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''Euplokamis'' is a genus of
ctenophore Ctenophora (; ctenophore ; ) comprise a phylum of marine invertebrates, commonly known as comb jellies, that inhabit sea waters worldwide. They are notable for the groups of cilia they use for swimming (commonly referred to as "combs"), and ...
s, or comb jellies, belonging to the monotypic family Euplokamididae. Despite living for hundreds of millions of years in marine environments, there is minimal research regarding ''Euplokamis'', primarily due to their body structure. Research on the evolution of the basic body structures of
diploblastic Diploblasty is a condition of the blastula in which there are two primary germ layers: the ectoderm and endoderm. Diploblastic organisms are organisms which develop from such a blastula, and include cnidaria and ctenophora, formerly grouped togeth ...
metazoans revealed that there are four major phyla, including the Ctenophores. Although the morphology of ''Euplokamis'' often resembles the medusa stage of
Cnidaria Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that th ...
ns, their eight rows of combs are one distinguishing feature that led to the official classification of Ctenophores. After being originally described by Chun (1879), the family '' Euplokamididae'' was expanded by Mills (1987) due to the discovery of a new species, ''
Euplokamis dunlapae ''Euplokamis dunlapae'' is a marine species of ctenophore. It is the first species of ctenophora reported to have giant axons controlling the comb rows. They control the ciliary beating, allowing for rapid change in the speed and direction of the ...
''. Further research indicated that ''Euplokamis'' should be identified from Mertensiidae due to the rows of combs and some compression. They may also be distinguished from the genus ''
Pleurobrachia ''Pleurobrachia'' is a common genus of Ctenophora(a exclusively marine phylum). Along with the genus '' Hormiphora'', it generally has the common name sea gooseberry. It contains the following species: *''Pleurobrachia arctica'' Wagner, 1885 *' ...
'' due to their more elongated shape. Additionally, various adaptations of ''Euplokamis'' have been observed such as the use of tentacles for movement/feeding, a complex nervous system, and bioluminescent capabilities. Other characteristics including a defined
mesoderm The mesoderm is the middle layer of the three germ layers that develops during gastrulation in the very early development of the embryo of most animals. The outer layer is the ectoderm, and the inner layer is the endoderm.Langman's Medical E ...
, lack of stinging cells, developmental differences, and symmetry supported the reclassification of these organisms.


Distribution & Habitat

''Euplokamis'' have been found in fossil records dating back to the
Cambrian period The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
, part of the Paleozoic Era, and it is estimated that some species may have evolved before this period. Originally,
Cnidaria Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that th ...
ns and
Ctenophores Ctenophora (; ctenophore ; ) comprise a phylum of marine invertebrates, commonly known as comb jellies, that inhabit sea waters worldwide. They are notable for the groups of cilia they use for swimming (commonly referred to as "combs"), a ...
were classified under the same phyla, Coelenterata. Ctenophore bodies are made up of a gelatin substance, similar to Cnidarians, but the multiple rows of combs present in fossil records are unique to ctenophores. Records of ''Euplokamis'' sp. indicate they are distributed widely around the world, but are most often found in warm coastal waters. ''Euplokamis'' prefer marine, or saltwater, environments and are classified as free swimmers, due to their ability to move through the water column. They have been identified in the Mediterranean Sea, the North Pacific, the Gulf of Maine, and off the coast of Sweden. However, since this genus was originally grouped in the family
Pleurobrachiidae Pleurobrachiidae is a family of ctenophores belonging to the order Cydippida. Genera: * ''Hormiphora'' L.Agassiz, 1860 * ''Minictena'' C.Carré & D.Carré, 1993 * ''Moseria (ctenophore), Moseria'' Ghigi, 1909 * ''Pleurobrachia'' Fleming, 182 ...
, there is limited information regarding their actual distribution and habitat.


Anatomy & Morphology

Ctenophores are divided into two classes based on either the presence(
Tentaculat ''XCOM'' (originally called ''X-COM'') is a science fiction video game franchise featuring an elite international organization tasked with countering alien invasions of Earth. The series began with the strategy video game '' X-COM: UFO Defense' ...
)) or the lack (
Nuda Beroidae is a family of ctenophores or comb jellies more commonly referred to as the beroids. It is the only family within the monotypic order Beroida and the class Nuda. They are distinguished from other comb jellies by the complete absence of ...
) of tentacles. Within each class, there are multiple orders to further distinguish their structures and characteristics. The class
Tentaculata Tentaculata is a class of comb jellies. The common feature of this class is a pair of long, feathery, contractile tentacles, which can be retracted into specialised ciliated sheaths. In some species, the primary tentacles are reduced and they h ...
contains the following orders:
Cydippida Cydippida is an order of comb jellies. They are distinguished from other comb jellies by their spherical or oval bodies, and the fact their tentacles are branched, and can be retracted into pouches on either side of the pharynx. The order is not ...
,
Lobata Lobata is an order of Ctenophora in the class Tentaculata with smaller tentacles than other ctenophores, and distinctive flattened lobes extending outwards from their bodies. They grow up to about long. Anatomy The lobates have a pair of lo ...
, and
Cestida Cestidae is a family of comb jellies. It is the only family in the monotypic order Cestida. Unlike other comb jellies, the body of cestids is greatly flattened, and drawn out into a long ribbon-like shape. The two tentacles are greatly shortened, ...
. The genus ''Euplokamis'' is part of the class tentaculate, which indicates that tentacles are present. They are also part of the order
Cydippida Cydippida is an order of comb jellies. They are distinguished from other comb jellies by their spherical or oval bodies, and the fact their tentacles are branched, and can be retracted into pouches on either side of the pharynx. The order is not ...
, distinguished by their tentacles and their round body shape. These tentacles are long, may display branching, and have a sheath allowing them to be retractile. The branching tentacles are known as
tentilla In zoology, a tentacle is a flexible, mobile, and elongated organ present in some species of animals, most of them invertebrates. In animal anatomy, tentacles usually occur in one or more pairs. Anatomically, the tentacles of animals work mainl ...
, which are held tightly in coils. Further, the widely spaced tentillia droplets allow for organisms to be classified to the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
level and are one of the only examples of striated muscle found in ctenophores. Additionally, these organisms have bi-radial symmetry with a mouth on their front end and a statocyst, or sense organ, at the other end. The sides of their stomachs are lined with distinct bulbs, shaped like tadpoles, and unlike other well-known jellyfish, ''Euplokamis'' do not have any
nematocytes A cnidocyte (also known as a cnidoblast or nematocyte) is an explosive cell containing one large secretory organelle called a cnidocyst (also known as a cnida () or nematocyst) that can deliver a sting to other organisms. The presence of this c ...
, known as stinging cells. The mouth is connected to the digestive tract via the
pharynx The pharynx (plural: pharynges) is the part of the throat behind the mouth and nasal cavity, and above the oesophagus and trachea (the tubes going down to the stomach and the lungs). It is found in vertebrates and invertebrates, though its struc ...
. The digestive system, or gastrovascular cavity, is made up of intricate canals that allow for both digestion and circulation to occur. They also lack an
anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, d ...
but are able to excrete some waste through pores on the
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