Madrean Sky Islands Mountain Ranges
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The Madrean Region (named after the
Sierra Madre Occidental The Sierra Madre Occidental is a major mountain range system of the North American Cordillera, that runs northwest–southeast through northwestern and western Mexico, and along the Gulf of California. The Sierra Madre is part of the American ...
) is a
floristic region A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both r ...
within the
Holarctic Kingdom The asterisk ( ), from Late Latin , from Ancient Greek , ''asteriskos'', "little star", is a typographical symbol. It is so called because it resembles a conventional image of a heraldic star. Computer scientists and mathematicians often voc ...
in North America, as delineated by Armen Takhtajan and Robert F. Thorne. It occupies arid or semiarid areas in the southwestern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
and northwestern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
and is bordered by the Rocky Mountain Region and
North American Atlantic Region North American Atlantic Region is a floristic region within the Holarctic Kingdom identified by Armen Takhtajan and Robert F. Thorne, spanning from the Atlantic and Gulf coasts to the Great Plains and comprising a major part of the United States ...
of the
Holarctic Kingdom The asterisk ( ), from Late Latin , from Ancient Greek , ''asteriskos'', "little star", is a typographical symbol. It is so called because it resembles a conventional image of a heraldic star. Computer scientists and mathematicians often voc ...
in the north and in the east,
Caribbean Region The Caribbean region of Colombia or Caribbean coast region is in the north of Colombia and is mainly composed of 8 departments located contiguous to the Caribbean.Neotropical Kingdom in the south. The Madrean Region is characterized by a very distinct
flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' ...
with at least three
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideal ...
( Fouquieriaceae,
Simmondsiaceae Simmondsiaceae or the jojoba family is a family of flowering plants. The family is not recognized by all taxonomic systems, the single species, ''Simmondsia chinensis'', often being treated as belonging to family Buxaceae. The APG II system, of ...
, and
Setchellanthaceae ''Setchellanthus caeruleus'' is a species of pungent shrub with large blue flowers. It is placed alone in the genus ''Setchellanthus'', which is in turn, is placed alone in the family Setchellanthaceae. It is endemic to Mexico. The genus and the ...
).
Crossosomataceae Crossosomataceae is a small plant family, consisting of four genera of shrubs found only in the dry parts of the American southwest and Mexico. This family has included up to ten species in the past, although as of 2021 six species are still ...
,
Garryaceae Garryaceae is a small family of plants known commonly as the silktassels.Garryaceae.
Flora o ...
,
Lennoaceae Lennooideae is a subfamily of parasitic flowering plants of southwestern North America and northwestern South America. The relationships of this subfamily to other plants remain uncertain. It was traditionally treated at family rank as Lennoacea ...
,
Limnanthaceae The Limnanthaceae are a small family of annual herbs occurring throughout temperate North America. There are eight species and nineteen taxa currently recognized. Members of this family are prominent in vernal pool communities of California. Some ...
and Stegnospermataceae have their principal development here; for
Onagraceae The Onagraceae are a family of flowering plants known as the willowherb family or evening primrose family. They include about 650 species of herbs, shrubs, and treesPolemoniaceae The Polemoniaceae (Jacob's-ladder or phlox family) are a family of flowering plants consisting of about 25 genera with 270–400 species of annuals and perennials native to the Northern Hemisphere and South America, with the center of diversit ...
and Hydrophyllaceae it is the major center of diversity. More than 250 genera and probably more than half of the
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of the region are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to it according to Takhtajan.


Floristic provinces

The region is subdivided into four
floristic provinces A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both r ...
: * Great Basin Province * Californian Province *Sonoran Province * Mexican Highlands Province


Great Basin Province

The
Great Basin Floristic Province The Great Basin Floristic Province is a floristic province of the Madrean Subkingdom (floristic region), in the Boreal Kingdom (floristic kingdom). It is located in the Western United States. A floristic province (otherwise known as a phytochor ...
includes most of the Great Basin, as well as the Colorado Plateau, the Snake River Plain, and Arizona north of the
Mogollon Rim The Mogollon Rim ( or or ) is a topographical and geological feature cutting across the northern half of the U.S. state of Arizona. It extends approximately , starting in northern Yavapai County and running eastward, ending near the border ...
. It shares much of its flora with the neighboring provinces and has only a few endemic genera.
Species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
endemism is also moderate (about 25%), but is much more considerable in such genera as ''
Astragalus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to tempe ...
'', ''
Eriogonum ''Eriogonum'' is a genus of flowering plants in the family Polygonaceae. The genus is found in North America and is known as wild buckwheat. This is a highly species-rich genus, and indications are that active speciation is continuing. It incl ...
'', ''
Penstemon ''Penstemon'' , the beardtongues, is a large genus of roughly 250 species of flowering plants native mostly to the Nearctic, but with a few species also found in the North American portion of the Neotropics. It is the largest genus of flowering ...
'', ''
Cymopterus ''Cymopterus'' is a genus of perennial plants in the family Apiaceae native to western North America. They are commonly known as the springparsleys.
'', '' Lomatium'', ''
Cryptantha ''Cryptantha'' is a genus of flowering plants in the borage family, Boraginaceae. They are known commonly as cat's eyes and popcorn flowers (the latter name is also used to refer to the closely related genus ''Plagiobothrys'',Hasenstab-Lehman, ...
'', '' Chrysothamnus'', ''
Erigeron ''Erigeron'' () is a large genus of plants in the family Asteraceae. It is closely related to the genus '' Aster'' and the true daisies in the genus '' Bellis''. The genus has a cosmopolitan distribution in dry, mountainous areas and grassland, ...
'', ''
Phacelia ''Phacelia'' (phacelia, scorpionweed, heliotrope) is a genus of about 200 species of annual or perennial herbaceous plants in the borage family, native to North and South America. California is particularly rich in species with over 90 recorded ...
'', ''
Castilleja ''Castilleja'', commonly known as paintbrush, Indian paintbrush, or prairie-fire, is a genus of about 200 species of annual and perennial herbaceous plants native to the west of the Americas from Alaska south to the Andes, northern Asia, and on ...
'', and '' Gilia''. The vegetation in the central
Great Basin shrub steppe The Great Basin Desert is part of the Great Basin between the Sierra Nevada and the Wasatch Range. The desert is a geographical region that largely overlaps the Great Basin shrub steppe defined by the World Wildlife Fund, and the Central Basin ...
part of the province is dominated by '' Artemisia'' species and
Chenopodiaceae Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus '' Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making i ...
genera. The Great Basin montane forests include ancient
Great Basin bristlecone pine ''Pinus longaeva'' (commonly referred to as the Great Basin bristlecone pine, intermountain bristlecone pine, or western bristlecone pine) is a long-living species of bristlecone pine tree found in the higher mountains of California, Nevada, and ...
(''Pinus longaeva'') trees.


Californian Province

The Californian Province occupies Central and Southern
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
from the seashore to the foothills of the Cascade Range, the Sierra Nevada range, and northern parts of
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
in
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. It possesses the most diverse flora within the floristic region. About half of the species are endemic. More than 50 genera (e.g. ''
Adenostoma ''Adenostoma'' is a genus of shrubs in the Rose family ( Rosaceae) containing only two species, chamise (''Adenostoma fasciculatum'') and redshanks (''Adenostoma sparsifolium''). Both are native to the Californias. Description Characteristi ...
'', '' Bergerocactus'', '' Carpenteria'', '' Cneoridium'', '' Dendromecon'', '' Fremontodendron'', '' Jepsonia'', ''
Lyonothamnus ''Lyonothamnus'' is a monotypic genus of trees in the rose family containing the single living species ''Lyonothamnus floribundus'', which is known by the common name Catalina ironwood, and the subspecies ''L. f.'' ssp. ''aspleniifolius'' and '' ...
'', '' Neostapfia'', '' Odontostomum'', ''
Ornithostaphylos ''Ornithostaphylos'' is a monotypic plant genus which contains the single species ''Ornithostaphylos oppositifolia'', commonly known as the Baja California birdbush or Baja California manzanita. A large, evergreen shrub in the heather family, th ...
'', '' Pickeringia'', '' Romneya'') are endemic or near-endemic as well. ''
Arctostaphylos ''Arctostaphylos'' (; from "bear" and "bunch of grapes") is a genus of plants comprising the manzanitas () and bearberries. They are shrubs or small trees. There are about 60 species, of ''Arctostaphylos'', ranging from ground-hugging arc ...
'', Brodiaeinae, ''
Calochortus ''Calochortus'' is a genus of flowering plants in the lily family. The group includes herbaceous, perennial and bulbous species, all native to North America (primarily the Western United States). The genus ''Calochortus'' includes mariposa ...
'', ''
Caulanthus ''Caulanthus'' is a genus of plants in the family Brassicaceae. Plants of this genus may be known as jewelflowers. They are also often referred to as wild cabbage, although this common name usually refers to wild variants of ''Brassica oleraceae' ...
'', '' Streptanthus'', '' Ceanothus'', ''
Cryptantha ''Cryptantha'' is a genus of flowering plants in the borage family, Boraginaceae. They are known commonly as cat's eyes and popcorn flowers (the latter name is also used to refer to the closely related genus ''Plagiobothrys'',Hasenstab-Lehman, ...
'', '' Downingia'', ''
Dudleya ''Dudleya'', commonly known as liveforevers (Spanish: ''siemprevivas'') is a genus of succulent plants in the stonecrop family, Crassulaceae, consisting of about 68 taxa in southwestern North America and Guadalupe Island. The species come in mu ...
'', Eritrichieae, Eriogonoideae, Gileae, Hydrophyllaceae,
Limnanthaceae The Limnanthaceae are a small family of annual herbs occurring throughout temperate North America. There are eight species and nineteen taxa currently recognized. Members of this family are prominent in vernal pool communities of California. Some ...
, '' Lotus'', Madiinae, ''
Mimulus Mimulus is a plant genus in the family Phrymaceae, which was traditionally placed in family Scrophulariaceae. The genus now contains only seven species, two native to eastern North America and the other five native to Asia, Australia, Africa, ...
'', Onagreae, Epilobieae, Orcuttieae, Eschscholzioideae, Platystemonoideae, ''
Astragalus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to tempe ...
'' and '' Cupressus'' have a principal center of diversity within the province. The flora of the Californian Province is partially shared with the spatially distant Chile-Patagonian Region of the Antarctic Kingdom and to a lesser extent
Mediterranean Region In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and wa ...
of the
Holarctic Kingdom The asterisk ( ), from Late Latin , from Ancient Greek , ''asteriskos'', "little star", is a typographical symbol. It is so called because it resembles a conventional image of a heraldic star. Computer scientists and mathematicians often voc ...
. The vegetation of the Californian Province is varied. Wetter northern parts of the Californian Province ( Northern California and
Oregon Oregon () is a state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington, while the Snake River delineates much of its eastern boundary with Idaho. T ...
) as defined by Peter Raven and D.I. Axelrod, as well as by Conservation International, fall under the Rocky Mountain Region in Takhtajan and Thorne's system and don't make parts of the Madrean Region.


Sonoran Province

The Sonoran Province comprises arid areas in the southwestern U.S. and northwestern Mexico from
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
and
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
to
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
and
Tamaulipas Tamaulipas (), officially the Free and Sovereign State of Tamaulipas ( es, Estado Libre y Soberano de Tamaulipas), is a state in the northeast region of Mexico; one of the 31 states which, along with Mexico City, comprise the 32 Federal Entiti ...
, including the Mojave (characterized by ''
Yucca brevifolia ''Yucca brevifolia'' is a plant species belonging to the genus ''Yucca''. It is tree-like in habit, which is reflected in its common names: Joshua tree, yucca palm, tree yucca, and palm tree yucca. This monocotyledonous tree is native to the ar ...
'', Joshua Tree), Sonora and Chihuahua Deserts. The vegetation is dominated by ''
Yucca ''Yucca'' is a genus of perennial plant, perennial shrubs and trees in the family (biology), family Asparagaceae, subfamily Agavoideae. Its 40–50 species are notable for their Rosette (botany), rosettes of evergreen, tough, sword-shaped Leaf, ...
'',
Cactaceae A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Gree ...
species ('' Opuntia'' spp. and other), as well as by ''
Larrea tridentata ''Larrea tridentata'', called creosote bush and greasewood as a plant, chaparral as a medicinal herb, and ''gobernadora'' (Spanish for "governess") in Mexico, due to its ability to secure more water by inhibiting the growth of nearby plants. In S ...
'' (Creosote bush).


Mexican Highlands Province

The Mexican Highlands Province comprises areas in the
Mexican Plateau The Central Mexican Plateau, also known as the Mexican Altiplano ( es, Altiplanicie Mexicana), is a large arid-to-semiarid plateau that occupies much of northern and central Mexico. Averaging above sea level, it extends from the United States b ...
region, including
Madrean pine-oak woodlands The Madrean Region (named after the Sierra Madre Occidental) is a floristic region within the Holarctic Kingdom in North America, as delineated by Armen Takhtajan and Robert F. Thorne. It occupies arid or semiarid areas in the southwestern United ...
.


See also

*
Madrean Sky Islands The Madrean Sky Islands are enclaves of Madrean pine–oak woodlands, found at higher elevations in a complex of small mountain ranges in southern and southeastern Arizona, southwestern New Mexico, and northwestern Mexico. The sky islands are s ...


Bibliography

*Thorne, Robert F
Phytogeography of North America North of Mexico
'' Flora of North America'', Vol. 1, Ch. 6. {{Flora of the United States by political division Floristic regions * Natural history of Mexico Plant communities of the Western United States Plant communities of California Mexican Plateau * Natural history of the Colorado Desert Natural history of the Mojave Desert Holarctic