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''Cteniogenys'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of choristodere, a morphologically diverse group of aquatic reptiles. It is part of the monotypic family Cteniogenidae. The type and only named species, ''C. antiquus'', was named in 1928 by Charles W. Gilmore. The holotype, VP.001088, was collected in the
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
(Como Bluff),
Wyoming Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
in 1881 by William H. Reed. More specimens have been discovered since then, including specimens from the Late Jurassic of Portugal and Middle Jurassic of Britain, which have not been assigned to species.


Description

''Cteniogenys'' was long, and probably weighed less than . The skull of this genus was long and slender, and the jaws had numerous conical teeth. ''Cteniogenys'' in the Morrison probably fed on insects and small fish. It is mostly known from freshwater sites (rivers and ponds), and is a rare find in the formation (only 60 specimens out of over 2,800 total vertebrate specimens known from the formation), mostly known from northern outcrops (particularly
Wyoming Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
); this may reflect a
preservation bias Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientist Ivan Efremov ...
against small animals in terrestrial settings, rather than an accurate reflection of ''Cteniogenys'' populations in the Morrison.


Distribution

Fossils of ''Cteniogenys'' are known from the upper Middle Jurassic (
Bathonian In the geologic timescale the Bathonian is an age (geology), age and stage (stratigraphy), stage of the Middle Jurassic. It lasted from approximately 168.2 ±1.2 annum, Ma to around 165.3 ±1.1 Ma (million years ago). The Bathonian Age succeeds ...
) aged Forest Marble Formation and Kilmaluag Formation of Britain, the
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
(
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
) aged
Alcobaça Formation The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit, is a geological Formation (geology), formation in Portugal. It dates back to the Kimmeridgian stage of the Late Jurassic. It is an impor ...
of Portugal, and the Late Jurassic-age
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
of western
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, indeterminate similar remains are also known from the
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
aged Balabansai Formation of
Kyrgyzstan Kyrgyzstan, officially the Kyrgyz Republic, is a landlocked country in Central Asia lying in the Tian Shan and Pamir Mountains, Pamir mountain ranges. Bishkek is the Capital city, capital and List of cities in Kyrgyzstan, largest city. Kyrgyz ...
and the Bathonian aged
Itat Formation The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from ...
of western Siberia, and the Berriasian aged
Angeac-Charente bonebed The Angeac-Charente bonebed is a fossil deposit located near Angeac-Charente in western France. It dates to the Berriasian stage of the Early Cretaceous, and is coeval with the Purbeck Group of Southern England. It has amongst the most diverse asse ...
in France.Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, et al.
Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
Geodiversitas, Museum National d’Histoire Naturelle Paris, In press. ffhal-03264773f
A handful of skull and jaw fragments from the Late Cretaceous Oldman Formation and
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
in Canada were assigned to ''Cteniogenys'' by Gao and Fox (1998). Given the long gap in time, however, Gao et al. (2005) cautioned that these remains could constitute a different, as-yet undetermined genus. In their description of new material of '' Khurendukhosaurus'', Matsumoto et al. (2009) agreed that the putative Cretaceous occurrence of ''Cteniogenys'' did not belong to that genus and represents an indeterminate choristoderan, possibly a new genus.


Systematics

The genus was named in 1928 by Charles W. Gilmore on the basis of a lower jaw collected during the late 19th century by
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of paleontology. A prolific fossil collector, Marsh was one of the preeminent paleontologists of the nineteenth century. Among his legacies are the discovery or ...
's workers at Como Bluff. He tentatively described the genus as a lizard, noting that it could instead be a frog. New material of ''Cteniogenys'' from Europe prompted
Susan E. Evans Susan Elizabeth Evans is a British palaeontologist and herpetologist. She is the author or co-author of over 100 peer-reviewed papers and book chapters. She received a BSc in Zoology at Bedford College (London), Bedford College in 1974, and i ...
to reclassify it as an early choristodere in 1989. To date it is only known from fragmentary specimens. In phylogenic analyses it has been found to the basal most choristodere.


See also

* Paleobiota of the Morrison Formation


References

{{Taxonbar, from1=Q5191621, from2=Q2727696 Choristodera Jurassic reptiles Late Jurassic reptiles of North America Jurassic Colorado Jurassic geology of Oklahoma Jurassic geology of South Dakota Jurassic geology of Utah Jurassic geology of Wyoming Late Cretaceous reptiles of North America Fossil taxa described in 1928 Prehistoric reptile genera