''Linheraptor'' is a
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
dromaeosaurid
Dromaeosauridae () is a family (biology), family of feathered coelurosaurian Theropoda, theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous period (geology), Period. The name Drom ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
which lived in what is now
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
in the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
. It was named by
Xu Xing and colleagues in 2010, and contains the species ''Linheraptor exquisitus''.
This bird-like dinosaur was less than 2 m (6.5 ft) long and was found in
Inner Mongolia
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an Autonomous regions of China, autonomous region of China. Its border includes two-thirds of the length of China's China–Mongolia border, border with the country of Mongolia. ...
. It is known from a single, nearly complete skeleton.
Discovery
Researchers announced the discovery of the genus after a nearly complete fossilised skeleton was found in 2008 by
Jonah N. Choiniere and Michael Pittman
in
Inner Mongolia
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an Autonomous regions of China, autonomous region of China. Its border includes two-thirds of the length of China's China–Mongolia border, border with the country of Mongolia. ...
; a more detailed publication is forthcoming.
The specimen was recovered from rocks at
Bayan Mandahu that belong to the
Wulansuhai Formation. The latter includes lithologies that are very similar to the Mongolian
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
-aged rocks of the
Djadokhta Formation
The Djadochta formation (sometimes transcribed and also known as Djadokhta, Djadokata, or Dzhadokhtskaya) is a highly fossiliferous geological formation in Central Asia, Gobi Desert, dating from the Late Cretaceous period, about 75 million ...
which have yielded the closely related dromaeosaurids ''
Tsaagan'' and ''Velociraptor''.
The
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''Linheraptor'', articulated and uncompressed, is one of the few nearly complete skeletons of dromaeosaurid dinosaurs worldwide.
The name of the genus refers to the district of
Linhe, Inner Mongolia, China where the specimen was discovered, while the
specific name, ''exquisitus'', refers to the well-preserved nature of the holotype (
IVPP V 16923).
Description

''Linheraptor'' was a bird-like
theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur. It was a dromaeosaurid which measured approximately in length,
and weighed up to approximately .
At that size, ''Linheraptor'' would have been a fast and agile predator, perhaps preying on small
ceratopsia
Ceratopsia or Ceratopia ( or ; Ancient Greek, Greek: "horned faces") is a group of herbivore, herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period (geology), Period, although ance ...
ns.
Like all dromaeosaurids, it had an elongated skull, a curved neck, an enlarged toe claw on each foot, and a long tail; ''Linheraptor'' was
bipedal
Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' ...
and
carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
. The large toe claws may have been used for capturing prey.
Taxonomy

Among its sister taxa, ''Linheraptor'' is believed to be most closely related to ''
Tsaagan mangas''. ''Linheraptor'' and ''Tsaagan'' were intermediate between basal and derived dromaeosaurids. The two share several skull details, among which a large maxillary fenestra — an opening in the
maxilla
In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
, an upper jaw bone — and lack various features of more
derived dromaeosaurids such as ''Velociraptor''.
Senter (2011) and Turner, Makovicky and Norell (2012) argue that ''Linheraptor exquisitus'' is a junior synonym of ''Tsaagan mangas'',
but Xu, Pittman ''et al''. (2015) reject this synonymy by responding to the counterarguments proposed using new and existing details of ''Linheraptors anatomy.
A monographic description of ''Linheraptor'' is currently in preparation.
See also
*
Timeline of dromaeosaurid research
References
External links
Fossil of ''Linheraptor''
{{Taxonbar, from=Q132521
Eudromaeosauria
Dinosaur genera
Campanian dinosaurs
Fossil taxa described in 2010
Taxa named by Mark Norell
Taxa named by Xu Xing
Taxa named by James M. Clark
Dinosaurs of China