrel
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
(Sciuridae) commonly just referred to as "squirrels." They include more than 100
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose num ...
species native to all continents except
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest co ...
and
Oceania Oceania (, , ) is a region, geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern Hemisphere, Eastern and Western Hemisphere, Western hemispheres, Oceania is estimated to have a land area of ...
. They do not form a single natural, or
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
, group; they are variously related to others in the squirrel family, including
ground squirrel Ground squirrels are members of the squirrel family of rodents (Sciuridae), which generally live on or in the ground, rather than trees. The term is most often used for the medium-sized ground squirrels, as the larger ones are more commonly known ...
s, flying squirrels, marmots, and chipmunks. The defining characteristic used to determine which species of Sciuridae are tree squirrels is dependent on their
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
rather than their
physiology Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemic ...
. Tree squirrels live mostly among trees, as opposed to those that live in
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of s ...
s in the ground or among rocks. An exception is the flying squirrel that also makes its home in trees, but has a physiological distinction separating it from its tree squirrel cousins: special flaps of skin called patagia, acting as glider wings, which allow gliding flight. The best known
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of tree squirrels is '' Sciurus'', which includes the
eastern gray squirrel The eastern gray squirrel (''Sciurus carolinensis''), also known, particularly outside of North America, as simply the grey squirrel, is a tree squirrel in the genus '' Sciurus''. It is native to eastern North America, where it is the most pro ...
of North America ( introduced to
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
in the 1870s), the red squirrel of Eurasia, and the North American
fox squirrel The fox squirrel (''Sciurus niger''), also known as the eastern fox squirrel or Bryant's fox squirrel, is the largest species of tree squirrel native to North America. Despite the differences in size and coloration, it is sometimes mistaken for ...
, among many others. Many tree squirrel species have adapted to human-altered environments such as rural
farms A farm (also called an agricultural holding) is an area of land that is devoted primarily to agricultural processes with the primary objective of producing food and other crops; it is the basic facility in food production. The name is used fo ...
, suburban backyards and urban parks.


Classification

Current
taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
, based on genetic data, splits the tree squirrels into several subfamilies. The following
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...
of the squirrel family are classified as tree squirrels. *Subfamily Ratufinae **Genus '' Ratufa'' (Asian giant squirrels) *Subfamily
Sciurillinae The Neotropical pygmy squirrel (''Sciurillus pusillus'') is a South American species of tree squirrel, being the only living species in the genus ''Sciurillus'' and the subfamily Sciurillinae. Genetic analysis has shown it to be the sister group ...
**Genus '' Sciurillus'' (South American pygmy squirrel) *Subfamily Sciurinae **Tribe Sciurini (mostly American tree squirrels) ***Genus '' Microsciurus'' (American dwarf squirrels) ***Genus ''
Rheithrosciurus The tufted ground squirrel or groove-toothed squirrel (''Rheithrosciurus macrotis'') is a species of rodent in the family Sciuridae. It is the sole species in the genus ''Rheithrosciurus''. It is found only on the island of Borneo. Confirmed elem ...
'' (Borneo tufted ground squirrel) ***Genus '' Sciurus'' (Eurasian and American tree squirrels) ***Genus '' Syntheosciurus'' (Central American mountain squirrel) ***Genus ''
Tamiasciurus Pine squirrels are squirrels of the genus ''Tamiasciurus'', in the Sciurini tribe, of the large family Sciuridae. Species This genus includes three species: *''Tamiasciurus hudsonicus'' — American red squirrel *''Tamiasciurus douglasi ...
'' (American pine squirrels) *Subfamily Callosciurinae (Asian tree squirrels) **Genus '' Callosciurus'' (Oriental tree squirrels, introduced into Europe and South America) **Genus '' Exilisciurus'' (Asian pygmy squirrels) **Genus '' Funambulus'' (Asian palm squirrels, introduced into Australia in the 1920s) **Genus ''
Glyphotes The sculptor squirrel or Bornean pygmy squirrel (''Glyphotes simus'') is a species of rodent in the family Sciuridae. It was described by Michael Rogers Oldfield Thomas in 1898. It is monotypic within the genus ''Glyphotes''. It is endemic to Mal ...
'' (sculptor squirrel) **Genus ''
Nannosciurus The black-eared squirrel (''Nannosciurus melanotis'') is a species of rodent in the family Sciuridae. It is monotypic within the genus ''Nannosciurus''. This tiny squirrel is found in forests in Borneo, Sumatra and Java. Except for its striking w ...
'' (Asian dwarf squirrel) **Genus '' Prosciurillus'' (Sulawesi dwarf squirrels) **Genus '' Rubrisciurus'' (Sulawesi giant squirrel) **Genus '' Sundasciurus'' (Sunda squirrels) **Genus '' Tamiops'' (Asian striped squirrels) *Subfamily Xerinae **Tribe Protoxerini (African tree squirrels) ***Genus '' Epixerus'' (African palm squirrels) ***Genus '' Funisciurus'' (rope squirrels) ***Genus '' Heliosciurus'' (sun squirrels) ***Genus '' Myosciurus'' (African pygmy squirrel) ***Genus '' Paraxerus'' (bush squirrels) ***Genus '' Protoxerus'' (African giant squirrels)


Relationship with humans

Squirrels are generally inquisitive and persistent animals. In residential neighborhoods, they are notorious for tenaciously trying to circumvent obstacles in order to eat from
bird feeder A birdfeeder, bird table, or tray feeder are devices placed outdoors to supply bird food to birds ( bird feeding). The success of a bird feeder in attracting birds depends upon its placement and the kinds of foods offered, as different species ...
s. Although they are expert climbers, and primarily
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose num ...
, some species of squirrels also thrive in urban environments, where they have adapted to humans.


As pests

Squirrels are sometimes considered
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
because of their propensity to chew on various edible and inedible objects, and their stubborn persistence in trying to get what they want. Their characteristic gnawing trait also aids in maintaining sharp
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, te ...
, and because their teeth grow continuously, prevents their over-growth. On occasion, squirrels will chew through plastic and even metal to get to food. Tree squirrels may bury food in the ground for later retrieval. Squirrels use their keen sense of smell to search for buried food, but can dig numerous holes in the process. This may become an annoyance to gardeners with strict landscape requirements, especially when the garden contains edibles. Homeowners in areas with a heavy squirrel population must be vigilant in keeping attics, basements, and sheds carefully sealed to prevent property damage caused by nesting squirrels. A squirrel nest is called a "
drey A drey is the nest of a tree squirrel, flying squirrel or ringtail possum. Dreys are usually built of twigs, dry leaves, and grass, and typically assembled in the forks of a tall tree. They are sometimes referred to as "drey nests" to dist ...
". Squirrels are a serious fire hazard when they break into buildings. They often treat exposed power cables as tree branches, and gnaw on the
electrical insulation An electrical insulator is a material in which electric current does not flow freely. The atoms of the insulator have tightly bound electrons which cannot readily move. Other materials—semiconductors and conductors—conduct electric current ...
. The resulting exposed conductors can short out, causing a fire. For this reason alone, squirrel nests inside buildings cannot be safely ignored. A squirrel nest will also cause problems with noise, excreta, unpleasant odors, and eventual structural damage. Some homeowners resort to more interesting ways of dealing with this problem, such as collecting and placing fur from pets such as domestic
cats The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
and dogs in attics. It is hoped that this fur would indicate to nesting squirrels that a potential predator roams, and will encourage evacuation. Odoriferous repellents, including mothballs and
ammonia Ammonia is an inorganic compound of nitrogen and hydrogen with the formula . A stable binary hydride, and the simplest pnictogen hydride, ammonia is a colourless gas with a distinct pungent smell. Biologically, it is a common nitrogeno ...
, are generally ineffective in expelling squirrels from buildings. Once established in a nest, squirrels stubbornly ignore fake owls and
scarecrow A scarecrow is a decoy or mannequin, often in the shape of a human. Humanoid scarecrows are usually dressed in old clothes and placed in open fields to discourage birds from disturbing and feeding on recently cast seed and growing crops.Lesle ...
s, along with bright flashing lights, loud noises, and ultrasonic or
electromagnetic In physics, electromagnetism is an interaction that occurs between particles with electric charge. It is the second-strongest of the four fundamental interactions, after the strong force, and it is the dominant force in the interactions of a ...
devices. However, squirrels must leave the nest to obtain food and water (usually daily, except in bad weather), affording an opportunity to trap them or exclude them from re-entering. To discourage chewing on an object, it can be coated or covered with something to make it distasteful: for instance a soft cloth doused with
chili pepper Chili peppers (also chile, chile pepper, chilli pepper, or chilli), from Nahuatl '' chīlli'' (), are varieties of the berry-fruit of plants from the genus '' Capsicum'', which are members of the nightshade family Solanaceae, cultivated for ...
paste or powder.
Capsaicin Capsaicin (8-methyl-''N''-vanillyl-6-nonenamide) ( or ) is an active component of chili peppers, which are plants belonging to the genus ''Capsicum''. It is a chemical irritant for mammals, including humans, and produces a sensation of burning ...
and Ro-pel are other forms of repellent. To remain effective, the coating must be reapplied regularly, especially if it is exposed to the weather.
Poisoning A poison can be any substance that is harmful to the body. It can be swallowed, inhaled, injected or absorbed through the skin. Poisoning is the harmful effect that occurs when too much of that substance has been taken. Poisoning is not to ...
squirrels can be problematic because of the risks to other animals or children in the building, and because the odor of a dead squirrel in an attic or wall cavity is very unpleasant and persistent. Trapping is often necessary to remove squirrels from residential structures. Effective baits include fruit,
peanut butter Peanut butter is a food paste or spread made from ground, dry-roasted peanuts. It commonly contains additional ingredients that modify the taste or texture, such as salt, sweeteners, or emulsifiers. Peanut butter is consumed in many countri ...
, nuts, seeds and vanilla extract. An alternative method is to wait until squirrels have left in search of food, and then close up all their access openings, or to install one-way trap doors or a carefully angled pipe. Attempting to get rid of all squirrels in a neighborhood is generally a futile goal; the focus instead should be on physically excluding them from places where they can do damage. There are other humane techniques to remove squirrels from buildings, but removal is ineffective unless steps are taken to prevent them from immediately breaking in again. Squirrels are often the cause of
power outage A power outage (also called a powercut, a power out, a power failure, a power blackout, a power loss, or a blackout) is the loss of the electrical power network supply to an end user. There are many causes of power failures in an electrici ...
s. They can readily climb a power pole and crawl or run along a power cable. The animals will climb onto power
transformer A transformer is a passive component that transfers electrical energy from one electrical circuit to another circuit, or multiple circuits. A varying current in any coil of the transformer produces a varying magnetic flux in the transformer' ...
s or
capacitor A capacitor is a device that stores electrical energy in an electric field by virtue of accumulating electric charges on two close surfaces insulated from each other. It is a passive electronic component with two terminals. The effect of a ...
s looking for food, or a place to cache
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera '' Quercus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and bo ...
s. If they touch a high voltage conductor and a
ground Ground may refer to: Geology * Land, the surface of the Earth not covered by water * Soil, a mixture of clay, sand and organic matter present on the surface of the Earth Electricity * Ground (electricity), the reference point in an electrical c ...
ed portion of the enclosure at the same time, they are electrocuted, and often cause a
short circuit A short circuit (sometimes abbreviated to short or s/c) is an electrical circuit that allows a current to travel along an unintended path with no or very low electrical impedance. This results in an excessive current flowing through the circuit ...
that shuts down equipment. Squirrels have brought down the high-tech
NASDAQ The Nasdaq Stock Market () (National Association of Securities Dealers Automated Quotations Stock Market) is an American stock exchange based in New York City. It is the most active stock trading venue in the US by volume, and ranked second ...
stock market twice and were responsible for a spate of power outages at the
University of Alabama The University of Alabama (informally known as Alabama, UA, or Bama) is a public research university in Tuscaloosa, Alabama. Established in 1820 and opened to students in 1831, the University of Alabama is the oldest and largest of the publi ...
. To sharpen their teeth, squirrels will often chew on tree branches or even the occasional live power line. Rubber or plastic plates, or freely rotating sleeves ("squirrel guards") are sometimes used to discourage access to these facilities. Squirrels otherwise appear to be safe and pose almost zero risk of transmitting
rabies Rabies is a viral disease that causes encephalitis in humans and other mammals. Early symptoms can include fever and tingling at the site of exposure. These symptoms are followed by one or more of the following symptoms: nausea, vomiting, vio ...
. Squirrels cause economic losses to homeowners, nut growers, and forest managers in addition to damage to electric transmission lines. These losses include direct damage to property, repairs, lost revenue and public relations. While dollar costs of these losses are sometimes calculated for isolated incidents, there is no tracking system to determine the total extent of the losses.


As roadkill and traffic hazards

In regions where squirrels are plentiful, tire-flattened roadkill is a common sight on roadways, especially in the spring and fall, when there is a fresh crop of young rodents. Motorists have caused serious accidents by attempting to swerve or stop to avoid a squirrel in the road. Evasive maneuvers are difficult since squirrels are much more agile and have much quicker reaction times than motorists in heavy vehicles; the majority of vehicular encounters end with no harm to either party. An effort to mitigate these hazards to both squirrels and humans is the Nutty Narrows Bridge in Longview, Washington, listed on the
National Register of Historic Places The National Register of Historic Places (NRHP) is the United States federal government's official list of districts, sites, buildings, structures and objects deemed worthy of preservation for their historical significance or "great artist ...
. It provides a way for squirrels to cross a busy street safely.


As urban wildlife

Tree squirrels are a common type of
urban wildlife Urban wildlife is wildlife that can live or thrive in urban/suburban environments or around densely populated human settlements such as townships. Some urban wildlife, such as house mice, are synanthropic, ecologically associated with and e ...
. They can be trained to be hand-fed and will take as much food as is available because they cache the surplus. Squirrels living in parks and campuses in cities have learned that humans are typically a ready source of food. Urban squirrels have learned to get a lot of food from generous or unknowingly 'careless' humans. Humans commonly offer various nuts and seeds; however, wildlife rehabilitators in the field have noted that neither raw nor roasted
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible Seed, seeds. It is widely grown in the tropics and subtropics, important to both small ...
s nor
sunflower seed The sunflower seed is the seed of the sunflower (''Helianthus annuus''). There are three types of commonly used sunflower seeds: linoleic (most common), high oleic, and sunflower oil seeds. Each variety has its own unique levels of monounsatu ...
s are healthy for squirrels, because they are deficient in several essential nutrients. This type of deficiency has been found to cause metabolic bone disease, a somewhat common ailment found in malnourished squirrels.


As food


In the US

In many areas of the U.S. squirrels are still hunted for food, as they were historically.It tasted like chicken
Retrieved December 19, 2008.
Recipes calling for squirrel even appear in cookbooks, including ''James Beard's American Cookery'' and pre-1997 copies of ''
The Joy of Cooking ''Joy of Cooking'', often known as "''The Joy of Cooking''", is one of the United States' most-published cookbooks. It has been in print continuously since 1936 and has sold more than 20 million copies. It was published privately during 1931 b ...
''. Squirrel meat can be substituted for
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit s ...
or
chicken The chicken (''Gallus gallus domesticus'') is a domestication, domesticated junglefowl species, with attributes of wild species such as the grey junglefowl, grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster ...
in many recipes and was an ingredient in the original recipe for Brunswick stew, a popular dish in various parts of the Southern U.S. Other similar stews were also based on squirrel meat, including burgoo and
Southern Illinois chowder Southern Illinois Chowder is a thick stew or soup, very different from the New England and Manhattan chowders. In Edwards County, Illinois, it refers to both the food and to the social gathering at which it is prepared and served. It is believ ...
. Although squirrel meat is low in fat content, unlike most game meat it has been found by the
American Heart Association The American Heart Association (AHA) is a nonprofit organization in the United States that funds cardiovascular medical research, educates consumers on healthy living and fosters appropriate cardiac care in an effort to reduce disability and death ...
to be high in
cholesterol Cholesterol is any of a class of certain organic molecules called lipids. It is a sterol (or modified steroid), a type of lipid. Cholesterol is biosynthesized by all animal cells and is an essential structural component of animal cell membr ...
. Squirrels Unlimited host a World Championship Squirrel cook-off each year in Bentonville, Arkansas.


In the UK

For most of the history of the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the European mainland, continental mainland. It comprises England, Scotlan ...
, squirrel has been a meat not commonly eaten, and even scorned by many. In the early 21st century however, wild squirrel has become a more popular meat to cook with, showing up in restaurants and shops more often in
Britain Britain most often refers to: * The United Kingdom, a sovereign state in Europe comprising the island of Great Britain, the north-eastern part of the island of Ireland and many smaller islands * Great Britain, the largest island in the United King ...
as a fashionable alternative meat. Specifically, Britons are cooking with the invasive gray squirrel, which is praised for its low fat content and the fact that it comes from free range sources. Additionally, the novelty of a meat considered unusual or special has contributed to the spread of squirrel consumption. Due to the difficulty of a clean kill and other factors, the majority of squirrel eaten in the UK is acquired from professional hunters, trappers and gamekeepers. Some Britons are eating gray squirrel as a direct attempt to help the native red squirrel, which has been dwindling since the 19th-century introduction of the gray squirrel, resulting in dramatic
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
for the indigenous red squirrels. This factor was marketed by a national "Save Our Squirrels" campaign that used the slogan, "Save a red, eat a grey!"


Risks of eating

As with other wild game and
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...
species, the consumption of squirrels that have been exposed to high levels of
pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the ...
or toxic waste poses a health risk to humans. In 2007 in the northern New Jersey community of Ringwood, the New Jersey Department of Health and Senior Services issued a warning to anyone who eats squirrel (especially children and those who are pregnant) to limit their consumption after a
lead Lead is a chemical element with the Symbol (chemistry), symbol Pb (from the Latin ) and atomic number 82. It is a heavy metals, heavy metal that is density, denser than most common materials. Lead is Mohs scale of mineral hardness#Intermediate ...
-contaminated squirrel was found near the Ringwood Mines Landfill. Toxic waste had been illegally dumped at this location for many years, before authorities cracked down on this practice in the 1980s. In 1997, doctors in
Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virgini ...
published a paper in the '' Lancet'' that considered a possible association between the local tradition of consuming squirrel brains and five cases of Creutzfeldt–Jakob disease, a rare but serious
prion Prions are misfolded proteins that have the ability to transmit their misfolded shape onto normal variants of the same protein. They characterize several fatal and transmissible neurodegenerative diseases in humans and many other animals. It ...
-based disorder. The authors posed this as a mere possibility, unconfirmed by either post-mortem analysis of the patients' brain tissue, or identification of a contagious prion agent in squirrels. Nonetheless, the ''Lancet'' article generated substantial media coverage, including articles in the New Yorker and
New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid ...
. A 2015 case of CJD in a
Pittsburgh Pittsburgh ( ) is a city in the Commonwealth (U.S. state), Commonwealth of Pennsylvania, United States, and the county seat of Allegheny County, Pennsylvania, Allegheny County. It is the most populous city in both Allegheny County and Wester ...
man who had eaten squirrel brains played out similarly: the media seized on the patient's unconventional food choice, positing squirrel brains as the source of his disease. The doctor who made the initial report later clarified that he had not meant to assert the squirrel meat was the cause. Analysis of the patient's brain tissue ruled out the possibility of CJD acquired from food. As of 2018, Creutzfeldt–Jakob disease had never been identified in squirrels, and the association between squirrel consumption and CJD remained speculative.


Relationship with trees

The biggest source of food for tree squirrels is tree nuts. Red squirrels store nuts in a single stash (a midden) that tends to dry out, so the seeds don't take root. Fox squirrels and gray squirrels bury nuts over a widespread area (
scatterhoarding Hoarding or caching in animal behavior is the storage of food in locations hidden from the sight of both conspecifics (animals of the same or closely related species) and members of other species. Most commonly, the function of hoarding or cachi ...
), and often forget them, resulting in new trees ( mutualism).


In culture

In the ''
Ramayana The ''Rāmāyana'' (; sa, रामायणम्, ) is a Sanskrit epic composed over a period of nearly a millennium, with scholars' estimates for the earliest stage of the text ranging from the 8th to 4th centuries BCE, and later stages e ...
'', an ancient
Sanskrit Sanskrit (; attributively , ; nominalization, nominally , , ) is a classical language belonging to the Indo-Aryan languages, Indo-Aryan branch of the Indo-European languages. It arose in South Asia after its predecessor languages had Trans-cul ...
epic poem An epic poem, or simply an epic, is a lengthy narrative poem typically about the extraordinary deeds of extraordinary characters who, in dealings with gods or other superhuman forces, gave shape to the mortal universe for their descendants. ...
, a squirrel assists in constructing a bridge from India to
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
to help
Rama Rama (; ), Ram, Raman or Ramar, also known as Ramachandra (; , ), is a major deity in Hinduism. He is the seventh and one of the most popular ''avatars'' of Vishnu. In Rama-centric traditions of Hinduism, he is considered the Supreme Being ...
rescue his wife
Sita Sita (; ) also called as Janaki and Vaidehi is a Hindu goddess and the female protagonist of the Hindu epic, '' Ramayana''. She is the consort of Rama, the avatar of the god Vishnu, and is regarded as a form of Vishnu's consort, Lakshmi ...
. Rama rewards the squirrel by stroking his back with his three middle fingers, thus giving the Indian palm squirrel the three white stripes that appear on its back. In
Norse mythology Norse, Nordic, or Scandinavian mythology is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia, and into the Nordic folklore of the modern peri ...
, the squirrel Ratatoskr is a messenger who scurries up and down the trunk of the world-tree
Yggdrasil Yggdrasil (from Old Norse ), in Norse cosmology, is an immense and central sacred tree. Around it exists all else, including the Nine Worlds. Yggdrasil is attested in the '' Poetic Edda'' compiled in the 13th century from earlier tradition ...
, carrying malicious gossip and insults back and forth between the dragon Níðhöggr, who sits at the bottom of the tree gnawing on its roots, and the hawk Veðrfölnir, who sits at the top of the tree keeping watch. According to Richard W. Thorington, Jr. and Katie E. Ferrell, this legend may have originated from the red squirrel's habit of giving a "scolding alarm call in response to danger", which some Norsemen may have imagined as insults. In
Irish mythology Irish mythology is the body of myths native to the island of Ireland. It was originally passed down orally in the prehistoric era, being part of ancient Celtic religion. Many myths were later written down in the early medieval era by ...
, the goddess
Medb Medb (), later spelled Meadhbh (), Méibh () and Méabh (), and often anglicised as Maeve ( ), is queen of Connacht in the Ulster Cycle of Irish mythology. Her husband in the core stories of the cycle is Ailill mac Máta, although she had seve ...
is said to always have a bird perched on one shoulder and a squirrel on the other, serving as her messengers to the sky and the earth respectively. In Europe during the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
, squirrels were sometimes used in bestiaries as symbols of greed and avarice on account of their storing of nuts, but, in the nineteenth century, British natural history books often praised them as thrifty for this same reason. A myth told by the
Ainu people The Ainu are the indigenous people of the lands surrounding the Sea of Okhotsk, including Hokkaido Island, Northeast Honshu Island, Sakhalin Island, the Kuril Islands, the Kamchatka Peninsula and Khabarovsk Krai, before the arrival of the ...
of Japan holds that squirrels are the discarded sandals of the ancestral deity Aioina, possibly because squirrels move in spurts like footsteps. The ''
Kalevala The ''Kalevala'' ( fi, Kalevala, ) is a 19th-century work of epic poetry compiled by Elias Lönnrot from Karelian and Finnish oral folklore and mythology, telling an epic story about the Creation of the Earth, describing the controversies an ...
'', a Finnish epic poem collected in the nineteenth century but rooted in much older
oral tradition Oral tradition, or oral lore, is a form of human communication wherein knowledge, art, ideas and cultural material is received, preserved, and transmitted orally from one generation to another. Vansina, Jan: ''Oral Tradition as History'' (1985 ...
, contains references to squirrels, including mention of a white squirrel being born of a virgin.
Literary Literature is any collection of written work, but it is also used more narrowly for writings specifically considered to be an art form, especially prose fiction, drama, and poetry. In recent centuries, the definition has expanded to includ ...
references to squirrels include the works of
Beatrix Potter Helen Beatrix Potter (, 28 July 186622 December 1943) was an English writer, illustrator, natural scientist, and conservationist. She is best known for her children's books featuring animals, such as '' The Tale of Peter Rabbit'', which was ...
, Brian Jacques' Redwall series (including Jess Squirrel and numerous other squirrels), Pattertwig in C. S. Lewis' '' Prince Caspian'', Michael Tod's Woodstock Saga of novels featuring squirrel communities in the style of ''
Watership Down ''Watership Down'' is an adventure novel by English author Richard Adams, published by Rex Collings Ltd of London in 1972. Set in Berkshire in southern England, the story features a small group of rabbits. Although they live in their natural w ...
'', and the Starwife and her subjects from Robin Jarvis's Deptford novels. The title character in Miriam Young's 1964 children's book ''Miss Suzy'' is a squirrel.
Anthropomorphic Anthropomorphism is the attribution of human traits, emotions, or intentions to non-human entities. It is considered to be an innate tendency of human psychology. Personification is the related attribution of human form and characteristics t ...
red squirrels were used in British road safety campaigns between the 1950s and 1980s. An episode of the radio program ''
This American Life ''This American Life'' (''TAL'') is an American monthly hour-long radio program produced in collaboration with Chicago Public Media and hosted by Ira Glass. It is broadcast on numerous public radio stations in the United States and internatio ...
'' called "Squirrel Cop" describes the unintentionally humorous misadventure of a newly hired policeman in trying to remove a frantic squirrel from a homeowner's living room, which results in personal injury and a small fire. First aired in 1998, this episode turned out to be one of the most popular ones of the series, prompting rebroadcasts and a lead position on the two-CD compilation '' Crimebusters + Crossed Wires: Stories from This American Life''.


Albino and white squirrels

One of the ways that squirrels affect human society is inspired by the fascination that people seem to have over local populations of white squirrels (often misidentified as being albino). This manifests itself by the creation of
social group In the social sciences, a social group can be defined as two or more people who interact with one another, share similar characteristics, and collectively have a sense of unity. Regardless, social groups come in a myriad of sizes and varieties ...
communities that form from a commonly shared interest in these rare animals. Other impacts on human society inspired by white squirrels include the creation of organizations that seek to protect them from human predation, and the use of the white squirrel image as a
cultural icon A cultural icon is a person or an artifact that is identified by members of a culture as representative of that culture. The process of identification is subjective, and "icons" are judged by the extent to which they can be seen as an authentic s ...
. Although these squirrels are commonly referred to as "albinos", most of them are likely non-albino squirrels that exhibit a rare white fur coloration known as leucism that is as a result of a recessive gene found within certain eastern gray squirrel (''Sciurus carolinensis'') populations, and so technically they ought to be referred to as ''white'' squirrels, instead of ''albino''."Debunking the Albino Squirrel: Lack of white, pink-eyed critters on campus may disprove superstition" by Hudson Lockett, The Daily Texan, May 1, 2009. A project run by Untamed Science is seeking to report and document the occurrence of both white squirrels, albinos, and other
piebald A piebald or pied animal is one that has a pattern of unpigmented spots (white) on a pigmented background of hair, feathers or scales. Thus a piebald black and white dog is a black dog with white spots. The animal's skin under the white backgro ...
morphs. Users are encouraged to submit their sightings.The White and Albino Squirrel Phenomenon + New US Maps!
/ref> UNTalbinosquirrel.jpg, A true albino squirrel. Note the pink eyes. Albino squirrel Minneapolis 2018.jpg, Albino squirrel head on, showing blue irises Wsquirrel.png, A white squirrel. Note the non-pink eyes.


Local pride

Olney, Illinois, known as the "White Squirrel Capital of the World", is home of the world's largest known white squirrel colony. These squirrels have the right of way on all streets in the town, with a $500 fine for hitting one. The Olney Police Department features the image of a white squirrel on its officers' uniform patches. Along with Olney, there are four other towns in North America that avidly compete with each other to be the official "Home of the White Squirrel", namely:
Marionville, Missouri Marionville is a city in Lawrence County, Missouri, United States. The population was 2,225 at the 2010 census. Marionville is known for its large population of white squirrels. History Marionville was platted in 1854. The city name has been ...
;
Brevard, North Carolina Brevard is a city in Transylvania County, North Carolina, United States, with a population of 7,609 as of the 2010 Census. It is the county seat of Transylvania County. Brevard is located at the entrance to Pisgah National Forest and has becom ...
; Exeter, Ontario; and Kenton, Tennessee, each of which holds an annual white squirrel festival, among other things designed to promote their claim of "White Squirrel Capital". A list of white squirrel sightings around the world is maintained by the White Squirrel Research Institute, a group based in Brevard, North Carolina. Other towns that have reported white squirrel populations in North America (although not necessarily competing to be the "official" white squirrel capital) include
Bowling Green, Kentucky Bowling Green is a home rule-class city and the county seat of Warren County, Kentucky, United States. Founded by pioneers in 1798, Bowling Green was the provisional capital of Confederate Kentucky during the American Civil War. As of the ...
; Columbia, Mississippi; DeForest, Wisconsin; Stratford, Connecticut; and some of the snowbelt cities in the
Western Western may refer to: Places *Western, Nebraska, a village in the US * Western, New York, a town in the US *Western Creek, Tasmania, a locality in Australia *Western Junction, Tasmania, a locality in Australia *Western world, countries that i ...
, Central and
Finger Lakes The Finger Lakes are a group of eleven long, narrow, roughly north–south lakes located south of Lake Ontario in an area called the ''Finger Lakes region'' in New York, in the United States. This region straddles the northern and transitional ...
regions of New York State ( Buffalo,
Rochester Rochester may refer to: Places Australia * Rochester, Victoria Canada * Rochester, Alberta United Kingdom *Rochester, Kent ** City of Rochester-upon-Medway (1982–1998), district council area ** History of Rochester, Kent ** HM Prison ...
, Ithaca and
Syracuse Syracuse may refer to: Places Italy *Syracuse, Sicily, or spelled as ''Siracusa'' *Province of Syracuse United States * Syracuse, New York ** East Syracuse, New York ** North Syracuse, New York * Syracuse, Indiana *Syracuse, Kansas *Syracuse, M ...
). The
Trinity Bellwoods Trinity-Bellwoods is an inner city neighbourhood in Toronto, Ontario, Canada. It is bounded on the east by Bathurst Street, on the north by College Street, on the south by Queen Street West, and by Dovercourt Road on the west. It has a large Por ...
neighborhood of Toronto, Ontario is locally known for white squirrel sightings.


Campus populations

In addition to the various towns that boast of their white squirrel populations, a number of university campuses in North America have white squirrels. The
University of Texas at Austin The University of Texas at Austin (UT Austin, UT, or Texas) is a public research university in Austin, Texas. It was founded in 1883 and is the oldest institution in the University of Texas System. With 40,916 undergraduate students, 11,075 ...
is home to a white squirrel population which has spurred the myth of the albino squirrel as a good luck charm. There are many versions of the tale; one of the more popular versions is if one spots the albino squirrel before an exam, they will ace it. The
University of North Texas The University of North Texas (UNT) is a public research university in Denton, Texas. It was founded as a nonsectarian, coeducational, private teachers college in 1890 and was formally adopted by the state 11 years later."Denton Normal Schoo ...
founded the
Albino Squirrel Preservation Society {{unreferenced, date=August 2011 The Albino Squirrel Preservation Society (ASPS) is an international collegiate organization dedicated to "fostering compassion and goodwill" toward albino squirrels. The ASPS has approximately 700 members in eight ...
in 2001, which has since acquired several "worldwide" chapters. In 2006, the University of North Texas held a student referendum to name their white squirrel as the university's secondary mascot, but the vote was narrowly defeated by the student body. University of Wisconsin - Eau Claire has a significant white squirrel population both on the campus and in other areas of the city of
Eau Claire Eau Claire (French for "clear water", ''pl.'' ''eaux claires'') is the name of a number of locations and features in North America. The name is pronounced as if it were spelled "O'Clare". Place names (Canada) Communities *Eau Claire, Calgary, a n ...
.
Michigan Technological University Michigan Technological University (Michigan Tech, MTU, or simply Tech) is a public research university in Houghton, Michigan, founded in 1885 as the Michigan Mining School, the first post-secondary institution in the Upper Peninsula of Michigan. ...
in
Houghton, Michigan Houghton (; ) is the largest city and seat of government of Houghton County in the U.S. state of Michigan. Located on the Keweenaw Peninsula, Houghton is the largest city in the Copper Country region. It is the fifth-largest city in the U ...
is home to frequently sighted white squirrels that live on and around the campus. A
Facebook Facebook is an online social media and social networking service owned by American company Meta Platforms. Founded in 2004 by Mark Zuckerberg with fellow Harvard College students and roommates Eduardo Saverin, Andrew McCollum, Dustin ...
group dedicated to these squirrels, called ''I've Seen the Albino Squirrel of Michigan Tech'', was created for people to post photographs and anecdotes of their encounters with the white squirrels, and includes some stories from Michigan Tech alumni that recall seeing white squirrels in Houghton dating back to the 1930s. In Kentucky, the
University of Louisville The University of Louisville (UofL) is a public research university in Louisville, Kentucky. It is part of the Kentucky state university system. When founded in 1798, it was the first city-owned public university in the United States and one of ...
has established its own chapter of the Albino Squirrel Preservation Society, which maintains contact with its members and interested parties through a Facebook group by that name. The university has an open policy to give away a free t-shirt to anyone who takes a photograph of a white squirrel on campus grounds and brings it to the administration offices. Other university campuses that have albino squirrel populations include Oberlin College in Ohio,
Ohio State University The Ohio State University, commonly called Ohio State or OSU, is a public land-grant research university in Columbus, Ohio. A member of the University System of Ohio, it has been ranked by major institutional rankings among the best pu ...
in
Columbus, Ohio Columbus () is the state capital and the most populous city in the U.S. state of Ohio. With a 2020 census population of 905,748, it is the 14th-most populous city in the U.S., the second-most populous city in the Midwest, after Chicago, an ...
,
Western Kentucky University Western Kentucky University is a public university in Bowling Green, Kentucky. It was founded by the Commonwealth of Kentucky in 1906, though its roots reach back a quarter-century earlier. It operates regional campuses in Glasgow, Elizabet ...
in
Bowling Green, Kentucky Bowling Green is a home rule-class city and the county seat of Warren County, Kentucky, United States. Founded by pioneers in 1798, Bowling Green was the provisional capital of Confederate Kentucky during the American Civil War. As of the ...
(which has had a population of albino squirrels since the 1960s), and Youngstown State University in
Youngstown, Ohio Youngstown is a city in the U.S. state of Ohio, and the largest city and county seat of Mahoning County, Ohio, Mahoning County. At the 2020 United States census, 2020 census, Youngstown had a city population of 60,068. It is a principal city of ...
. Michael Stokes, a biology professor at Western Kentucky University, commented that the probable cause for the abundance of white squirrels on university campuses was because they were originally introduced by someone: "We're not sure how they got here, but I'll tell you how it usually happens...When you see them, especially around a college campus or parks, somebody brought them in because they thought it would be neat to have white squirrels around." Albert Meier, another biology professor at Western Kentucky University, added that: "... white squirrels rarely survive in the wild because they can't easily hide. But on a college campus, they are less likely to be consumed by other animals."


In folklore

A story in which a
Nāga The Nagas (IAST: ''nāga''; Devanāgarī: नाग) are a divine, or semi-divine, race of half-human, half-serpent beings that reside in the netherworld (Patala), and can occasionally take human or part-human form, or are so depicted in art. ...
shapeshifts into a white or albino squirrel, is killed by a hunter, and is magically transformed into meat equal to 8,000
cart A cart or dray (Australia and New Zealand) is a vehicle designed for transport, using two wheels and normally pulled by one or a pair of draught animals. A handcart is pulled or pushed by one or more people. It is different from the flatbed tr ...
loads figures prominently in the
folklore Folklore is shared by a particular group of people; it encompasses the traditions common to that culture, subculture or group. This includes oral traditions such as Narrative, tales, legends, proverbs and jokes. They include material culture, r ...
of rocket festival
tradition A tradition is a belief or behavior (folk custom) passed down within a group or society with symbolic meaning or special significance with origins in the past. A component of cultural expressions and folklore, common examples include holidays ...
s and the origin of Nong Han Kumphawapi Lake in Northeast Thailand.


Red and gray squirrels in the UK

A decline of the red squirrel and the rise of the
eastern gray squirrel The eastern gray squirrel (''Sciurus carolinensis''), also known, particularly outside of North America, as simply the grey squirrel, is a tree squirrel in the genus '' Sciurus''. It is native to eastern North America, where it is the most pro ...
, an
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived the ...
from North America, has been widely remarked upon in British popular culture. It is mostly regarded as the invading grays driving out the native red species. Evidence also shows that gray squirrels are vectors of the squirrel parapoxvirus for which no vaccine is currently available, and which is deadly to red squirrels but does not seem to affect the non-native host. Currently, the red squirrel's range has been reduced to the coniferous forests in Scotland, and in England's
Formby Formby is a town and civil parish in the Metropolitan Borough of Sefton, Merseyside, England, which had a population of 22,419 at the 2011 Census. Historically in Lancashire, three manors are recorded in the Domesday Book of 1086 under "Forn ...
, the Lake District,
Brownsea Island Brownsea Island is the largest of the islands in Poole Harbour in the county of Dorset, England. The island is owned by the National Trust with the northern half managed by the Dorset Wildlife Trust. Much of the island is open to the public an ...
, and the
Isle of Wight The Isle of Wight ( ) is a Counties of England, county in the English Channel, off the coast of Hampshire, from which it is separated by the Solent. It is the List of islands of England#Largest islands, largest and List of islands of England#Mo ...
. The majority of England's red squirrels are found in the county of
Northumberland Northumberland () is a ceremonial counties of England, county in Northern England, one of two counties in England which border with Scotland. Notable landmarks in the county include Alnwick Castle, Bamburgh Castle, Hadrian's Wall and Hexham Ab ...
. Special measures are in place to contain and remove any infiltration of gray squirrels into these areas. Though the population has dramatically decreased, they remain listed on the IUCN Red List as Least Concern. As of 2008, the eastern gray squirrel was regarded as
vermin Vermin (colloquially varmint(s) or varmit(s)) are pests or nuisance animals that spread diseases or destroy crops or livestock. Since the term is defined in relation to human activities, which species are included vary by region and enterpri ...
and it was illegal to release any into the wild; any caught could be releases only if one applied for and was granted a licence to do so. As of 2015, any caught in Scotland had to be humanely killed.


See also

* American red squirrel *
Black squirrel Black squirrels are a melanistic subgroup of squirrels with black coloration on their fur. The phenomenon occurs with several species of squirrels, although it is most frequent with the eastern gray squirrel (''Sciurus carolinensis'') and the ...
*
Eastern gray squirrel The eastern gray squirrel (''Sciurus carolinensis''), also known, particularly outside of North America, as simply the grey squirrel, is a tree squirrel in the genus '' Sciurus''. It is native to eastern North America, where it is the most pro ...
*
Fox squirrel The fox squirrel (''Sciurus niger''), also known as the eastern fox squirrel or Bryant's fox squirrel, is the largest species of tree squirrel native to North America. Despite the differences in size and coloration, it is sometimes mistaken for ...
* Red squirrel * Squirrel * Western gray squirrel


References

;Footnotes ;Sources


Further reading

* *Nowak, Ronald M. 1999. ''Walker's Mammals of the World'', 6th edition. Johns Hopkins University Press, 1936 pp., *Young, Miriam. 1964. ''Miss Suzy'', Purple House Press,


External links

{{commons category, Sciurus
The Squirrel Lovers Club
an international club of squirrel enthusiasts.
Squirrel-Rehab.org
includes advice and links for helping orphaned baby squirrels. Squirrels Rodents by common name af:Boomeekhoring