A cloven hoof, cleft hoof, divided hoof, or split hoof is a

hoof The hoof (: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits; the ruminants with ...

split into two toes. Members of the

mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...

ian order

Artiodactyla Artiodactyls are placental mammals belonging to the order (biology), order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof ...

that possess this type of hoof include

cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...

deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...

, pigs,

antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...

gazelle A gazelle is one of many antelope species in the genus ''Gazella'' . There are also seven species included in two further genera; '' Eudorcas'' and '' Nanger'', which were formerly considered subgenera of ''Gazella''. A third former subgenus, ' ...

goat The goat or domestic goat (''Capra hircus'') is a species of Caprinae, goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the ...

s, and

sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...

. The two digits of cloven-hooved animals are homologous to the third and fourth fingers of the

hand A hand is a prehensile, multi-fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the Koala#Characteristics, koala (which has two thumb#O ...

. They are called claws and are named for their relative location on the foot: the outer, or

lateral Lateral is a geometric term of location which may also refer to: Biology and healthcare * Lateral (anatomy), a term of location meaning "towards the side" * Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx * Lateral release ( ...

, claw and the inner, or medial claw. The space between the two claws is called the interdigital cleft; the area of skin is called the interdigital skin. The hard outer covering of the hoof is called the hoof wall or horn. It is a hard surface, similar to the human fingernail. The almost finger-like dexterity available to cloven-hooved mammals such as

mountain goat The mountain goat (''Oreamnos americanus''), also known as the Rocky Mountain goat, is a cloven-footed mammal that is endemic to the remote and rugged mountainous areas of western North America. A subalpine to truly alpine species, it is a s ...

s and wild sheep combined with a hard outer shell and soft and flexible inner pads provides excellent traction in their precarious habitats.

Evolution

It is speculated by paleontologists that during the

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

period hooved marsh dwellers carried their body weight mainly on two of the middle toes, which grew to equal size, becoming the

or even-toed hooved animals. Before the end of the Eocene period the side toes of some had dwindled and practically disappeared (mainly in the form of a

dewclaw A dewclaw is a digit – vestigial in some animals – on the foot of many mammals, birds, and reptiles (including some extinct orders, like certain theropods). It commonly grows higher on the leg than the rest of the foot, such that in di ...

) while the basal pieces or metapodium of the pair of supporting toes became fused together, thus producing the appearance of a cloven hoof. The mammal with a cloven hoof is an even-toed

ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...

of order Artiodactyla as opposed to the

odd-toed ungulate Perissodactyla (, ), or odd-toed ungulates, is an order of ungulates. The order includes about 17 living species divided into three families: Equidae (horses, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They t ...

s of Perissidactyla, such as the

horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...

, which have one toe, or the

rhinoceros A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant taxon, extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family (biology), famil ...

, which has three toes. The five-toed ancestors of the earliest

had already developed feet that suggest odd-toed and even-toed descendants to the modern viewer. Even Phenacodus, the most generalized of the early mammals, has a foot in which the central toe is somewhat larger than the others and could be placed in the division of odd-toed ungulates, Perissidactyla.

Jewish dietary laws

The distinction between cloven and uncloven hooves is highly relevant for dietary laws of

Judaism Judaism () is an Abrahamic religions, Abrahamic, Monotheism, monotheistic, ethnic religion that comprises the collective spiritual, cultural, and legal traditions of the Jews, Jewish people. Religious Jews regard Judaism as their means of o ...

(''

kashrut (also or , ) is a set of Food and drink prohibitions, dietary laws dealing with the foods that Jewish people are permitted to eat and how those foods must be prepared according to halakha, Jewish law. Food that may be consumed is deemed ko ...

''), as set forth in the

Torah The Torah ( , "Instruction", "Teaching" or "Law") is the compilation of the first five books of the Hebrew Bible, namely the books of Genesis, Exodus, Leviticus, Numbers and Deuteronomy. The Torah is also known as the Pentateuch () ...

and the

Talmud The Talmud (; ) is the central text of Rabbinic Judaism and the primary source of Jewish religious law (''halakha'') and Jewish theology. Until the advent of Haskalah#Effects, modernity, in nearly all Jewish communities, the Talmud was the cen ...

.Shulchan Aruch, Yoreh De'ah 79 . Animals that ''both'' chew their cud ( ruminate, i.e. regurgitate partly digested food from a specialised multi-chambered stomach back to the mouth to be chewed for a second time as part of their ordinary digestive process) and have split true cloven hooves (a hoof being hard or rubbery sole and a hard wall formed by a thick nail) are allowed (''

kosher (also or , ) is a set of dietary laws dealing with the foods that Jewish people are permitted to eat and how those foods must be prepared according to Jewish law. Food that may be consumed is deemed kosher ( in English, ), from the Ashke ...

'', lit. "fit") for Jewish consumption. Those animals that have neither of these two characteristics, or only one of the characteristics, are considered

unclean animals In some religions, an unclean animal is an animal whose consumption or handling is taboo. According to these religions, persons who handle such animals may need to Ritual purification, ritually purify themselves to get rid of their uncleanline ...

('' treif'', not fit for Jewish consumption) and Jews are forbidden to eat them. This rule thus excludes the

camel A camel (from and () from Ancient Semitic: ''gāmāl'') is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as livestock, they provid ...

from the list of kosher animals because although the camel does ruminate it does not possess true "hooves" – it walks on soft toes which have little more than a nail merely giving an appearance of a "hoof". Similarly the pig, although it has cloven true hooves, does not ruminate. Other animals such as horses, dogs and cats exhibit neither characteristic and are thus also forbidden for Jewish consumption.

References

External links

Food laws of Judaism concerning cloven hooves in the ''Torah ''and in Maimonides' Code of Jewish Law (''Mishneh Torah'')
{{DEFAULTSORT:Cloven Hoof Artiodactyls Mammal anatomy