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Extinction is the termination of an
organism An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have be ...
by the
death Death is the end of life; the irreversible cessation of all biological functions that sustain a living organism. Death eventually and inevitably occurs in all organisms. The remains of a former organism normally begin to decompose sh ...
of its last member. A
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
may become functionally extinct before the death of its last member if it loses the capacity to reproduce and recover. As a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
) after a period of apparent absence. Over five billion species are estimated to have died out. It is estimated that there are currently around 8.7 million species of
eukaryote The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms ...
s globally, possibly many times more if
microorganism A microorganism, or microbe, is an organism of microscopic scale, microscopic size, which may exist in its unicellular organism, single-celled form or as a Colony (biology)#Microbial colonies, colony of cells. The possible existence of unseen ...
s are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, and
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus.'' They lived from the late Miocene epoch (from around 6.2 million years ago) into the Holocene until about 4,000 years ago, with mammoth species at various times inhabi ...
s. Through
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
, species arise through the process of speciation. Species become extinct when they are no longer able to survive in changing conditions or against superior
competition Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indi ...
. The relationship between animals and their ecological niches has been firmly established. A typical species becomes extinct within 10 million years of its first appearance, although some species, called
living fossil A living fossil is a Deprecation, deprecated term for an extant taxon that phenotypically resembles related species known only from the fossil record. To be considered a living fossil, the fossil species must be old relative to the time of or ...
s, survive with little to no morphological change for hundreds of millions of years. Mass extinctions are relatively rare events; however, isolated extinctions of species and clades are quite common, and are a natural part of the evolutionary process. Only recently have extinctions been recorded with scientists alarmed at the current high rate of extinctions. Most species that become extinct are never scientifically documented. Some scientists estimate that up to half of presently existing plant and animal species may become extinct by 2100. Wilson, E.O., ''The Future of Life'' (2002) (). See also: Leakey, Richard, ''The Sixth Extinction : Patterns of Life and the Future of Humankind'', A 2018 report indicated that the phylogenetic diversity of 300 mammalian species erased during the human era since the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
would require 5 to 7 million years to recover. According to the 2019 '' Global Assessment Report on Biodiversity and Ecosystem Services'' by IPBES, the biomass of wild mammals has fallen by 82%, natural ecosystems have lost about half their area and a million species are at risk of extinction—all largely as a result of human actions. Twenty-five percent of plant and animal species are threatened with extinction. In a subsequent report, IPBES listed unsustainable fishing, hunting and logging as being some of the primary drivers of the global extinction crisis. In June 2019, one million species of plants and animals were at risk of extinction. At least 571 plant species have been lost since 1750. The main cause of the extinctions is the destruction of natural habitats by human activities, such as cutting down forests and converting land into fields for farming. A dagger symbol (†) placed next to the name of a species or other taxon normally indicates its status as extinct.


Definition

A species is extinct when the last existing member dies. Extinction therefore becomes a certainty when there are no surviving individuals that can reproduce and create a new generation. A species may become functionally extinct when only a handful of individuals survive, which cannot reproduce due to poor health, age, sparse distribution over a large range, a lack of individuals of both sexes (in sexually reproducing species), or other reasons. Pinpointing the extinction (or pseudoextinction) of a species requires a clear definition of that species. If it is to be declared extinct, the species in question must be uniquely distinguishable from any ancestor or daughter species, and from any other closely related species. Extinction of a species (or replacement by a daughter species) plays a key role in the punctuated equilibrium hypothesis of
Stephen Jay Gould Stephen Jay Gould ( ; September 10, 1941 – May 20, 2002) was an American Paleontology, paleontologist, Evolutionary biology, evolutionary biologist, and History of science, historian of science. He was one of the most influential and widely re ...
and Niles Eldredge. In
ecology Ecology () is the natural science of the relationships among living organisms and their Natural environment, environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosystem, and biosphere lev ...
, ''extinction'' is sometimes used informally to refer to
local extinction Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with extinction, global extinctions. Local extinctions ...
, in which a species ceases to exist in the chosen area of study, despite still existing elsewhere. Local extinctions may be made good by the reintroduction of individuals of that species taken from other locations; wolf reintroduction is an example of this. Species that are not globally extinct are termed
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
. Those species that are extant, yet are threatened with extinction, are referred to as threatened or
endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
. Currently, an important aspect of extinction is human attempts to preserve critically endangered species. These are reflected by the creation of the conservation status "extinct in the wild" (EW). Species listed under this status by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
(IUCN) are not known to have any living specimens in the wild and are maintained only in zoos or other artificial environments. Some of these species are functionally extinct, as they are no longer part of their natural habitat and it is unlikely the species will ever be restored to the wild.Maas, Peter.
Extinct in the Wild" ''The Extinction Website''. URL accessed January 26 2007.
When possible, modern zoological institutions try to maintain a viable population for species preservation and possible future reintroduction to the wild, through use of carefully planned breeding programs. The extinction of one species' wild population can have knock-on effects, causing further extinctions. These are also called "chains of extinction". This is especially common with extinction of keystone species. A 2018 study indicated that the sixth mass extinction started in the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
could take up to 5 to 7 million years to restore mammal diversity to what it was before the human era.


Pseudoextinction

Extinction of a parent species where daughter species or subspecies are still extant is called pseudoextinction or phyletic extinction. Effectively, the old taxon vanishes, transformed ( anagenesis) into a successor, or split into more than one ( cladogenesis). Pseudoextinction is difficult to demonstrate unless one has a strong chain of evidence linking a living species to members of a pre-existing species. For example, it is sometimes claimed that the extinct '' Hyracotherium'', which was an early horse that shares a common ancestor with the modern
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
, is pseudoextinct, rather than extinct, because there are several extant species of '' Equus'', including zebra and
donkey The donkey or ass is a domesticated equine. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a separate species, ''Equus asinus''. It was domes ...
; however, as fossil species typically leave no genetic material behind, one cannot say whether ''Hyracotherium'' evolved into more modern horse species or merely evolved from a common ancestor with modern horses. Pseudoextinction is much easier to demonstrate for larger taxonomic groups.


Lazarus taxa

A Lazarus taxon or Lazarus species refers to instances where a species or taxon was thought to be extinct, but was later rediscovered. It can also refer to instances where large gaps in the fossil record of a taxon result in fossils reappearing much later, although the taxon may have ultimately become extinct at a later point. The coelacanth, a fish related to lungfish and
tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...
s, is an example of a Lazarus taxon that was known only from the fossil record and was considered to have been extinct since the end of the Cretaceous Period. In 1938, however, a living specimen was found off the Chalumna River (now Tyolomnqa) on the east coast of South Africa. '' Calliostoma bullatum'', a species of deepwater
sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...
originally described from fossils in 1844 proved to be a Lazarus species when extant individuals were described in 2019. Attenborough's long-beaked echidna (''Zaglossus attenboroughi'') is an example of a Lazarus species from
Papua New Guinea Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
that had last been sighted in 1962 and believed to be possibly extinct, until it was recorded again in November 2023. Some species thought to be extinct have had ongoing speculation that they may still exist, and in the event of rediscovery would be considered Lazarus species. Examples include the
thylacine The thylacine (; binomial name ''Thylacinus cynocephalus''), also commonly known as the Tasmanian tiger or Tasmanian wolf, was a carnivorous marsupial that was native to the Mainland Australia, Australian mainland and the islands of Tasmani ...
, or Tasmanian tiger (''Thylacinus cynocephalus''), the last known example of which died in Hobart Zoo in Tasmania in 1936; the Japanese wolf (''Canis lupus hodophilax''), last sighted over 100 years ago; the American ivory-billed woodpecker (''Campephilus principalis''), with the last universally accepted sighting in 1944; and the slender-billed curlew (''Numenius tenuirostris''), not seen since 2007.


Causes

As long as species have been evolving, species have been going extinct. It is estimated that over 99.9% of all species that ever lived are extinct. The average lifespan of a species is 1–10 million years, although this varies widely between taxa. A variety of causes can contribute directly or indirectly to the extinction of a species or group of species. "Just as each species is unique", write Beverly and Stephen C. Stearns, "so is each extinction ... the causes for each are varied—some subtle and complex, others obvious and simple". Most simply, any species that cannot survive and reproduce in its environment and cannot move to a new environment where it can do so, dies out and becomes extinct. Extinction of a species may come suddenly when an otherwise healthy species is wiped out completely, as when toxic
pollution Pollution is the introduction of contaminants into the natural environment that cause harm. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the component ...
renders its entire
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
unliveable; or may occur gradually over thousands or millions of years, such as when a species gradually loses out in competition for food to better adapted competitors. Extinction may occur a long time after the events that set it in motion, a phenomenon known as extinction debt. Assessing the relative importance of genetic factors compared to environmental ones as the causes of extinction has been compared to the debate on nature and nurture. The question of whether more extinctions in the
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
record have been caused by
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
or by competition or by predation or by disease or by catastrophe is a subject of discussion; Mark Newman, the author of ''Modeling Extinction'', argues for a mathematical model that falls in all positions. By contrast,
conservation biology Conservation biology is the study of the conservation of nature and of Earth's biodiversity with the aim of protecting species, their habitats, and ecosystems from excessive rates of extinction and the erosion of biotic interactions. It is an i ...
uses the extinction vortex model to classify extinctions by cause. When concerns about human extinction have been raised, for example in Sir Martin Rees' 2003 book '' Our Final Hour'', those concerns lie with the effects of
climate change Present-day climate change includes both global warming—the ongoing increase in Global surface temperature, global average temperature—and its wider effects on Earth's climate system. Climate variability and change, Climate change in ...
or
technological Technology is the application of conceptual knowledge to achieve practical goals, especially in a reproducible way. The word ''technology'' can also mean the products resulting from such efforts, including both tangible tools such as ute ...
disaster. Human-driven extinction started as humans migrated out of Africa more than 60,000 years ago. Currently, environmental groups and some governments are concerned with the extinction of species caused by humanity, and they try to prevent further extinctions through a variety of conservation programs. Humans can cause extinction of a species through
overharvesting Overexploitation, also called overharvesting or ecological overshoot, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to ...
,
pollution Pollution is the introduction of contaminants into the natural environment that cause harm. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the component ...
, habitat destruction, introduction of
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
(such as new
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s and food
competitors Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indi ...
), overhunting, and other influences. Explosive, unsustainable human
population growth Population growth is the increase in the number of people in a population or dispersed group. The World population, global population has grown from 1 billion in 1800 to 8.2 billion in 2025. Actual global human population growth amounts to aroun ...
and increasing per capita consumption are essential drivers of the extinction crisis. According to the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
(IUCN), 784 extinctions have been recorded since the year 1500, the arbitrary date selected to define "recent" extinctions, up to the year 2004; with many more likely to have gone unnoticed. Several species have also been listed as extinct since 2004.


Genetics and demographic phenomena

If
adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the p ...
increasing population fitness is slower than
environmental degradation Environment most often refers to: __NOTOC__ * Natural environment, referring respectively to all living and non-living things occurring naturally and the physical and biological factors along with their chemical interactions that affect an organism ...
plus the accumulation of slightly deleterious
mutation In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, ...
s, then a population will go extinct. Smaller populations have fewer beneficial mutations entering the population each generation, slowing adaptation. It is also easier for slightly deleterious mutations to fix in small populations; the resulting positive feedback loop between small population size and low fitness can cause mutational meltdown. Limited geographic range is the most important determinant of
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
extinction at background rates but becomes increasingly irrelevant as mass extinction arises. Limited geographic range is a cause both of small population size and of greater vulnerability to local environmental catastrophes. Extinction rates can be affected not just by population size, but by any factor that affects evolvability, including balancing selection, cryptic genetic variation, phenotypic plasticity, and robustness. A diverse or deep gene pool gives a population a higher chance in the short term of surviving an adverse change in conditions. Effects that cause or reward a loss in
genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is d ...
can increase the chances of extinction of a species. Population bottlenecks can dramatically reduce genetic diversity by severely limiting the number of reproducing individuals and make
inbreeding Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely genetic distance, related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genet ...
more frequent.


Genetic pollution

Extinction sometimes results for species evolved to specific ecologies that are subjected to genetic pollution—i.e., uncontrolled hybridization, introgression and genetic swamping that lead to homogenization or out-competition from the introduced ( or hybrid) species. Endemic populations can face such extinctions when new populations are imported or selectively bred by people, or when habitat modification brings previously isolated species into contact. Extinction is likeliest for rare species coming into contact with more abundant ones; interbreeding can swamp the rarer gene pool and create hybrids, depleting the purebred gene pool (for example, the endangered wild water buffalo is most threatened with extinction by genetic pollution from the abundant domestic water buffalo). Such extinctions are not always apparent from morphological (non-genetic) observations. Some degree of
gene flow In population genetics, gene flow (also known as migration and allele flow) is the transfer of genetic variation, genetic material from one population to another. If the rate of gene flow is high enough, then two populations will have equivalent ...
is a normal evolutionary process; nevertheless, hybridization (with or without introgression) threatens rare species' existence. The gene pool of a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
or a
population Population is a set of humans or other organisms in a given region or area. Governments conduct a census to quantify the resident population size within a given jurisdiction. The term is also applied to non-human animals, microorganisms, and pl ...
is the variety of genetic information in its living members. A large gene pool (extensive
genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is d ...
) is associated with robust populations that can survive bouts of intense selection. Meanwhile, low genetic diversity (see
inbreeding Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely genetic distance, related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genet ...
and population bottlenecks) reduces the range of adaptions possible. Replacing native with alien genes narrows genetic diversity within the original population, thereby increasing the chance of extinction.


Habitat degradation

Habitat degradation is currently the main anthropogenic cause of species extinctions. The main cause of habitat degradation worldwide is agriculture, with
urban sprawl Urban sprawl (also known as suburban sprawl or urban encroachment) is defined as "the spreading of urban developments (such as houses and shopping centers) on undeveloped land near a city". Urban sprawl has been described as the unrestricted ...
, logging, mining, and some fishing practices close behind. The degradation of a species'
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
may alter the fitness landscape to such an extent that the species is no longer able to survive and becomes extinct. This may occur by direct effects, such as the environment becoming toxic, or indirectly, by limiting a species' ability to compete effectively for diminished resources or against new competitor species. Habitat destruction, particularly the removal of vegetation that stabilizes soil, enhances erosion and diminishes nutrient availability in terrestrial ecosystems. This degradation can lead to a reduction in agricultural productivity. Furthermore, increased erosion contributes to poorer water quality by elevating the levels of sediment and pollutants in rivers and streams. Habitat degradation through toxicity can kill off a species very rapidly, by killing all living members through
contamination Contamination is the presence of a constituent, impurity, or some other undesirable element that renders something unsuitable, unfit or harmful for the physical body, natural environment, workplace, etc. Types of contamination Within the scien ...
or sterilizing them. It can also occur over longer periods at lower toxicity levels by affecting life span, reproductive capacity, or competitiveness. Habitat degradation can also take the form of a physical destruction of niche habitats. The widespread destruction of
tropical rainforest Tropical rainforests are dense and warm rainforests with high rainfall typically found between 10° north and south of the Equator. They are a subset of the tropical forest biome that occurs roughly within the 28° latitudes (in the torrid zo ...
s and replacement with open pastureland is widely cited as an example of this; elimination of the dense forest eliminated the infrastructure needed by many species to survive. For example, a
fern The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...
that depends on dense shade for protection from direct sunlight can no longer survive without forest to shelter it. Another example is the destruction of ocean floors by
bottom trawling Bottom trawling is trawling (towing a trawl, which is a fishing net) along the seafloor. It is also referred to as "dragging". The scientific community divides bottom trawling into benthic trawling and Demersal zone, demersal trawling. Benthic tra ...
. Diminished resources or introduction of new competitor species also often accompany habitat degradation.
Global warming Present-day climate change includes both global warming—the ongoing increase in global average temperature—and its wider effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes ...
has allowed some species to expand their range, bringing competition to other species that previously occupied that area. Sometimes these new competitors are predators and directly affect prey species, while at other times they may merely outcompete vulnerable species for limited resources. Vital resources including
water Water is an inorganic compound with the chemical formula . It is a transparent, tasteless, odorless, and Color of water, nearly colorless chemical substance. It is the main constituent of Earth's hydrosphere and the fluids of all known liv ...
and food can also be limited during habitat degradation, leading to extinction.


Predation, competition, and disease

In the natural course of events, species become extinct for a number of reasons, including but not limited to: extinction of a necessary host, prey or pollinator, interspecific competition, inability to deal with evolving diseases and changing environmental conditions (particularly sudden changes) which can act to introduce novel predators, or to remove prey. Recently in geological time, humans have become an additional cause of extinction of some species, either as a new mega-predator or by transporting
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s and
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s from one part of the world to another. Such introductions have been occurring for thousands of years, sometimes intentionally (e.g.
livestock Livestock are the Domestication, domesticated animals that are raised in an Agriculture, agricultural setting to provide labour and produce diversified products for consumption such as meat, Egg as food, eggs, milk, fur, leather, and wool. The t ...
released by sailors on islands as a future source of food) and sometimes accidentally (e.g. rats escaping from boats). In most cases, the introductions are unsuccessful, but when an invasive alien species does become established, the consequences can be catastrophic. Invasive alien species can affect native species directly by eating them, competing with them, and introducing
pathogen In biology, a pathogen (, "suffering", "passion" and , "producer of"), in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a Germ theory of d ...
s or
parasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted str ...
s that sicken or kill them; or indirectly by destroying or degrading their habitat. Human populations may themselves act as invasive predators. According to the "overkill hypothesis", the swift extinction of the megafauna in areas such as Australia (40,000 years before present),
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
(12,000 years before present),
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
,
Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
(AD 300–1000), and New Zealand (AD 1300–1500), resulted from the sudden introduction of human beings to environments full of animals that had never seen them before and were therefore completely unadapted to their predation techniques.Lee, Anita.
The Pleistocene Overkill Hypothesis
." ''University of California at Berkeley Geography Program.'' Retrieved January 11, 2007.


Coextinction

Coextinction refers to the loss of a species due to the extinction of another; for example, the extinction of parasitic insects following the loss of their hosts. Coextinction can also occur when a species loses its
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female carpel, stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are ...
, or to
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s in a
food chain A food chain is a linear network of links in a food web, often starting with an autotroph (such as grass or algae), also called a producer, and typically ending at an apex predator (such as grizzly bears or killer whales), detritivore (such as ...
who lose their prey. "Species coextinction is a manifestation of one of the interconnectednesses of organisms in complex ecosystems ... While coextinction may not be the most important cause of species extinctions, it is certainly an insidious one." Coextinction is especially common when a keystone species goes extinct. Models suggest that coextinction is the most common form of
biodiversity loss Biodiversity loss happens when plant or animal species disappear completely from Earth (extinction) or when there is a decrease or disappearance of species in a specific area. Biodiversity loss means that there is a reduction in Biodiversity, b ...
. There may be a cascade of coextinction across the trophic levels. Such effects are most severe in mutualistic and parasitic relationships. An example of coextinction is the Haast's eagle and the moa: the Haast's eagle was a predator that became extinct because its food source became extinct. The moa were several species of flightless birds that were a food source for the Haast's eagle.


Climate change

Extinction as a result of
climate change Present-day climate change includes both global warming—the ongoing increase in Global surface temperature, global average temperature—and its wider effects on Earth's climate system. Climate variability and change, Climate change in ...
has been confirmed by fossil studies. Particularly, the extinction of amphibians during the Carboniferous Rainforest Collapse, 305 million years ago. A 2003 review across 14 biodiversity research centers predicted that, because of climate change, 15–37% of land species would be "committed to extinction" by 2050. The ecologically rich areas that would potentially suffer the heaviest losses include the
Cape Floristic Region The Cape Floral Region is a floristic region located near the southern tip of South Africa. It is the only floristic region of the Cape Floristic Kingdom, and includes only one floristic province, known as the Cape Floristic Province. The Cap ...
and the Caribbean Basin. These areas might see a doubling of present carbon dioxide levels and rising temperatures that could eliminate 56,000 plant and 3,700 animal species. Climate change has also been found to be a factor in
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
and
desertification Desertification is a type of gradual land degradation of Soil fertility, fertile land into arid desert due to a combination of natural processes and human activities. The immediate cause of desertification is the loss of most vegetation. This i ...
.


Sexual selection and male investment

Studies of fossils following species from the time they evolved to their extinction show that species with high
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
, especially characteristics in males that are used to compete for mating, are at a higher risk of extinction and die out faster than less sexually dimorphic species, the least sexually dimorphic species surviving for millions of years while the most sexually dimorphic species die out within mere thousands of years. Earlier studies based on counting the number of currently living species in modern taxa have shown a higher number of species in more sexually dimorphic taxa which have been interpreted as higher survival in taxa with more sexual selection, but such studies of modern species only measure indirect effects of extinction and are subject to error sources such as dying and doomed taxa speciating more due to splitting of habitat ranges into more small isolated groups during the habitat retreat of taxa approaching extinction. Possible causes of the higher extinction risk in species with more sexual selection shown by the comprehensive fossil studies that rule out such error sources include expensive sexually selected ornaments having negative effects on the ability to survive
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generation ...
, as well as
sexual selection Sexual selection is a mechanism of evolution in which members of one sex mate choice, choose mates of the other sex to mating, mate with (intersexual selection), and compete with members of the same sex for access to members of the opposite sex ...
removing a diversity of genes that under current ecological conditions are neutral for natural selection but some of which may be important for surviving climate change.


Mass extinctions

There have been at least five mass extinctions in the history of life on earth, and four in the last 350 million years in which many species have disappeared in a relatively short period of geological time. A massive eruptive event that released large quantities of
tephra Tephra is fragmental material produced by a Volcano, volcanic eruption regardless of composition, fragment size, or emplacement mechanism. Volcanologists also refer to airborne fragments as pyroclasts. Once clasts have fallen to the ground, ...
particles into the atmosphere is considered to be one likely cause of the "
Permian–Triassic extinction event The Permian–Triassic extinction event (also known as the P–T extinction event, the Late Permian extinction event, the Latest Permian extinction event, the End-Permian extinction event, and colloquially as the Great Dying,) was an extinction ...
" about 250 million years ago, which is estimated to have killed 90% of species then existing. There is also evidence to suggest that this event was preceded by another mass extinction, known as Olson's Extinction. The
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event, also known as the K–T extinction, was the extinction event, mass extinction of three-quarters of the plant and animal species on Earth approximately 66 million years ago. The event cau ...
(K–Pg) occurred 66 million years ago, at the end of the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
period; it is best known for having wiped out non-avian
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s, among many other species.


Modern extinctions

According to a 1998 survey of 400 biologists conducted by New York's American Museum of Natural History, nearly 70% believed that the Earth is currently in the early stages of a human-caused mass extinction, American Museum of Natural History.
National Survey Reveals Biodiversity Crisis – Scientific Experts Believe We are in the Midst of the Fastest Mass Extinction in Earth's History
". Retrieved September 20, 2006.
known as the
Holocene extinction The Holocene extinction, also referred to as the Anthropocene extinction or the sixth mass extinction, is an ongoing extinction event caused exclusively by human activities during the Holocene epoch. This extinction event spans numerous families ...
. In that survey, the same proportion of respondents agreed with the prediction that up to 20% of all living populations could become extinct within 30 years (by 2028). A 2014 special edition of ''
Science Science is a systematic discipline that builds and organises knowledge in the form of testable hypotheses and predictions about the universe. Modern science is typically divided into twoor threemajor branches: the natural sciences, which stu ...
'' declared there is widespread consensus on the issue of human-driven mass species extinctions. A 2020 study published in ''
PNAS ''Proceedings of the National Academy of Sciences of the United States of America'' (often abbreviated ''PNAS'' or ''PNAS USA'') is a peer-reviewed multidisciplinary scientific journal. It is the official journal of the National Academy of S ...
'' stated that the contemporary extinction crisis "may be the most serious environmental threat to the persistence of civilization, because it is irreversible." A 2025 study found that human activities are to blame for biodiversity loss across all species and ecosystems. Biologist E. O. Wilson estimated in 2002 that if current rates of human destruction of the biosphere continue, one-half of all plant and animal species of life on earth will be extinct in 100 years. E.O. Wilson repeats his estimation in 2012. More significantly, the current rate of global species extinctions is estimated as 100 to 1,000 times "background" rates (the average extinction rates in the
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
ary time scale of planet Earth), faster than at any other time in human history, while future rates are likely 10,000 times higher. However, some groups are going extinct much faster. Biologists Paul R. Ehrlich and Stuart Pimm, among others, contend that human
population growth Population growth is the increase in the number of people in a population or dispersed group. The World population, global population has grown from 1 billion in 1800 to 8.2 billion in 2025. Actual global human population growth amounts to aroun ...
and
overconsumption Overconsumption describes a situation where consumers overuse their available goods and services to where they can't, or don't want to, replenish or reuse them. In microeconomics, this is the point where the marginal cost of a consumer is greater ...
are the main drivers of the modern extinction crisis. In January 2020, the UN's Convention on Biological Diversity drafted a plan to mitigate the contemporary extinction crisis by establishing a deadline of 2030 to protect 30% of the Earth's land and oceans and reduce pollution by 50%, with the goal of allowing for the restoration of ecosystems by 2050. The 2020
United Nations The United Nations (UN) is the Earth, global intergovernmental organization established by the signing of the Charter of the United Nations, UN Charter on 26 June 1945 with the stated purpose of maintaining international peace and internationa ...
' ''Global Biodiversity Outlook'' report stated that of the 20 biodiversity goals laid out by the Aichi Biodiversity Targets in 2010, only 6 were "partially achieved" by the deadline of 2020. The report warned that biodiversity will continue to decline if the status quo is not changed, in particular the "currently unsustainable patterns of production and consumption, population growth and technological developments". In a 2021 report published in the journal ''Frontiers in Conservation Science'', some top scientists asserted that even if the Aichi Biodiversity Targets set for 2020 had been achieved, it would not have resulted in a significant mitigation of biodiversity loss. They added that failure of the global community to reach these targets is hardly surprising given that biodiversity loss is "nowhere close to the top of any country's priorities, trailing far behind other concerns such as employment, healthcare, economic growth, or currency stability."


History of scientific understanding

For much of history, the modern understanding of extinction as the end of a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
was incompatible with the prevailing worldview. Prior to the 19th century, much of Western society adhered to the belief that the world was created by God and as such was complete and perfect. This concept reached its heyday in the 1700s with the peak popularity of a theological concept called the
great chain of being The great chain of being is a hierarchical structure of all matter and life, thought by medieval Christianity to have been decreed by God. The chain begins with God and descends through angels, Human, humans, Animal, animals and Plant, plants to ...
, in which all life on earth, from the tiniest microorganism to God, is linked in a continuous chain. The extinction of a species was impossible under this model, as it would create gaps or missing links in the chain and destroy the natural order.
Thomas Jefferson Thomas Jefferson (, 1743July 4, 1826) was an American Founding Fathers of the United States, Founding Father and the third president of the United States from 1801 to 1809. He was the primary author of the United States Declaration of Indepe ...
was a firm supporter of the great chain of being and an opponent of extinction, famously denying the extinction of the woolly mammoth on the grounds that nature never allows a race of animals to become extinct. A series of fossils were discovered in the late 17th century that appeared unlike any living species. As a result, the scientific community embarked on a voyage of creative rationalization, seeking to understand what had happened to these species within a framework that did not account for total extinction. In October 1686, Robert Hooke presented an impression of a
nautilus A nautilus (; ) is any of the various species within the cephalopod family Nautilidae. This is the sole extant family of the superfamily Nautilaceae and the suborder Nautilina. It comprises nine living species in two genera, the type genus, ty ...
to the
Royal Society The Royal Society, formally The Royal Society of London for Improving Natural Knowledge, is a learned society and the United Kingdom's national academy of sciences. The society fulfils a number of roles: promoting science and its benefits, re ...
that was more than two feet in diameter, and morphologically distinct from any known living species. Hooke theorized that this was simply because the species lived in the deep ocean and no one had discovered them yet. While he contended that it was possible a species could be "lost", he thought this highly unlikely. Similarly, in 1695, Sir Thomas Molyneux published an account of enormous antlers found in
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
that did not belong to any extant taxa in that area. Molyneux reasoned that they came from the North American
moose The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...
and that the animal had once been common on the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
. Rather than suggest that this indicated the possibility of species going extinct, he argued that although organisms could become locally extinct, they could never be entirely lost and would continue to exist in some unknown region of the globe. The antlers were later confirmed to be from the extinct deer '' Megaloceros''. Hooke and Molyneux's line of thinking was difficult to disprove. When parts of the world had not been thoroughly examined and charted, scientists could not rule out that animals found only in the fossil record were not simply "hiding" in unexplored regions of the Earth.''Ideas: A History from Fire to Freud'' ( Peter Watson Weidenfeld & Nicolson )
Georges Cuvier Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...
is credited with establishing the modern conception of extinction in a 1796 lecture to the French Institute, though he would spend most of his career trying to convince the wider scientific community of his theory. Cuvier was a well-regarded geologist, lauded for his ability to reconstruct the anatomy of an unknown species from a few fragments of bone. His primary evidence for extinction came from mammoth skulls found near
Paris Paris () is the Capital city, capital and List of communes in France with over 20,000 inhabitants, largest city of France. With an estimated population of 2,048,472 residents in January 2025 in an area of more than , Paris is the List of ci ...
. Cuvier recognized them as distinct from any known living species of elephant, and argued that it was highly unlikely such an enormous animal would go undiscovered. In 1798, he studied a fossil from the Paris Basin that was first observed by Robert de Lamanon in 1782, first hypothesizing that it belonged to a canine but then deciding that it instead belonged to an animal that was unlike living ones. His study paved the way to his naming of the extinct mammal genus '' Palaeotherium'' in 1804 based on the skull and additional fossil material along with another extinct contemporary mammal genus '' Anoplotherium''. In both genera, he noticed that their fossils shared some similarities with other mammals like
ruminant Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microb ...
s and
rhinoceros A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant taxon, extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family (biology), famil ...
es but still had distinct differences. In 1812, Cuvier, along with Alexandre Brongniart and Geoffroy Saint-Hilaire, mapped the
strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
of the Paris basin. They saw alternating saltwater and freshwater deposits, as well as patterns of the appearance and disappearance of fossils throughout the record. From these patterns, Cuvier inferred historic cycles of catastrophic flooding, extinction, and repopulation of the earth with new species. Cuvier's fossil evidence showed that very different life forms existed in the past than those that exist today, a fact that was accepted by most scientists. The primary debate focused on whether this turnover caused by extinction was gradual or abrupt in nature. Cuvier understood extinction to be the result of cataclysmic events that wipe out huge numbers of species, as opposed to the gradual decline of a species over time. His catastrophic view of the nature of extinction garnered him many opponents in the newly emerging school of uniformitarianism.
Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biologi ...
, a gradualist and colleague of Cuvier, saw the fossils of different life forms as evidence of the mutable character of species. While Lamarck did not deny the possibility of extinction, he believed that it was exceptional and rare and that most of the change in species over time was due to gradual change. Unlike Cuvier, Lamarck was skeptical that catastrophic events of a scale large enough to cause total extinction were possible. In his geological history of the earth titled Hydrogeologie, Lamarck instead argued that the surface of the earth was shaped by gradual erosion and deposition by water, and that species changed over time in response to the changing environment. Charles Lyell, a noted geologist and founder of uniformitarianism, believed that past processes should be understood using present day processes. Like Lamarck, Lyell acknowledged that extinction could occur, noting the total extinction of the
dodo The dodo (''Raphus cucullatus'') is an extinction, extinct flightless bird that was endemism, endemic to the island of Mauritius, which is east of Madagascar in the Indian Ocean. The dodo's closest relative was the also-extinct and flightles ...
and the extirpation of indigenous horses to the British Isles. He similarly argued against mass extinctions, believing that any extinction must be a gradual process. Lyell also showed that Cuvier's original interpretation of the Parisian strata was incorrect. Instead of the catastrophic floods inferred by Cuvier, Lyell demonstrated that patterns of saltwater and freshwater deposits, like those seen in the Paris basin, could be formed by a slow rise and fall of
sea level Mean sea level (MSL, often shortened to sea level) is an mean, average surface level of one or more among Earth's coastal Body of water, bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical ...
s. The concept of extinction was integral to
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
's ''
On the Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life'')The book's full original title was ''On the Origin of Species by M ...
'', with less fit lineages disappearing over time. For Darwin, extinction was a constant side effect of
competition Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indi ...
. Because of the wide reach of ''On the Origin of Species'', it was widely accepted that extinction occurred gradually and evenly (a concept now referred to as background extinction). It was not until 1982, when David Raup and Jack Sepkoski published their seminal paper on mass extinctions, that Cuvier was vindicated and catastrophic extinction was accepted as an important mechanism. The current understanding of extinction is a synthesis of the cataclysmic extinction events proposed by Cuvier, and the background extinction events proposed by Lyell and Darwin.


Human attitudes and interests

Extinction is an important research topic in the field of
zoology Zoology ( , ) is the scientific study of animals. Its studies include the anatomy, structure, embryology, Biological classification, classification, Ethology, habits, and distribution of all animals, both living and extinction, extinct, and ...
, and
biology Biology is the scientific study of life and living organisms. It is a broad natural science that encompasses a wide range of fields and unifying principles that explain the structure, function, growth, History of life, origin, evolution, and ...
in general, and has also become an area of concern outside the scientific community. A number of organizations, such as the Worldwide Fund for Nature, have been created with the goal of preserving species from extinction.
Government A government is the system or group of people governing an organized community, generally a State (polity), state. In the case of its broad associative definition, government normally consists of legislature, executive (government), execu ...
s have attempted, through enacting laws, to avoid habitat destruction, agricultural over-harvesting, and
pollution Pollution is the introduction of contaminants into the natural environment that cause harm. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the component ...
. While many human-caused extinctions have been accidental, humans have also engaged in the deliberate destruction of some species, such as dangerous
virus A virus is a submicroscopic infectious agent that replicates only inside the living Cell (biology), cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Viruses are ...
es, and the total destruction of other problematic species has been suggested. Other species were deliberately driven to extinction, or nearly so, due to
poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...
or because they were "undesirable", or to push for other human agendas. One example was the near extinction of the
American bison The American bison (''Bison bison''; : ''bison''), commonly known as the American buffalo, or simply buffalo (not to be confused with Bubalina, true buffalo), is a species of bison that is endemic species, endemic (or native) to North America. ...
, which was nearly wiped out by mass hunts sanctioned by the United States government, to force the removal of Native Americans, many of whom relied on the bison for food. Biologist Bruce Walsh states three reasons for scientific interest in the preservation of species: genetic resources, ecosystem stability, and
ethics Ethics is the philosophy, philosophical study of Morality, moral phenomena. Also called moral philosophy, it investigates Normativity, normative questions about what people ought to do or which behavior is morally right. Its main branches inclu ...
; and today the scientific community "stress sthe importance" of maintaining biodiversity. Walsh, Bruce
Extinction
. Bioscience at University of Arizona. Retrieved July 26, 2006.
Committee on Recently Extinct Organisms.

". Retrieved July 30, 2006.
In modern times, commercial and industrial interests often have to contend with the effects of production on plant and animal life. However, some technologies with minimal, or no, proven harmful effects on ''
Homo sapiens Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
'' can be devastating to wildlife (for example, DDT).International Programme on Chemical Safety (1989).
DDT and its Derivatives – Environmental Aspects
". Environmental Health Criteria 83. Retrieved September 20, 2006.
Biogeographer Jared Diamond notes that while big business may label environmental concerns as "exaggerated", and often cause "devastating damage", some corporations find it in their interest to adopt good conservation practices, and even engage in preservation efforts that surpass those taken by
national park A national park is a nature park designated for conservation (ethic), conservation purposes because of unparalleled national natural, historic, or cultural significance. It is an area of natural, semi-natural, or developed land that is protecte ...
s. Governments sometimes see the loss of native species as a loss to
ecotourism Ecotourism is a form of nature-oriented tourism intended to contribute to the Ecological conservation, conservation of the natural environment, generally defined as being minimally impactful, and including providing both contributions to conserv ...
,Drewry, Rachel.
Ecotourism: Can it save the orangutans?
" ''Inside Indonesia''. Retrieved January 26, 2007.
and can enact laws with severe punishment against the trade in native species in an effort to prevent extinction in the wild. Nature preserves are created by governments as a means to provide continuing habitats to species crowded by human expansion. The 1992 Convention on Biological Diversity has resulted in international Biodiversity Action Plan programmes, which attempt to provide comprehensive guidelines for government biodiversity conservation. Advocacy groups, such as The Wildlands ProjectThe Wildlands Project
. Retrieved January 26, 2007.
and the Alliance for Zero Extinctions, work to educate the public and pressure governments into action. People who live close to nature can be dependent on the survival of all the species in their environment, leaving them highly exposed to extinction
risk In simple terms, risk is the possibility of something bad happening. Risk involves uncertainty about the effects/implications of an activity with respect to something that humans value (such as health, well-being, wealth, property or the environ ...
s. However, people prioritize day-to-day survival over species conservation; with
human overpopulation Human overpopulation (or human population overshoot) is the idea that human populations may become too large to be sustainability, sustained by their environment or resources in the long term. The topic is usually discussed in the context of wor ...
in tropical
developing countries A developing country is a sovereign state with a less-developed Secondary sector of the economy, industrial base and a lower Human Development Index (HDI) relative to developed countries. However, this definition is not universally agreed upon. ...
, there has been enormous pressure on forests due to subsistence agriculture, including
slash-and-burn Slash-and-burn agriculture is a form of shifting cultivation that involves the cutting and burning of plants in a forest or woodland to create a Field (agriculture), field called a swidden. The method begins by cutting down the trees and woody p ...
agricultural techniques that can reduce endangered species's habitats. Antinatalist philosopher David Benatar concludes that any popular concern about non-human species extinction usually arises out of concern about how the loss of a species will impact human wants and needs, that "we shall live in a world impoverished by the loss of one aspect of faunal diversity, that we shall no longer be able to behold or use that species of animal." He notes that typical concerns about possible human extinction, such as the loss of individual members, are not considered in regards to non-human species extinction. Anthropologist Jason Hickel speculates that the reason humanity seems largely indifferent to anthropogenic mass species extinction is that we see ourselves as separate from the natural world and the organisms within it. He says that this is due in part to the logic of
capitalism Capitalism is an economic system based on the private ownership of the means of production and their use for the purpose of obtaining profit. This socioeconomic system has developed historically through several stages and is defined by ...
: "that the world is not really alive, and it is certainly not our kin, but rather just stuff to be extracted and discarded – and that includes most of the human beings living here too."


Planned extinction


Completed

* The
smallpox Smallpox was an infectious disease caused by Variola virus (often called Smallpox virus), which belongs to the genus '' Orthopoxvirus''. The last naturally occurring case was diagnosed in October 1977, and the World Health Organization (W ...
virus is now extinct in the wild, although samples are retained in laboratory settings. * The
rinderpest Rinderpest (also cattle plague or steppe murrain) was an infectious viral disease of cattle, domestic water buffalo, and many other species of even-toed ungulates, including gaurs, African Buffalo, buffaloes, large antelope, deer, giraffes, wilde ...
virus, which infected domestic cattle, is now extinct in the wild.


Proposed


= Disease agents

= The poliovirus is now confined to small parts of the world due to extermination efforts. '' Dracunculus medinensis'', or Guinea worm, a parasitic worm which causes the disease dracunculiasis, is now close to eradication thanks to efforts led by the Carter Center. '' Treponema pallidum pertenue'', a bacterium which causes the disease yaws, is in the process of being eradicated.


= Disease vectors

= Biologist Olivia Judson has advocated the deliberate extinction of certain disease-carrying
mosquito Mosquitoes, the Culicidae, are a Family (biology), family of small Diptera, flies consisting of 3,600 species. The word ''mosquito'' (formed by ''Musca (fly), mosca'' and diminutive ''-ito'') is Spanish and Portuguese for ''little fly''. Mos ...
species. In an article in ''
The New York Times ''The New York Times'' (''NYT'') is an American daily newspaper based in New York City. ''The New York Times'' covers domestic, national, and international news, and publishes opinion pieces, investigative reports, and reviews. As one of ...
'' on 25 September 2003, she advocated "specicide" of thirty mosquito species by introducing a genetic element that can insert itself into another crucial gene, to create recessive " knockout genes". She says that the '' Anopheles'' mosquitoes (which spread
malaria Malaria is a Mosquito-borne disease, mosquito-borne infectious disease that affects vertebrates and ''Anopheles'' mosquitoes. Human malaria causes Signs and symptoms, symptoms that typically include fever, Fatigue (medical), fatigue, vomitin ...
) and ''
Aedes ''Aedes'' (also known as the tiger mosquito) is a genus of mosquitoes originally found in tropical and subtropical zones, but now found on all continents except Antarctica. Some species have been spread by human activity: ''Aedes albopictus'', ...
'' mosquitoes (which spread
dengue fever Dengue fever is a mosquito-borne disease caused by dengue virus, prevalent in tropical and subtropical areas. Asymptomatic infections are uncommon, mild cases happen frequently; if symptoms appear, they typically begin 3 to 14 days after i ...
, yellow fever, elephantiasis, and other diseases) represent only 30 of around 3,500 mosquito species; eradicating these would save at least one million human lives per year, at a cost of reducing the
genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is d ...
of the family
Culicidae Mosquitoes, the Culicidae, are a Family (biology), family of small Diptera, flies consisting of 3,600 species. The word ''mosquito'' (formed by ''Musca (fly), mosca'' and diminutive ''-ito'') is Spanish and Portuguese for ''little fly''. Mos ...
by only 1%. She further argues that since species become extinct "all the time" the disappearance of a few more will not destroy the
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
: "We're not left with a wasteland every time a species vanishes. Removing one species sometimes causes shifts in the populations of other species—but different need not mean worse." In addition, anti-malarial and mosquito control programs offer little realistic hope to the 300 million people in developing nations who will be infected with acute illnesses this year. Although trials are ongoing, she writes that if they fail "we should consider the ultimate swatting". Biologist E. O. Wilson has advocated the eradication of several species of mosquito, including malaria vector '' Anopheles gambiae''. Wilson stated, "I'm talking about a very small number of species that have co-evolved with us and are preying on humans, so it would certainly be acceptable to remove them. I believe it's just common sense." There have been many campaigns – some successful – to locally eradicate tsetse flies and their trypanosomes in areas, countries, and islands of Africa (including
Príncipe Príncipe (; ) is the smaller, northern major island of the country of São Tomé and Príncipe lying off the west coast of Africa in the Gulf of Guinea. It has an area of (including offshore islets) and a population of 7,324 at the 2012 Cens ...
). (Other ) There are currently serious efforts to do away with them all across Africa, and this is generally viewed as beneficial and morally necessary, although not always.


Cloning

Some, such as Harvard geneticist George M. Church, believe that ongoing technological advances will let us "bring back to life" an extinct species by
cloning Cloning is the process of producing individual organisms with identical genomes, either by natural or artificial means. In nature, some organisms produce clones through asexual reproduction; this reproduction of an organism by itself without ...
, using
DNA Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...
from the remains of that species. Proposed targets for cloning include the
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus.'' They lived from the late Miocene epoch (from around 6.2 million years ago) into the Holocene until about 4,000 years ago, with mammoth species at various times inhabi ...
, the
thylacine The thylacine (; binomial name ''Thylacinus cynocephalus''), also commonly known as the Tasmanian tiger or Tasmanian wolf, was a carnivorous marsupial that was native to the Mainland Australia, Australian mainland and the islands of Tasmani ...
, and the Pyrenean ibex. For this to succeed, enough individuals would have to be cloned, from the DNA of different individuals (in the case of sexually reproducing organisms) to create a viable population. Though bioethical and philosophical objections have been raised, the cloning of extinct creatures seems theoretically possible. In 2003, scientists tried to clone the extinct Pyrenean ibex (''C. p. pyrenaica''). This attempt failed: of the 285 embryos reconstructed, 54 were transferred to 12 Spanish ibexes and ibex–domestic
goat The goat or domestic goat (''Capra hircus'') is a species of Caprinae, goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the ...
hybrids, but only two survived the initial two months of gestation before they, too, died. In 2009, a second attempt was made to clone the Pyrenean ibex: one clone was born alive, but died seven minutes later, due to physical defects in the lungs.


See also


References


Further reading

* * * *


External links


Committee on recently extinct organisms

The age of extinction
series in ''
The Guardian ''The Guardian'' is a British daily newspaper. It was founded in Manchester in 1821 as ''The Manchester Guardian'' and changed its name in 1959, followed by a move to London. Along with its sister paper, ''The Guardian Weekly'', ''The Guardi ...
''
Extincion events connected with New theory of evolution
{{Authority control Biota by conservation status Environmental conservation Evolutionary biology IUCN Red List