A songbird is a

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

belonging to the

suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...

Passeri of the perching birds (

Passeriformes A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...

). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

''oscen'', "songbird". The Passeriformes contains 5,000 or so speciesEdwards, Scott V. and John Harshman. 2013. Passeriformes. Perching Birds, Passerine Birds. Version 06 February 2013 (under construction). http://tolweb.org/Passeriformes/15868/2013.02.06 in The Tree of Life Web Project, http://tolweb.org/ ccessed 2017/12/11 found all over the world, in which the vocal organ typically is developed in such a way as to produce a diverse and elaborate bird song. Songbirds form one of the two major lineages of extant perching birds (~4,000 species), the other being the Tyranni (~1,000 species), which are most diverse in the

Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeog ...

and absent from many parts of the world. The Tyranni have a simpler

syrinx In classical Greek mythology, Syrinx () was an Arcadian nymph and a follower of Artemis, known for her chastity. Being pursued by Pan, she fled into the river Ladon, and at her own request was metamorphosed into a reed from which Pan then mad ...

musculature, and while their vocalizations are often just as complex and striking as those of songbirds, they are altogether more mechanical sounding. There is a third perching bird lineage, the Acanthisitti of

New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...

, of which only two species remain alive today. Recent estimates indicate that songbirds originated 50 million years ago. The distribution of their basal lineages suggest that their origin and initial diversification occurred exclusively in the Australian continent and only about 40 million years ago, oscines started to colonize

Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

, and eventually the

Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...

Description

The song in this clade is essentially territorial, because it communicates the identity and whereabouts of an individual to other birds, and also signals sexual intentions. Sexual selection among songbirds is highly based on mimetic vocalization. Female preference has shown in some populations to be based on the extent of a male's song repertoire. The larger a male's repertoire, the more females a male individual attracts. It is not to be confused with bird calls that are used for alarms and contact and are especially important in birds that feed or migrate in flocks. While almost all living birds give calls of some sort, well-developed songs are only given by a few lineages outside the songbirds. And still, not all songbirds proffer a call that is distinctly melodious. Songbirds do, however, possess a highly developed vocal organ, the

, that enables their sonorous activity. This organ, also known as a song box, can be found where the windpipe meets diverging bronchial tubes which lead to the lungs. The organ is a solid, bony structure lined with a film of membranes which air passes through as the songbird calls. While the song boxes of songbirds vary in size and intricacy, this does not necessarily determine the songbird's ability to voice their song. Researchers believe this has more to do with the length of the windpipe. Other birds (especially non-passeriforms) sometimes have

songs A song is a musical composition performed by the human voice. The voice often carries the melody (a series of distinct and fixed pitches) using patterns of sound and silence. Songs have a structure, such as the common ABA form, and are usuall ...

to attract mates or hold territory, but these are usually simple and repetitive, lacking the variety of many oscine songs. The monotonous repetition of the

common cuckoo The cuckoo, common cuckoo, European cuckoo or Eurasian cuckoo (''Cuculus canorus'') is a member of the cuckoo order of birds, Cuculiformes, which includes the Geococcyx, roadrunners, the ani (bird), anis and the coucals. This species is a widesp ...

or little crake can be contrasted with the variety of a nightingale or marsh warbler. However, although many songbirds have songs that are pleasant to the human ear, this is not invariably the case. Many members of the

crow A crow is a bird of the genus ''Corvus'', or more broadly, a synonym for all of ''Corvus''. The word "crow" is used as part of the common name of many species. The related term "raven" is not linked scientifically to any certain trait but is rathe ...

family (

Corvidae Corvidae is a Cosmopolitan distribution, cosmopolitan Family (biology), family of Songbird, oscine passerine birds that contains the crows, ravens, Rook (bird), rooks, magpies, jackdaws, jays, treepies, choughs, and Nutcracker (bird), nutcrackers ...

) communicate with croaks or screeches, which sound harsh to humans. Even these, however, have a song of sorts, a softer twitter that is given between courting partners. And even though some

parrot Parrots (Psittaciformes), also known as psittacines (), are birds with a strong curved beak, upright stance, and clawed feet. They are classified in four families that contain roughly 410 species in 101 genus (biology), genera, found mostly in ...

s (which are not songbirds) can be taught to repeat human speech, vocal mimicry among birds is almost completely restricted to songbirds, some of which (such as the lyrebirds or the aptly named

mockingbird Mockingbirds are a group of New World passerine birds from the family (biology), family Mimidae. They are best known for the habit of some species Mimicry, mimicking the songs of other birds and the sounds of insects and amphibians, often loudly ...

s) excel in imitating the sounds of other birds or even environmental noises. The birds from higher altitudes have evolved thicker downs (also known as jackets) to protect themselves from colder temperatures. Their feathers have outer and inner portions, with the lower down being fluffier and warmer to provide increased warmth.

Song repertoire and courtship

Sexual selection can be broken down into several different studies regarding different aspects of a bird's song. As a result, songs can vary even within a single species. Many believe that song repertoire and cognition have a direct relationship. However, a study published in 2013 has shown that cognitive abilities may not all be directly related to the song repertoire of a songbird. Specifically, spatial learning is said to have an inverse relationship with song repertoire. So for example, this would be an individual who does not migrate as far as others in the species but has a better song repertoire. This suggests an evolutionary trade-off between possible alleles. With natural selection choosing traits best fit for reproductive success, there could be a trade-off in either direction depending on which trait would produce a higher fitness at that time period. Song repertoire can be attributed to male songbirds as it is one of the main mechanisms of courtship. Song repertoires differ from male individual to male individual and species to species. Some species may typically have large repertoires while others may have significantly smaller ones.

Mate choice Mate choice is one of the primary mechanisms under which evolution can occur. It is characterized by a "selective response by animals to particular stimuli" which can be observed as behavior.Bateson, Paul Patrick Gordon. "Mate Choice." Mate Choi ...

in female songbirds is a significant realm of study as song abilities are continuously evolving. Males often sing to assert their dominance over other males in competition for a female, sometimes in lieu of a combative episode, and to arouse the female by announcing a readiness to mate. Though less frequent, females have also been known to sing occasionally a duet with a mate as an affirmation of their partnership. While some will sing their song from a familiar perch, other species common to grasslands will sing a familiar song each time they fly. Currently, there have been numerous studies involving songbird repertoires, unfortunately, there has not yet been a concrete evidence to confirm that every songbird species prefers larger repertoires. A conclusion can be made that it can vary between species on whether a larger repertoire is connected to better fitness. With this conclusion, it can be inferred that evolution via natural selection, or sexual selection, favors the ability to retain larger repertoires for these certain species as it leads to higher reproductive success. During times of courtship, it is said that male songbirds increase their repertoire by mimicking other species songs. The better the mimicking ability, retaining ability, and the quantity of other species mimicked has been proven to have a positive relationship with mating success. Female preferences cause the constant improvement of accuracy and presentation of the copied songs. Another theory known as the "song-sharing hypothesis" suggests that females prefer simpler, more homogenous songs that signal a male of familiar territory. As birdsong can be broken into regional dialects through this process of mimicry, the foreign song of a newcomer suggests the lack of territorial possession. This can be costly in the wake of territorial conflicts between disparate songbird populations and may compel a female to prefer a male spouting a familiar song of the area.

Taxonomy and systematics

Sibley and Alquist divided songbirds into two " parvorders", Corvida and Passerida (standard taxonomic practice would rank these as

infraorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classific ...

s), distributed in Australo-Papua and

respectively.Selvatti, A.P. ''et al''. (2015
A Paleogene origin for crown passerines and the diversification of the Oscines in the New World
''Molecular Phylogenetics and Evolution'', 88:1-15. Subsequent molecular studies, however, show this treatment to be somewhat erroneous. Passerida is a highly diverse lineage, uniting over one-third of all bird species to include (in 2015) 3,885 species). These are divided into three major superfamilies (though not exactly corresponding to the Sibley-Ahlquist arrangement), in addition to some minor lineages. In contrast, Sibley & Alquist's "Corvida" is a phylogenetic grade and an artefact of the phenetic methodology. The bulk of the "Corvida" make up the large clade Corvides (812 species as of 2015), which is a

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to the Passerida. The remaining 15 oscine families (343 species in 2015) form a series of basally branching sister groups to the Corvoid - Passerid clade. All of these groups, which form at least six successively branching basal clades, are found exclusively or predominantly in Australasia. Australian endemics are also prominent among basal lineages in both Corvoids and Passerids, suggesting that songbirds originated and diverged in Australia. Scrubbirds and lyrebirds, of which there are just two species of each, represent the oldest lineage of songbirds on Earth. The rufous scrubbird, ''Atrichornis rufescens'', is essentially confined to the Gondwana Rainforests of Australia World Heritage Area, occurring in both Queensland and New South Wales sections. It is now only found at elevations above . One of the earliest known fossil songbirds is '' Resoviaornis'' from the

Early Oligocene The Rupelian is, in the geologic timescale, the older of two age (geology), ages or the lower of two stage (stratigraphy), stages of the Oligocene epoch (geology), Epoch/series (stratigraphy), Series. It spans the time between . It is preceded b ...

of Poland.

Families

* Menuroidea ** Menuridae: lyrebirds ** Atrichornithidae: scrub birds * Bowerbirds and Australian treecreepers ** Climacteridae: Australian treecreepers **

Ptilonorhynchidae Bowerbirds () make up the bird family (biology), family Ptilonorhynchidae. They are renowned for their unique courtship behaviour, where males build a structure and decorate it with sticks and brightly coloured objects in an attempt to attract a ...

bowerbird Bowerbirds () make up the bird family Ptilonorhynchidae. They are renowned for their unique courtship behaviour, where males build a structure and decorate it with sticks and brightly coloured objects in an attempt to attract a mate. The family ...

s * Meliphagoidea: honeyeaters and allies ** Maluridae: fairy-wrens, emu-wrens and grasswrens ** Meliphagidae: true

honeyeater The honeyeaters are a large and diverse family, Meliphagidae, of small to medium-sized birds. The family includes the Australian chats, myzomelas, friarbirds, wattlebirds, miners and melidectes. They are most common in Australia and New Gui ...

s and chats ** Dasyornithidae: bristlebirds ** Pardalotidae: pardalotes ** Acanthizidae: scrubwrens, thornbills, and gerygones * Australopapuan babblers ** Pomatostomidae: Australasian babblers * Logrunners ** Orthonychidae: logrunners * Other basal lineages ** Cnemophilidae: satinbirds '' Cnemophilus'' and '' Loboparadisea'' ** Melanocharitidae: berrypeckers and longbills ** Callaeidae: New Zealand wattlebirds kōkako, saddleback and † huia ** Notiomystidae: stitchbird * Corvides ** Paramythiidae: tit berrypeckers and crested berrypeckers ** Psophodidae: whipbirds, jewel-babblers and quail-thrushes ** Platysteiridae: wattle-eyes and batis ** Malaconotidae: bush-shrikes ** Machaerirynchidae: boatbills **

Vangidae The family Vangidae (from ''vanga'', Malagasy language, Malagasy for the hook-billed vanga, ''Vanga curvirostris'') comprises a group of often shrike-like medium-sized birds distributed from Asia to Africa, including the vangas of Madagascar to ...

: vangas, woodshrikes, and helmetshrikes ** Pityriasidae: Bornean bristlehead ** Artamidae: butcherbirds, currawongs and Australian magpie (formerly in Cracticidae) ** Rhagologidae: mottled whistlers ** Aegithinidae: ioras ** Campephagidae: cuckooshrikes and trillers ** Mohouidae: whiteheads ** Neosittidae: sittellas ** Eulacestomidae: ploughbills ** Oreoicidae: Australo-Papuan bellbirds ** Pachycephalidae: whistlers, shrike-thrushes, pitohuis and allies ** Laniidae: shrikes **

Vireonidae The vireos make up a family (biology), family, Vireonidae, of small to medium-sized passerine birds found in the New World (Canada to Argentina, including Bermuda and the West Indies) and Southeast Asia. The family contains 62 species and is d ...

: vireos ** Oriolidae: orioles, figbirds and † piopio (formerly Turnagridae) ** Dicruridae: drongos ** Rhipiduridae: fantails ** Monarchidae:

monarch A monarch () is a head of stateWebster's II New College Dictionary. "Monarch". Houghton Mifflin. Boston. 2001. p. 707. Life tenure, for life or until abdication, and therefore the head of state of a monarchy. A monarch may exercise the highest ...

s and allies ** Platylophidae: jayshrikes **

magpie Magpies are birds of various species of the family Corvidae. Like other members of their family, they are widely considered to be intelligent creatures. The Eurasian magpie, for instance, is thought to rank among the world's most intelligent c ...

s, and jays ** Corcoracidae: white-winged chough and apostlebird ** Melampittidae: melampittas ** Ifritidae: ifritabirds ** Paradisaeidae: birds of paradise * Passerida ** Petroicidae: Australasian robins ** Picathartidae: rockfowl ** Chaetopidae: rockjumpers ** Eupetidae: rail-babbler ** Bombycillidae: waxwings and allies ** Ptiliogonatidae: silky-flycatchers ** Hypocoliidae: hypocolius ** Dulidae: palmchat ** † Mohoidae: some Hawaiian honeyeaters, '' Moho'' and '' Chaetoptila'' genera, not closely related to Meliphagidae ** Hylocitreidae: hylocitrea ** Stenostiridae: fairy-flycatcher and allies ** Paridae: tits, chickadees, and titmouse ** Remizidae: penduline-tits ** Nicatoridae: nicators ** Panuridae: bearded reedling ** Alaudidae:

lark Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...

s ** Pycnonotidae:

bulbul The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropic ...

s ** Hirundinidae:

swallow The swallows, martins, and saw-wings, or Hirundinidae are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica. Highly adapted to aerial feeding, they have a distinctive appearance. The ...

s and martins ** Pnoepygidae: wren-babblers ** Macrosphenidae: crombecs and African warblers ** Cettiidae: bush-warblers and allies ** Scotocercidae: streaked scrub-warbler **

Erythrocercidae ''Erythrocercus'' is a genus of birds containing three flycatchers that are found in Africa. The genus is placed in its own family (biology), family Erythrocercidae that was introduced by Silke Fregin and collaborators in 2012. Species The ge ...

: yellow flycatchers ** Aegithalidae: long-tailed tits ** Phylloscopidae: leaf-warblers and allies. Recently split from Sylviidae. ** Acrocephalidae: reed warblers and allies ** Locustellidae: grassbirds and allies ** Donacobiidae: black-capped donacobius ** Bernieridae: Malagasy warblers **

Cisticolidae The family Cisticolidae is a group of about 160 warblers, small passerine birds found mainly in warmer southern regions of the Old World. They were formerly included within the Old World warbler family Sylviidae. This family probably originated ...

: cisticolas and allies ** Timaliidae: babblers ** Pellorneidae: ground babblers ** Leiothrichidae: laughingthrushes and allies ** Sylviidae:

Old World warbler The Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. They are not closely related to the New World warblers. The family held over 400 species in over 70 genera, and were the source of much taxo ...

s **

Zosteropidae The white-eyes are a family, Zosteropidae, of small passerine birds native to tropical, subtropical and temperate Sub-Saharan Africa, southern and eastern Asia, and Australasia. White-eyes inhabit most tropical islands in the Indian Ocean, the ...

: white-eyes ** Arcanatoridae: dapple-throat and allies ** Promeropidae: sugarbirds ** Irenidae: fairy-bluebirds ** Regulidae: kinglets ** Elachuridae: elachuras ** Hyliotidae: hyliotas ** Troglodytidae: wrens ** Polioptilidae: gnatcatchers ** Sittidae:

nuthatch The nuthatches () constitute a genus, ''Sitta'', of small passerine birds belonging to the family Sittidae. Characterised by large heads, short tails, and powerful bills and feet, nuthatches advertise their territory using loud, simple songs. Mo ...

es ** Tichodromidae: wallcreeper ** Certhiidae: treecreepers ** Mimidae:

s and thrashers ** Sturnidae:

starling Starlings are small to medium-sized passerine (perching) birds known for the often dark, glossy iridescent sheen of their plumage; their complex vocalizations including mimicking; and their distinctive, often elaborate swarming behavior, know ...

s ** Buphagidae: oxpeckers ** Turdidae: thrushes and allies **

Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...

: Old World flycatchers and chats ** Cinclidae: dippers ** Chloropseidae:

leafbird The leafbirds (Chloropseidae) are a family of small passerine bird species found in the Indian Subcontinent and Southeast Asia. They were formerly grouped with the ioras and fairy-bluebirds in the family Irenidae. As presently defined, the leaf ...

s ** Dicaeidae: flowerpeckers ** Nectariniidae: sunbirds **

Passeridae Old World sparrows are a group of small passerine birds forming the family Passeridae. They are also known as true sparrows, a name also used for a particular genus of the family, '' Passer''. They are distinct from both the New World sparrows, ...

: true sparrows **

Ploceidae Ploceidae is a family of small passerine birds, many of which are called weavers, weaverbirds, weaver finches, or bishops. These names come from the nests of intricately woven vegetation created by birds in this family. In most recent classificat ...

: weavers and widowbirds ** Estrildidae: estrildid finches ( waxbills, munias, etc.) ** Viduidae: indigo birds and whydahs ** Peucedramidae: olive warbler ** Prunellidae: accentor **

Motacillidae The wagtails, longclaws, and pipits are a family (biology), family, Motacillidae, of small passerine birds with medium to long tails. Around 70 species occur in five genus, genera. The longclaws are entirely restricted to the Afrotropics, and the ...

: wagtails and pipits ** Urocynchramidae: Przevalski's finch ** Fringillidae: true finches and Hawaiian honeycreepers (formerly Drepanididae) **

Parulidae The New World warblers or wood-warblers are a group of small, often colorful, passerine birds that make up the family Parulidae and are restricted to the New World. The family contains 120 species. They are not closely related to Old World warb ...

: New World warblers, for example the black-throated blue warblers and allies ** Icteridae: American blackbirds,

New World oriole New World orioles are a group of birds in the genus ''Icterus'' of the Icterid, blackbird family. Although they are not closely related to Old World orioles of the family Oriolidae, they are strikingly similar in size, diet, behavior, and strong ...

s, grackles and cowbirds. ** Coerebidae: bananaquit ** Emberizidae:

buntings Bunting may refer to: Birds * ''Emberiza'', a group of Old World passerine birds * ''Passerina'', a group of birds in the Cardinalidae family known as the North American buntings * Blue bunting, ''Cyanocompsa parellina'' * Lark bunting, ''Calam ...

** Passerellidae: New World sparrows **

Thraupidae The tanagers (singular ) comprise the bird family (biology), family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12 ...

: tanagers, true honeycreepers and allies **

Calcariidae Calcariidae is a small family (biology), family of passerine birds. It includes longspurs and snow buntings. There are six species in three genera worldwide, found mainly in North America and Eurasia. They are migratory and can live in a variety ...

: snow buntings and longspurs **

Cardinalidae Cardinalidae (sometimes referred to as "cardinal-grosbeaks" or simply "cardinals") is a family of New World-Endemism, endemic passerine birds that consists of Cardinalis, cardinals, grosbeaks, and Passerina, buntings. It also includes several ot ...

: cardinals and allies

References

External links