In
biology
Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditar ...
, an organ is a collection of
tissues joined in a structural unit to serve a common function. In the
hierarchy of life, an organ lies between
tissue and an
organ system. Tissues are formed from same type
cells to act together in a function. Tissues of different types combine to form an organ which has a specific function. The
intestinal wall for example is formed by
epithelial tissue and
smooth muscle tissue.
Two or more organs working together in the execution of a specific body function form an organ system, also called a
biological system or body system.
An organ's tissues can be broadly categorized as
parenchyma, the functional tissue, and
stroma, the structural tissue with supportive, connective, or ancillary functions. For example, the
gland
In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream ( endocrine gland) or into cavities inside the body or its outer surface ( exocrine gland).
Structure
...
's tissue that makes the
hormones is the parenchyma, whereas the stroma includes the
nerves that innervate the parenchyma, the
blood vessel
Blood vessels are the structures of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away from ...
s that oxygenate and nourish it and carry away its metabolic wastes, and the
connective tissue
Connective tissue is one of the four primary types of animal tissue, along with epithelial tissue, muscle tissue, and nervous tissue. It develops from the mesenchyme derived from the mesoderm the middle embryonic germ layer. Connective tissue ...
s that provide a suitable place for it to be situated and anchored. The main tissues that make up an organ tend to have common
embryologic origins, such as arising from the same
germ layer. Organs exist in most multicellular
organism
In biology, an organism () is any life, living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy (biology), taxonomy into groups such as Multicellular o ...
s. In
single-celled organisms such as
bacteria
Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were am ...
, the
functional analogue of an organ is known as an
organelle
In cell biology, an organelle is a specialized subunit, usually within a cell, that has a specific function. The name ''organelle'' comes from the idea that these structures are parts of cells, as organs are to the body, hence ''organelle,'' t ...
. In plants, there are three main organs.
In the study of
anatomy
Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
, viscera (singular viscus) refers to the internal organs of the
abdominal,
thoracic, and
pelvic cavities.
The abdominal organs may be classified as solid organs, or hollow organs. The solid organs are the
liver
The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it ...
,
pancreas
The pancreas is an organ of the digestive system and endocrine system of vertebrates. In humans, it is located in the abdomen behind the stomach and functions as a gland. The pancreas is a mixed or heterocrine gland, i.e. it has both an en ...
,
spleen,
kidney
The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; bloo ...
s, and
adrenal glands. The hollow organs of the abdomen are the
stomach,
intestines,
gallbladder,
bladder, and
rectum.
In the
thoracic cavity the
heart
The heart is a muscular organ found in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon diox ...
is a hollow, muscular organ.
The number of organs in any organism depends on the definition used. By one widely adopted definition, 79 organs have been identified in the human body.
Animals

Except for
placozoans,
multicellular animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motilit ...
s including humans have a variety of
organ systems. These specific systems are widely studied in
human anatomy. The functions of these organ systems often share significant overlap. For instance, the
nervous and
endocrine system both operate via a shared organ, the
hypothalamus. For this reason, the two systems are combined and studied as the
neuroendocrine system. The same is true for the
musculoskeletal system because of the relationship between the
muscular and
skeletal systems.
*
Cardiovascular system: pumping and channeling
blood
Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood in th ...
to and from the
body and
lungs with
heart
The heart is a muscular organ found in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon diox ...
,
blood
Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood in th ...
and
blood vessel
Blood vessels are the structures of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away from ...
s.
*
Digestive system:
digestion and processing food with
salivary glands,
esophagus,
stomach,
liver
The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it ...
,
gallbladder,
pancreas
The pancreas is an organ of the digestive system and endocrine system of vertebrates. In humans, it is located in the abdomen behind the stomach and functions as a gland. The pancreas is a mixed or heterocrine gland, i.e. it has both an en ...
,
intestines,
colon,
rectum and
anus.
*
Endocrine system: communication within the body using
hormones made by
endocrine glands such as the
hypothalamus,
pituitary gland,
pineal body or pineal gland,
thyroid,
parathyroids and
adrenals, i.e., adrenal glands.
*
Excretory system:
kidney
The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; bloo ...
s,
ureter
The ureters are tubes made of smooth muscle that propel urine from the kidneys to the urinary bladder. In a human adult, the ureters are usually long and around in diameter. The ureter is lined by urothelial cells, a type of transitional e ...
s,
bladder and
urethra involved in fluid balance,
electrolyte balance and excretion of
urine.
*
Lymphatic system: structures involved in the transfer of
lymph between
tissues and the
blood stream, the lymph and the
nodes and
vessels that transport it including the
immune system
The immune system is a network of biological processes that protects an organism from diseases. It detects and responds to a wide variety of pathogens, from viruses to parasitic worms, as well as Tumor immunology, cancer cells and objects such ...
: defending against
disease
A disease is a particular abnormal condition that negatively affects the structure or function of all or part of an organism, and that is not immediately due to any external injury. Diseases are often known to be medical conditions that a ...
-causing agents with
leukocytes,
tonsils,
adenoids,
thymus and
spleen.
*
Integumentary system
The integumentary system is the set of organs forming the outermost layer of an animal's body. It comprises the skin and its appendages, which act as a physical barrier between the external environment and the internal environment that it serves ...
:
skin
Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation.
Other cuticle, animal coverings, such as the arthropod exoskeleton, have diffe ...
,
hair and
nails of mammals. Also
scales of
fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...
,
reptiles, and
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, and
feathers of birds.
*
Muscular system: movement with
muscle
Skeletal muscles (commonly referred to as muscles) are Organ (biology), organs of the vertebrate muscular system and typically are attached by tendons to bones of a skeleton. The muscle cells of skeletal muscles are much longer than in the other ...
s.
*
Nervous system
In Biology, biology, the nervous system is the Complex system, highly complex part of an animal that coordinates its Behavior, actions and Sense, sensory information by transmitting action potential, signals to and from different parts of its ...
: collecting, transferring and processing information with
brain
The brain is an organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It consists of nervous tissue and is typically located in the head ( cephalization), usually near organs for special ...
,
spinal cord
The spinal cord is a long, thin, tubular structure made up of nervous tissue, which extends from the medulla oblongata in the brainstem to the lumbar region of the vertebral column (backbone). The backbone encloses the central canal of the spin ...
and
nerves.
*
Reproductive system: the
sex organs, such as
ovaries,
fallopian tubes,
uterus
The uterus (from Latin ''uterus'', plural ''uteri'') or womb () is the organ in the reproductive system of most female mammals, including humans that accommodates the embryonic and fetal development of one or more embryos until birth. The ...
,
vulva,
vagina,
testes,
vas deferens,
seminal vesicles,
prostate and
penis
A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males d ...
.
*
Respiratory system
The respiratory system (also respiratory apparatus, ventilatory system) is a biological system consisting of specific organs and structures used for gas exchange in animals and plants. The anatomy and physiology that make this happen varies grea ...
: the organs used for
breathing
Breathing (or ventilation) is the process of moving air into and from the lungs to facilitate gas exchange with the internal environment, mostly to flush out carbon dioxide and bring in oxygen.
All aerobic creatures need oxygen for cell ...
, the
pharynx
The pharynx (plural: pharynges) is the part of the throat behind the mouth and nasal cavity, and above the oesophagus and trachea (the tubes going down to the stomach and the lungs). It is found in vertebrates and invertebrates, though its ...
,
larynx,
trachea,
bronchi,
lungs and
diaphragm
Diaphragm may refer to:
Anatomy
* Thoracic diaphragm, a thin sheet of muscle between the thorax and the abdomen
* Pelvic diaphragm or pelvic floor, a pelvic structure
* Urogenital diaphragm or triangular ligament, a pelvic structure
Other
* Diap ...
.
*
Skeletal system: structural support and protection with
bone
A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, an ...
s,
cartilage,
ligaments and
tendons.
Viscera
In the study of
anatomy
Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
, viscera (singular viscus) refers to the internal organs of the
abdominal,
thoracic, and
pelvic cavities.
The abdominal organs may be classed as solid organs, or hollow organs. The solid organs are the
liver
The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it ...
,
pancreas
The pancreas is an organ of the digestive system and endocrine system of vertebrates. In humans, it is located in the abdomen behind the stomach and functions as a gland. The pancreas is a mixed or heterocrine gland, i.e. it has both an en ...
,
spleen,
kidney
The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; bloo ...
s, and
adrenal glands. The hollow organs are the
stomach,
intestines,
gallbladder,
bladder, and
rectum.
In the
thoracic cavity the
heart
The heart is a muscular organ found in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon diox ...
is a hollow, muscular organ.
Splanchnology is the study of the viscera.
The term "visceral" is contrasted with the term "", meaning "of or relating to the wall of a body part,
organ or cavity"
The two terms are often used in describing a membrane or piece of connective tissue, referring to the opposing sides.
Origin and evolution

The organ level of organisation in
animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motilit ...
s can be first detected in
flatworm
The flatworms, flat worms, Platyhelminthes, or platyhelminths (from the Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") are a phylum of relatively simple bilaterian, unsegme ...
s and the more derived
phyla Phyla, the plural of ''phylum'', may refer to:
* Phylum, a biological taxon between Kingdom and Class
* by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another
Phyl ...
, i.e. the
bilaterians. The less-advanced
taxa (i.e. ''
Placozoa'', ''
Porifera'', ''
Ctenophora'' and ''
Cnidaria
Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in Fresh water, freshwater and Marine habitats, marine environments, predominantly the latter.
Their distinguishing feature is cnidocyt ...
'') do not show consolidation of their tissues into organs.
More complex animals are composed of different organs, which have evolved over time. For example, the liver and heart evolved in the
chordates about 550-500 million years ago, while the gut and brain are even more ancient, arising in the ancestor of vertebrates, insects, molluscs, and worms about 700-650 million years ago.
Given the ancient origin of most vertebrate organs, researchers have looked for model systems, where organs have evolved more recently, and ideally have evolved multiple times independently. An outstanding model for this kind of research is the
placenta
The placenta is a temporary embryonic and later fetal organ (anatomy), organ that begins embryonic development, developing from the blastocyst shortly after implantation (embryology), implantation. It plays critical roles in facilitating nutrien ...
, which has evolved more than 100 times independently in vertebrates, has evolved relatively recently in some lineages, and exists in intermediate forms in extant taxa.
Studies on the evolution of the placenta have identified a variety of genetic and physiological processes that contribute to the origin and evolution of organs, these include the re-purposing of existing animal tissues, the acquisition of new functional properties by these tissues, and novel interactions of distinct tissue types.
Plants
The study of plant organs is covered in
plant morphology
Phytomorphology is the study of the physical form and external structure of plants.Raven, P. H., R. F. Evert, & S. E. Eichhorn. ''Biology of Plants'', 7th ed., page 9. (New York: W. H. Freeman, 2005). . This is usually considered distinct from ...
. Organs of
plant
Plants are predominantly Photosynthesis, photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all curr ...
s can be divided into vegetative and reproductive. Vegetative plant organs include
roots,
stems, and
leaves. The reproductive organs are variable. In
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
s, they are represented by the
flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
,
seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
and
fruit
In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering.
Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
. In
conifers, the organ that bears the reproductive structures is called a
cone. In other divisions (
phyla Phyla, the plural of ''phylum'', may refer to:
* Phylum, a biological taxon between Kingdom and Class
* by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another
Phyl ...
) of plants, the reproductive organs are called
strobili, in ''
Lycopodiophyta'', or simply gametophores in
mosses. Common organ system designations in plants include the differentiation of shoot and root. All parts of the plant above ground (in non-
epiphytes), including the functionally distinct leaf and flower organs, may be classified together as the shoot organ system.
The vegetative organs are essential for maintaining the life of a plant. While there can be 11 organ systems in animals, there are far fewer in plants, where some perform the vital functions, such as
photosynthesis, while the reproductive organs are essential in
reproduction. However, if there is
asexual vegetative reproduction, the vegetative organs are those that create the new generation of plants (see
clonal colony).
Society and culture
Many societies have a system for
organ donation, in which a living or deceased donor's organ are
transplanted into a person with a failing organ. The transplantation of larger solid organs often requires
immunosuppression to prevent
organ rejection or
graft-versus-host disease.
There is considerable interest throughout the world in creating
laboratory-grown or
artificial organs.
Organ transplants
Beginning in the 20th century
organ transplants began to take place as scientists knew more about the anatomy of organs. These came later in time as procedures were often dangerous and difficult. Both the source and method of obtaining the organ to transplant are major ethical issues to consider, and because organs as resources for transplant are always more limited than demand for them, various notions of justice, including
distributive justice
Distributive justice concerns the socially just allocation of resources. Often contrasted with just process, which is concerned with the administration of law, distributive justice concentrates on outcomes. This subject has been given considerab ...
, are developed in the ethical analysis. This situation continues as long as transplantation relies upon organ donors rather than technological innovation, testing, and industrial manufacturing.
History

The English word "organ" dates back to the twelfth century and refers to any musical instrument. By the late 14th century, the musical term's meaning had narrowed to refer specifically to the
keyboard-based instrument. At the same time, a second meaning arose, in reference to a "body part adapted to a certain function".
Plant organs are made from tissue composed of different types of tissue. The three tissue types are ground, vascular, and dermal. When three or more organs are present, it is called an organ system.
The adjective ''
visceral'', also ''
splanchnic'', is used for anything pertaining to the internal organs. Historically, viscera of animals were examined by
Roman pagan
priest
A priest is a religious leader authorized to perform the sacred rituals of a religion, especially as a mediatory agent between humans and one or more deities. They also have the authority or power to administer religious rites; in particu ...
s like the
haruspices or the
augurs in order to divine the future by their shape, dimensions or other factors. This practice remains an important ritual in some remote, tribal societies.
The term "visceral" is contrasted with the term "", meaning "of or relating to the wall of a body part,
organ or cavity"
The two terms are often used in describing a membrane or piece of connective tissue, referring to the opposing sides.
Antiquity
Aristotle
Aristotle (; grc-gre, Ἀριστοτέλης ''Aristotélēs'', ; 384–322 BC) was a Greek philosopher and polymath during the Classical Greece, Classical period in Ancient Greece. Taught by Plato, he was the founder of the Peripatet ...
used the word frequently in his philosophy, both to describe the organs of plants or animals (e.g. the roots of a tree, the heart or liver of an animal), and to describe more abstract "parts" of an interconnected whole (e.g. his logical works, taken as a whole, are referred to as the ''
Organon
The ''Organon'' ( grc, Ὄργανον, meaning "instrument, tool, organ") is the standard collection of Aristotle's six works on logical analysis and dialectic. The name ''Organon'' was given by Aristotle's followers, the Peripatetics.
The si ...
'').
Some alchemists (e.g.
Paracelsus) adopted the
Hermetic Qabalah assignment between the seven vital organs and the seven
classical planets as follows:
See also
*
Organoid
*
Organ-on-a-chip
References
External links
*
{{Authority control
Levels of organization (Biology)