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Feces (also known as faeces or fæces; : faex) are the solid or semi-solid remains of food that was not digested in the
small intestine The small intestine or small bowel is an organ (anatomy), organ in the human gastrointestinal tract, gastrointestinal tract where most of the #Absorption, absorption of nutrients from food takes place. It lies between the stomach and large intes ...
, and has been broken down by
bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...
in the
large intestine The large intestine, also known as the large bowel, is the last part of the gastrointestinal tract and of the Digestion, digestive system in tetrapods. Water is absorbed here and the remaining waste material is stored in the rectum as feces befor ...
. Feces contain a relatively small amount of
metabolic waste Metabolic wastes or excrements are substances left over from metabolic processes (such as cellular respiration) which cannot be used by the organism (they are surplus or toxic), and must therefore be excreted. This includes nitrogen compounds ...
products such as bacterially-altered
bilirubin Bilirubin (BR) (adopted from German, originally bili—bile—plus ruber—red—from Latin) is a red-orange compound that occurs in the normcomponent of the straw-yellow color in urine. Another breakdown product, stercobilin, causes the brown ...
and dead epithelial cells from the lining of the gut. Feces are discharged through the
anus In mammals, invertebrates and most fish, the anus (: anuses or ani; from Latin, 'ring' or 'circle') is the external body orifice at the ''exit'' end of the digestive tract (bowel), i.e. the opposite end from the mouth. Its function is to facil ...
or
cloaca A cloaca ( ), : cloacae ( or ), or vent, is the rear orifice that serves as the only opening for the digestive (rectum), reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles, birds, cartilagin ...
during
defecation Defecation (or defaecation) follows digestion and is the necessary biological process by which organisms eliminate a solid, semisolid, or liquid metabolic waste, waste material known as feces (or faeces) from the digestive tract via the anus o ...
. Feces can be used as
fertilizer A fertilizer or fertiliser is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from liming materials or other non-nutrient soil amendments. Man ...
or
soil conditioner A soil conditioner is a product which is added to soil to improve the soil’s physical qualities, usually its fertility (ability to provide nutrition for plants) and sometimes its mechanics. In general usage, the term "soil conditioner" is often ...
in agriculture. They can also be burned as
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work (physics), work. The concept was originally applied solely to those materials capable of releasing chem ...
or dried and used for
construction Construction are processes involved in delivering buildings, infrastructure, industrial facilities, and associated activities through to the end of their life. It typically starts with planning, financing, and design that continues until the a ...
. Some
medicinal Medicine is the science and Praxis (process), practice of caring for patients, managing the Medical diagnosis, diagnosis, prognosis, Preventive medicine, prevention, therapy, treatment, Palliative care, palliation of their injury or disease, ...
uses have been found. In the case of
human feces Human feces (American English) or faeces (British English), commonly and in medical literature more often called stool, are the solid or semisolid remains of food that could not be digested or absorbed in the small intestine of humans, but has b ...
, fecal transplants or
fecal bacteriotherapy Fecal microbiota transplant (FMT), also known as a stool transplant, is the process of transferring fecal bacteria and other microbes from a healthy individual into another individual. FMT is an effective treatment for Clostridioides diffic ...
are in use.
Urine Urine is a liquid by-product of metabolism in humans and many other animals. In placental mammals, urine flows from the Kidney (vertebrates), kidneys through the ureters to the urinary bladder and exits the urethra through the penile meatus (mal ...
and feces together are called
excreta Excretion is elimination of metabolic waste, which is an essential process in all organisms. In vertebrates, this is primarily carried out by the lungs, kidneys, and skin. This is in contrast with secretion, where the substance may have specifi ...
.


Characteristics

The distinctive
odor An odor (American English) or odour ( Commonwealth English; see spelling differences) is a smell or a scent caused by one or more volatilized chemical compounds generally found in low concentrations that humans and many animals can perceive ...
of feces is due to
skatole Skatole or 3-methylindole is an organic compound belonging to the indole family. It occurs naturally in the feces of mammals and birds and is the primary contributor to fecal odor. In low concentrations, it has a flowery smell and is found in sever ...
, and
thiol In organic chemistry, a thiol (; ), or thiol derivative, is any organosulfur compound of the form , where R represents an alkyl or other organic substituent. The functional group itself is referred to as either a thiol group or a sulfhydryl grou ...
s (
sulfur Sulfur ( American spelling and the preferred IUPAC name) or sulphur ( Commonwealth spelling) is a chemical element; it has symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms ...
-containing compounds), as well as amines and carboxylic acids. Skatole is produced from
tryptophan Tryptophan (symbol Trp or W) is an α-amino acid that is used in the biosynthesis of proteins. Tryptophan contains an α-amino group, an α-carboxylic acid group, and a side chain indole, making it a polar molecule with a non-polar aromat ...
via indoleacetic acid.
Decarboxylation Decarboxylation is a chemical reaction that removes a carboxyl group and releases carbon dioxide (CO2). Usually, decarboxylation refers to a reaction of carboxylic acids, removing a carbon atom from a carbon chain. The reverse process, which is ...
gives skatole. The perceived bad odor of feces has been hypothesized to be a deterrent for humans, as consuming or touching it may result in sickness or infection.


Physiology

Feces are discharged through the
anus In mammals, invertebrates and most fish, the anus (: anuses or ani; from Latin, 'ring' or 'circle') is the external body orifice at the ''exit'' end of the digestive tract (bowel), i.e. the opposite end from the mouth. Its function is to facil ...
or
cloaca A cloaca ( ), : cloacae ( or ), or vent, is the rear orifice that serves as the only opening for the digestive (rectum), reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles, birds, cartilagin ...
during
defecation Defecation (or defaecation) follows digestion and is the necessary biological process by which organisms eliminate a solid, semisolid, or liquid metabolic waste, waste material known as feces (or faeces) from the digestive tract via the anus o ...
. This process requires pressures that may reach (13.3 kPa) in humans and (60 kPa) in penguins. The forces required to expel the feces are generated through muscular contractions and a build-up of gases inside the gut, prompting the sphincter to relieve the pressure and release the feces.


Ecology

After an animal has digested eaten material, the remains of that material are discharged from its body as waste. Although it is lower in energy than the food from which it is derived, feces may retain a large amount of energy, often 50% of that of the original food. This means that of all food eaten, a significant amount of energy remains for the decomposers of ecosystems. Many organisms feed on feces, from bacteria to fungi to insects such as
dung beetle Dung beetles are beetles that feed on feces. All species of dung beetle belong to the superfamily Scarabaeoidea, most of them to the subfamilies Scarabaeinae and Aphodiinae of the family Scarabaeidae (scarab beetles). As most species of Scara ...
s, who can
sense A sense is a biological system used by an organism for sensation, the process of gathering information about the surroundings through the detection of Stimulus (physiology), stimuli. Although, in some cultures, five human senses were traditio ...
odors from long distances. Some may specialize in feces, while others may eat other foods. Feces serve not only as a basic food, but also as a supplement to the usual diet of some animals. This process is known as
coprophagia Coprophagia ( ) or coprophagy ( ) is the consumption of feces. The word is derived from the Ancient Greek "feces" and "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of o ...
, and occurs in various animal species such as young elephants eating the feces of their mothers to gain essential
gut flora Gut microbiota, gut microbiome, or gut flora are the microorganisms, including bacteria, archaea, fungi, and viruses, that live in the digestive tracts of animals. The gastrointestinal metagenome is the aggregate of all the genomes of the g ...
, or by other animals such as dogs, rabbits, and monkeys. Feces and urine, which reflect
ultraviolet Ultraviolet radiation, also known as simply UV, is electromagnetic radiation of wavelengths of 10–400 nanometers, shorter than that of visible light, but longer than X-rays. UV radiation is present in sunlight and constitutes about 10% of ...
light, are important to raptors such as
kestrel The term kestrel (from , derivative from , i.e. ratchet) is the common name given to several species of predatory birds from the falcon genus ''Falco''. Kestrels are most easily distinguished by their typical hunting behaviour which is to hover ...
s, who can see the
near ultraviolet Ultraviolet radiation, also known as simply UV, is electromagnetic radiation of wavelengths of 10–400 nanometers, shorter than that of visible light, but longer than X-rays. UV radiation is present in sunlight and constitutes about 10% of th ...
and thus find their prey by their
midden A midden is an old dump for domestic waste. It may consist of animal bones, human excrement, botanical material, mollusc shells, potsherds, lithics (especially debitage), and other artifacts and ecofacts associated with past human oc ...
s and
territorial A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
markers.
Seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s also may be found in feces. Animals who eat
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
are known as
frugivore A frugivore ( ) is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance ...
s. An advantage for a plant in having fruit is that animals will eat the fruit and unknowingly disperse the seed in doing so. This mode of
seed dispersal In spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
is highly successful, as seeds dispersed around the base of a plant are unlikely to succeed and often are subject to heavy
predation Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
. Provided the seed can withstand the pathway through the digestive system, it is not only likely to be far away from the parent plant, but is even provided with its own fertilizer. Organisms that subsist on dead organic matter or ''
detritus In biology, detritus ( or ) is organic matter made up of the decomposition, decomposing remains of organisms and plants, and also of feces. Detritus usually hosts communities of microorganisms that colonize and decomposition, decompose (Reminera ...
'' are known as
detritivore Detritivores (also known as detrivores, detritophages, detritus feeders or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrates, ...
s, and play an important role in ecosystems by recycling organic matter back into a simpler form that plants and other
autotroph An autotroph is an organism that can convert Abiotic component, abiotic sources of energy into energy stored in organic compounds, which can be used by Heterotroph, other organisms. Autotrophs produce complex organic compounds (such as carbohy ...
s may absorb once again. This cycling of matter is known as the
biogeochemical cycle A biogeochemical cycle, or more generally a cycle of matter, is the movement and transformation of chemical elements and compounds between living organisms, the atmosphere, and the Earth's crust. Major biogeochemical cycles include the carbon cyc ...
. To maintain nutrients in soil it is therefore important that feces returns to the area from which they came, which is not always the case in human society where food may be transported from rural areas to urban populations and then feces disposed of into a river or sea.


Human feces

Depending on the individual and the circumstances, human beings may defecate several times a day, every day, or once every two or three days. Extensive hardening of the feces that interrupts this routine for several days or more is called
constipation Constipation is a bowel dysfunction that makes bowel movements infrequent or hard to pass. The Human feces, stool is often hard and dry. Other symptoms may include abdominal pain, bloating, and feeling as if one has not completely passed the ...
. The appearance of human fecal matter varies according to diet and health. Normally it is semisolid, with a
mucus Mucus (, ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both Serous fluid, serous and muc ...
coating. A combination of
bile Bile (from Latin ''bilis''), also known as gall, is a yellow-green/misty green fluid produced by the liver of most vertebrates that aids the digestion of lipids in the small intestine. In humans, bile is primarily composed of water, is pro ...
and
bilirubin Bilirubin (BR) (adopted from German, originally bili—bile—plus ruber—red—from Latin) is a red-orange compound that occurs in the normcomponent of the straw-yellow color in urine. Another breakdown product, stercobilin, causes the brown ...
, which comes from dead
red blood cells Red blood cells (RBCs), referred to as erythrocytes (, with -''cyte'' translated as 'cell' in modern usage) in academia and medical publishing, also known as red cells, erythroid cells, and rarely haematids, are the most common type of blood cel ...
, gives feces the typical brown color. After the
meconium Meconium is the earliest stool of a mammalian infant resulting from defecation. Unlike later feces, meconium is composed of materials ingested during the time the infant spends in the uterus: intestinal epithelial cells, lanugo, mucus, am ...
, the first stool expelled, a newborn's feces contains only
bile Bile (from Latin ''bilis''), also known as gall, is a yellow-green/misty green fluid produced by the liver of most vertebrates that aids the digestion of lipids in the small intestine. In humans, bile is primarily composed of water, is pro ...
, which gives it a yellow-green color.
Breast feeding Breastfeeding, also known as nursing, is the process where breast milk is fed to a child. Infants may suck the milk directly from the breast, or milk may be extracted with a pump and then fed to the infant. The World Health Organization (WH ...
babies expel soft, pale yellowish, and not quite malodorous matter; but once the baby begins to eat, and the body starts expelling bilirubin from dead red blood cells, its matter acquires the familiar brown color. At different times in their life, human beings will expel feces of different colors and textures. A stool that passes rapidly through the intestines will look greenish; lack of bilirubin will make the stool look like clay.


Uses of animal feces


Fertilizer

The feces of animals, e.g.
guano Guano (Spanish from ) is the accumulated excrement of seabirds or bats. Guano is a highly effective fertiliser due to the high content of nitrogen, phosphate, and potassium, all key nutrients essential for plant growth. Guano was also, to a le ...
and
manure Manure is organic matter that is used as organic fertilizer in agriculture. Most manure consists of animal feces; other sources include compost and green manure. Manures contribute to the fertility of soil by adding organic matter and nut ...
, often are used as
fertilizer A fertilizer or fertiliser is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from liming materials or other non-nutrient soil amendments. Man ...
.


Energy

Dry animal dung, such as that of
camel A camel (from and () from Ancient Semitic: ''gāmāl'') is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as livestock, they provid ...
,
bison A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised. Of the two surviving species, the American ...
and
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
, is burned as fuel in many countries. Animals such as the
giant panda The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear or simply panda, is a bear species endemic to China. It is characterised by its white animal coat, coat with black patches around the eyes, ears, legs and shoulders. ...
and
zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: Grévy's zebra (''Equus grevyi''), the plains zebra (''E. quagga''), and the mountain zebra (''E. ...
possess gut bacteria capable of producing biofuel. The bacterium in question, Brocadia anammoxidans, can be used to synthesize the rocket fuel
hydrazine Hydrazine is an inorganic compound with the chemical formula . It is a simple pnictogen hydride, and is a colourless flammable liquid with an ammonia-like odour. Hydrazine is highly hazardous unless handled in solution as, for example, hydraz ...
.


Coprolites and paleofeces

A
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name ...
is
fossilized A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
feces and is classified as a
trace fossil A trace fossil, also called an ichnofossil (; ), is a fossil record of biological activity by lifeforms, but not the preserved remains of the organism itself. Trace fossils contrast with body fossils, which are the fossilized remains of part ...
. In
paleontology Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure ge ...
they give evidence about the diet of an animal. They were first described by
William Buckland William Buckland Doctor of Divinity, DD, Royal Society, FRS (12 March 1784 – 14 August 1856) was an English theologian, geologist and paleontology, palaeontologist. His work in the early 1820s proved that Kirkdale Cave in North Yorkshire h ...
in 1829. Prior to this, they were known as "fossil fir cones" and "
bezoar A bezoar stone ( ) is a mass often found trapped in the gastrointestinal system, though it can occur in other locations. A pseudobezoar is an indigestible object introduced intentionally into the digestive system. There are several varieties o ...
stones". They serve a valuable purpose in paleontology because they provide direct evidence of the predation and diet of extinct organisms. Coprolites may range in size from a few millimetres to more than 60 centimetres. Palaeofeces are ancient
feces Feces (also known as faeces American and British English spelling differences#ae and oe, or fæces; : faex) are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the ...
, often found as part of
archaeological Archaeology or archeology is the study of human activity through the recovery and analysis of material culture. The archaeological record consists of Artifact (archaeology), artifacts, architecture, biofact (archaeology), biofacts or ecofacts, ...
excavations or surveys. Intact paleofeces of ancient people may be found in caves in arid climates and in other locations with suitable preservation conditions. These are studied to determine the
diet Diet may refer to: Food * Diet (nutrition), the sum of the food consumed by an organism or group * Dieting, the deliberate selection of food to control body weight or nutrient intake ** Diet food, foods that aid in creating a diet for weight loss ...
and
health Health has a variety of definitions, which have been used for different purposes over time. In general, it refers to physical and emotional well-being, especially that associated with normal functioning of the human body, absent of disease, p ...
of the people who produced them through the analysis of seeds, small bones, and parasite eggs found inside. Feces may contain information about the person excreting the material as well as information about the material. They also may be analyzed chemically for more in-depth information on the individual who excreted them, using
lipid Lipids are a broad group of organic compounds which include fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids include storing ...
analysis and ancient
DNA Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...
analysis. The success rate of usable DNA extraction is relatively high in paleofeces, making it more reliable than skeletal DNA retrieval. The reason this analysis is possible at all is due to the digestive system not being entirely efficient, in the sense that not everything that passes through the digestive system is destroyed. Not all of the surviving material is recognizable, but some of it is. Generally, this material is the best indicator archaeologists can use to determine ancient diets, as no other part of the archaeological record is so direct an indicator. A process that preserves feces in a way that they may be analyzed later is the Maillard reaction. This reaction creates a casing of sugar that preserves the feces from the elements. To extract and analyze the information contained within, researchers generally have to freeze the feces and grind it up into powder for analysis.


Other uses

Animal dung occasionally is used as a
cement A cement is a binder, a chemical substance used for construction that sets, hardens, and adheres to other materials to bind them together. Cement is seldom used on its own, but rather to bind sand and gravel ( aggregate) together. Cement mi ...
to make
adobe Adobe (from arabic: الطوب Attub ; ) is a building material made from earth and organic materials. is Spanish for mudbrick. In some English-speaking regions of Spanish heritage, such as the Southwestern United States, the term is use ...
(
mudbrick Mudbrick or mud-brick, also known as unfired brick, is an air-dried brick, made of a mixture of mud (containing loam, clay, sand and water) mixed with a binding material such as rice husks or straw. Mudbricks are known from 9000 BCE. From ...
) huts, or even in throwing sports, especially with cow and camel dung.
Kopi luwak Kopi luwak, also known as civet coffee, is a coffee that consists of partially digested coffee cherry, coffee cherries, which have been eaten and defecation, defecated by the Asian palm civet (''Paradoxurus hermaphroditus''). The cherries are Fer ...
, or civet coffee, is coffee made from coffee beans that have been eaten and excreted by
Asian palm civet The Asian palm civet (''Paradoxurus hermaphroditus''), also called common palm civet, toddy cat and musang, is a viverrid native to South and Southeast Asia. Since 2008, it is IUCN Red Listed as Least Concern as it accommodates to a broad range o ...
s (''Paradoxurus hermaphroditus'').
Giant pandas The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear or simply panda, is a bear species endemic to China. It is characterised by its white coat with black patches around the eyes, ears, legs and shoulders. Its body is ...
provide fertilizer for the world's most expensive
green tea Green tea is a type of tea made from the leaves and buds of the '' Camellia sinensis'' that have not undergone the withering and oxidation process that creates oolong teas and black teas. Green tea originated in China in the late 1st millenn ...
. In
Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
, tea is made from the droppings of
stick insect The Phasmatodea (also known as Phasmida or Phasmatoptera) are an order of insects whose members are variously known as stick insects, stick bugs, walkingsticks, stick animals, or bug sticks. They are also occasionally referred to as Devil's da ...
s fed on
guava Guava ( ), also known as the 'guava-pear', is a common tropical fruit cultivated in many tropical and subtropical regions. The common guava '' Psidium guajava'' (lemon guava, apple guava) is a small tree in the myrtle family (Myrtaceae), nativ ...
leaves. In northern
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
,
elephant Elephants are the largest living land animals. Three living species are currently recognised: the African bush elephant ('' Loxodonta africana''), the African forest elephant (''L. cyclotis''), and the Asian elephant ('' Elephas maximus ...
s are used to digest coffee beans in order to make Black Ivory coffee, which is among the world's most expensive coffees.
Paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, Textile, rags, poaceae, grasses, Feces#Other uses, herbivore dung, or other vegetable sources in water. Once the water is dra ...
is also made from elephant dung in Thailand. Haathi Chaap is a brand of paper made from elephant dung.
Dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
feces was used in the tanning process of
leather Leather is a strong, flexible and durable material obtained from the tanning (leather), tanning, or chemical treatment, of animal skins and hides to prevent decay. The most common leathers come from cattle, sheep, goats, equine animals, buffal ...
during the
Victorian era In the history of the United Kingdom and the British Empire, the Victorian era was the reign of Queen Victoria, from 20 June 1837 until her death on 22 January 1901. Slightly different definitions are sometimes used. The era followed the ...
. Collected dog feces, known as "pure", "puer", or "pewer", were mixed with water to form a substance known as "bate", because
proteolytic Proteolysis is the breakdown of proteins into smaller polypeptides or amino acids. Protein degradation is a major regulatory mechanism of gene expression and contributes substantially to shaping mammalian proteomes. Uncatalysed, the hydrolysis o ...
enzyme An enzyme () is a protein that acts as a biological catalyst by accelerating chemical reactions. The molecules upon which enzymes may act are called substrate (chemistry), substrates, and the enzyme converts the substrates into different mol ...
s in the dog feces helped to relax the fibrous structure of the hide before the final stages of tanning. Dog feces collectors were known as ''pure finders''. Elephants,
hippos A hippo or hippopotamus is either of two species of large African mammal which live mainly in and near water: * Hippopotamus * Pygmy hippopotamus Hippo or Hippos may also refer to: Toponymy * The ancient city of Hippo Regius (modern Annaba, Alg ...
,
koala The koala (''Phascolarctos cinereus''), sometimes inaccurately called the koala bear, is an arboreal herbivorous marsupial native to Australia. It is the only Extant taxon, extant representative of the Family (biology), family ''Phascolar ...
s and pandas are born with sterile intestines, and require bacteria obtained from eating the feces of their mothers to digest vegetation. In India,
cow dung Cow dung, also known as cow pats, cow poop or cow manure, is the waste product (faeces) of bovine animal species. These species include domestic cattle ("cows"), bison ("buffalo"), yak, and water buffalo. Cow dung is the undigested residue of p ...
and
cow urine Cow urine, gomutra or gōmēz is a liquid by-product of metabolism in cows. It has a sacred role in Zoroastrianism and some forms of Hinduism. Urophagia, the consumption of urine, was used in several ancient cultures for various health, heal ...
are major ingredients of the traditional
Hindu Hindus (; ; also known as Sanātanīs) are people who religiously adhere to Hinduism, also known by its endonym Sanātana Dharma. Jeffery D. Long (2007), A Vision for Hinduism, IB Tauris, , pp. 35–37 Historically, the term has also be ...
drink ''
Panchagavya Panchagavya or panchakavyam is a mixture used in traditional Hindu rituals that is prepared by mixing five ingredients. The three direct constituents are cow dung, cow urine, and milk; the two derived products are curd and ghee. These are mixe ...
''. Politician Shankarbhai Vegad stated that they can cure
cancer Cancer is a group of diseases involving Cell growth#Disorders, abnormal cell growth with the potential to Invasion (cancer), invade or Metastasis, spread to other parts of the body. These contrast with benign tumors, which do not spread. Po ...
.


Terminology

''Feces'' is the scientific terminology, while the term ''stool'' is also commonly used in medical contexts. Outside of scientific contexts, these terms are less common, with the most common layman's term being ''poop'' or ''poo''. The term ''
shit ''Shit'' is an English-language profanity. As a noun, it refers to fecal matter, and as a verb it means to defecate; in the plural ("the shits"), it means diarrhea. ''Shite'' is a common variant in British and Irish English. As a slang ...
'' is also in common use, although it is widely considered vulgar or offensive. There are many other terms, see below.


Etymology

The word ''faeces'' is the plural of the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word '' faex'' meaning "dregs". In most English-language
usage The usage of a language is the ways in which its written and spoken variations are routinely employed by its speakers; that is, it refers to "the collective habits of a language's native speakers", as opposed to idealized models of how a languag ...
, there is no singular form, making the word a
plurale tantum A ; ) is a noun that appears only in the plural form and does not have a singular variant for referring to a single object. In a less strict usage of the term, it can also refer to nouns whose singular form is rarely used. In English, are oft ...
; out of various major dictionaries, only one enters variation from plural agreement.


Synonyms

"Feces" is used more in biology and medicine than in other fields (reflecting
science Science is a systematic discipline that builds and organises knowledge in the form of testable hypotheses and predictions about the universe. Modern science is typically divided into twoor threemajor branches: the natural sciences, which stu ...
's tradition of classical Latin and
Neo-Latin Neo-LatinSidwell, Keith ''Classical Latin-Medieval Latin-Neo Latin'' in ; others, throughout. (also known as New Latin and Modern Latin) is the style of written Latin used in original literary, scholarly, and scientific works, first in Italy d ...
) *In hunting and tracking, terms such as ''dung'', ''scat'', ''spoor'', and ''droppings'' normally are used to refer to non-human animal feces *In husbandry and farming, ''
manure Manure is organic matter that is used as organic fertilizer in agriculture. Most manure consists of animal feces; other sources include compost and green manure. Manures contribute to the fertility of soil by adding organic matter and nut ...
'' is common. *''Stool'' is a common term in reference to
human feces Human feces (American English) or faeces (British English), commonly and in medical literature more often called stool, are the solid or semisolid remains of food that could not be digested or absorbed in the small intestine of humans, but has b ...
. For example, in
medicine Medicine is the science and Praxis (process), practice of caring for patients, managing the Medical diagnosis, diagnosis, prognosis, Preventive medicine, prevention, therapy, treatment, Palliative care, palliation of their injury or disease, ...
, to diagnose the presence or absence of a medical condition, a
stool sample A stool test is a medical diagnostic technique that involves the collection and analysis of fecal matter. Microbial analysis (culturing), microscopy and chemical tests are among the tests performed on stool samples. Collection Stool samples shou ...
sometimes is requested for testing purposes. *The term ''bowel movement(s)'' (with each movement a defecation event) is also common in
health care Health care, or healthcare, is the improvement or maintenance of health via the preventive healthcare, prevention, diagnosis, therapy, treatment, wikt:amelioration, amelioration or cure of disease, illness, injury, and other disability, physic ...
. There are many synonyms in informal registers for feces, just like there are for
urine Urine is a liquid by-product of metabolism in humans and many other animals. In placental mammals, urine flows from the Kidney (vertebrates), kidneys through the ureters to the urinary bladder and exits the urethra through the penile meatus (mal ...
. Many are euphemistic,
colloquial Colloquialism (also called ''colloquial language'', ''colloquial speech'', ''everyday language'', or ''general parlance'') is the linguistic style used for casual and informal communication. It is the most common form of speech in conversation amo ...
, or both; some are profane (such as ''
shit ''Shit'' is an English-language profanity. As a noun, it refers to fecal matter, and as a verb it means to defecate; in the plural ("the shits"), it means diarrhea. ''Shite'' is a common variant in British and Irish English. As a slang ...
''), whereas most belong chiefly to
child-directed speech Baby talk is a type of speech associated with an older person speaking to a child or infant. It is also called caretaker speech, infant-directed speech (IDS), child-directed speech (CDS), child-directed language (CDL), caregiver register, parente ...
) or to crude humor (such as ''crap'', ''dump'', ''load'' and ''turd'').


Feces of animals

The feces of animals often have special names (some of them are slang), for example: *Non-human animals **As bulk material – dung **Individually – droppings *
Cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
**Bulk material –
cow dung Cow dung, also known as cow pats, cow poop or cow manure, is the waste product (faeces) of bovine animal species. These species include domestic cattle ("cows"), bison ("buffalo"), yak, and water buffalo. Cow dung is the undigested residue of p ...
**Individual droppings – cow pats, meadow muffins, etc. *
Deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
(and formerly other quarry animals) – fewmets * Wild
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they ar ...
s – scat *
Otter Otters are carnivorous mammals in the subfamily Lutrinae. The 13 extant otter species are all semiaquatic, aquatic, or marine. Lutrinae is a branch of the Mustelidae family, which includes weasels, badgers, mink, and wolverines, among ...
spraint *
Bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s (individual) – droppings (also include urine as white crystals of
uric acid Uric acid is a heterocyclic compound of carbon, nitrogen, oxygen, and hydrogen with the Chemical formula, formula C5H4N4O3. It forms ions and salts known as urates and acid urates, such as ammonium acid urate. Uric acid is a product of the meta ...
) *
Seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
s or
bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s (large accumulations) –
guano Guano (Spanish from ) is the accumulated excrement of seabirds or bats. Guano is a highly effective fertiliser due to the high content of nitrogen, phosphate, and potassium, all key nutrients essential for plant growth. Guano was also, to a le ...
*Herbivorous
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s, such as
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s and
leaf beetle The beetle family Chrysomelidae, commonly known as leaf beetles, includes over 37,000 (and probably at least 50,000) species in more than 2,500 genera, making it one of the largest and most commonly encountered of all beetle families. Numerous s ...
s –
frass Frass refers loosely to the more or less solid excreta of insects, and to certain other related matter. Definition and etymology ''Frass'' is an informal term and accordingly it is variously used and variously defined. It is derived from the ...
*
Earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s,
lugworm ''Arenicola'', also known as sandworms, is a genus of capitellid annelid worms comprising the lugworms and black lugs. ''A.cristata'' is the dominant warm-water lugworm on the shores of North America and Humboldt Bay, California. ''A. caroledna' ...
s etc. – worm castings (feces extruded at ground surface) *Feces when used as
fertilizer A fertilizer or fertiliser is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from liming materials or other non-nutrient soil amendments. Man ...
(usually mixed with animal bedding and urine) –
manure Manure is organic matter that is used as organic fertilizer in agriculture. Most manure consists of animal feces; other sources include compost and green manure. Manures contribute to the fertility of soil by adding organic matter and nut ...
*
Horses The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 milli ...
– horse manure, roadapple (before motor vehicles became common, horse droppings were a big part of the rubbish communities needed to clean off roads)


Society and culture


Feelings of disgust

In all human cultures, feces elicit varying degrees of
disgust Disgust (, from Latin , ) is an emotional response of rejection or revulsion to something potentially contagious or something considered offensive, distasteful or unpleasant. In ''The Expression of the Emotions in Man and Animals'', Charles D ...
in adults. Children under two years typically have no disgust response to it, suggesting it is culturally derived. Disgust toward feces appears to be strongest in cultures where flush toilets make olfactory contact with human feces minimal. Disgust is experienced primarily in relation to the sense of taste (either perceived or imagined) and, secondarily to anything that causes a similar feeling by sense of smell, touch, or vision.


Social media

There is a Pile of Poo emoji represented in
Unicode Unicode or ''The Unicode Standard'' or TUS is a character encoding standard maintained by the Unicode Consortium designed to support the use of text in all of the world's writing systems that can be digitized. Version 16.0 defines 154,998 Char ...
as , called ' or ''unchi-kun'' in Japan.


Jokes

Poop is the center of
toilet humor Toilet humour or potty humour is a type of off-colour humour dealing with: defecation (including diarrhea and constipation), in which case it is called scatological humour (compare scatology); urination; flatulence, in which case it is called fla ...
, and is commonly an interest of young children and teenagers.


See also

*
Coprophagia Coprophagia ( ) or coprophagy ( ) is the consumption of feces. The word is derived from the Ancient Greek "feces" and "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of o ...
*
Coprophilia Coprophilia (from Greek κόπρος, ''kópros'' 'excrement' and φιλία, ''philía'' 'liking, fondness'), also called scatophilia or scat (Greek: σκατά, ''skatá'' 'feces'), is the paraphilia involving sexual arousal and pleasure from ...
*
Coprophilous fungi A coprophilous fungus (''dung-loving'' fungus) is a type of saprobic fungus that grows on animal dung. The hardy spores of coprophilous species are unwittingly consumed by herbivores from vegetation, and are excreted along with the plant matter. ...
*
Cow dung Cow dung, also known as cow pats, cow poop or cow manure, is the waste product (faeces) of bovine animal species. These species include domestic cattle ("cows"), bison ("buffalo"), yak, and water buffalo. Cow dung is the undigested residue of p ...
*
Excretory system The excretory system is a passive biological system that removes excess, unnecessary materials from the body fluids of an organism, so as to help maintain internal chemical homeostasis and prevent damage to the body. The dual function of excret ...
* Fecal microbiota * Fecal plug *
Jenkem Jenkem is a purported inhalant and hallucinogen created from fermented human waste. In the mid-1990s, it was reported to be a popular street drug among Zambian youth, created by placing feces and urine in a bottle or a bucket, sealing it with a ...
*
Manure Manure is organic matter that is used as organic fertilizer in agriculture. Most manure consists of animal feces; other sources include compost and green manure. Manures contribute to the fertility of soil by adding organic matter and nut ...
*
Metabolic waste Metabolic wastes or excrements are substances left over from metabolic processes (such as cellular respiration) which cannot be used by the organism (they are surplus or toxic), and must therefore be excreted. This includes nitrogen compounds ...
*
Night soil Night soil is a historical euphemism for Human waste, human excreta collected from cesspit, cesspools, privies, pail closets, pit latrines, privy middens, septic tanks, etc. This material was removed from the immediate area, usually at night, by ...
*
Sanitation Sanitation refers to public health conditions related to clean drinking water and treatment and disposal of human excreta and sewage. Preventing human contact with feces is part of sanitation, as is hand washing with soap. Sanitation systems ...
*
Scatology In medicine and biology, scatology or coprology is the study of faeces. Scatological studies allow one to determine a wide range of biological information about a creature, including its diet (and thus where it has been), health and diseases ...
* Whale feces * Zooplankton fecal pellets


References


External links


Article on Feces
– MedFriendly {{Authority control Animal physiology