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Choristodere
Choristodera (from the Greek χωριστός ''chōristos'' + δέρη ''dérē'', 'separated neck') is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the late Miocene (168 to 11.6 million years ago). Choristoderes are morphologically diverse, with the best known members being the crocodile-like neochoristoderes such as ''Champsosaurus''. Other choristoderans had lizard-like or long necked morphologies. Choristoderes appear to have been confined to the Northern Hemisphere, having been found in North America, Asia, and Europe, and possibly also North Africa. Choristoderes are generally thought to be derived neodiapsids that are close relatives or members of Sauria. History of Discovery Choristodera was erected in 1876, originally as a suborder of Rhynchocephalia by Edward Drinker Cope to contain ''Champsosaurus,'' which was described from Late Cretaceous strata of Montana by Cope in the same paper. A year l ...
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Neochoristodera
Neochoristodera is a lineage of specialised crocodile-like fully aquatic choristodere reptiles. Noted for their long jaws and large size, these animals were predominant across the Northern Hemisphere, occurring in freshwater and coastal environments across the Cretaceous and early Cenozoic. Systematics Neochoristoderes form a monophyletic group, however there is no consensus about the relationships of the genera, which have been recovered as a polytomy in recent studies. Neochoristodera contains the named genera ''Champsosaurus'', '' Ikechosaurus'', '' Kosmodraco'', '' Liaoxisaurus'', '' Mengshanosaurus'', ''Simoedosaurus'' and '' Tchoiria''. Various taxa of uncertain affinities within this group are known, including a partial femur of a choristodere, possibly of a neochoristodere from the Cedar Mountain Formation of the United States and an indeterminate partial skeleton from the Kuwajima Formation of Japan. Evolution Neochoristoderes first appear in the Early Cretaceous of A ...
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Cteniogenys
''Cteniogenys'' is a genus of choristodere, a morphologically diverse group of aquatic reptiles. It is part of the monotypic family Cteniogenidae. The type and only named species, ''C. antiquus'', was named in 1928 by Charles W. Gilmore. The holotype, VP.001088, was collected in the Morrison Formation (Como Bluff), Wyoming in 1881 by William H. Reed. More specimens have been discovered since then, including specimens from the Late Jurassic of Portugal and Middle Jurassic of Britain, which have not been assigned to species. Description ''Cteniogenys'' was long, and probably weighed less than . The skull of this genus was long and slender, and the jaws had numerous conical teeth. ''Cteniogenys'' in the Morrison probably fed on insects and small fish. It is mostly known from freshwater sites (rivers and ponds), and is a rare find in the formation (only 60 specimens out of over 2,800 total vertebrate specimens known from the formation), mostly known from northern outc ...
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Simoedosaurus
''Simoedosaurus'' is an extinct reptile known from the Paleocene of North America, Europe and western Asia, and a member of the Choristodera, a group of aquatic reptiles that lived in the Northern Hemisphere from the Jurassic to the early Cenozoic. Described other species, ''S. dakotensis'' later got its own genus, '' Kosmodraco''. Taxonomy French paleontologist Paul Gervais described ''Simoedosaurus'' in 1877. Though similar to and contemporaneous, ''Simoedosaurus'' is not closely related to the North American ''Champsosaurus'', instead it appears to be most closely related to '' Tchoiria'' and '' Ikechosaurus'' from the Early Cretaceous of Asia. It therefore may represent a species that immigrated into North America from Asia in the wake of the Cretaceous-Tertiary mass extinction event, though the absence of choristoderes in the Late Cretaceous of Asia makes this merely a paleogeographical speculation. Biology ''Simoedosaurus'' was an aquatic predator, specialised to a full ...
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Kosmodraco
''Kosmodraco'' is a genus of large bodied choristodere from the Paleocene of North America. Originally described as a species of ''Simoedosaurus'', it was found to represent a distinct genus in 2022. Multiple fossil skulls show a relatively short and robust snout and a skull that is considerably wider behind the eyes. Two species are currently recognized, ''K. dakotensis'' and ''K. magnicornis''. History and naming The first specimen now known to belong to ''Kosmodraco'' was discovered in 1964 and 1968 in the Polecat Bench Formation (Wyoming). These two skulls, alongside others from Montana's Bear Creek, were reported on briefly by Sigogneau-Russell and Donald in 1978, who regarded them as evidence for the presence of the Eurasian genus ''Simoedosaurus'' in North America. However, at the time, these four specimens, although thought to be diagonstic at a genus level, were still unprepared and not assigned to a species. They were subsequently stored in the collection of the Princ ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the la ...
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Lazarussuchus
''Lazarussuchus'' (meaning " Lazarus's crocodile") is an extinct genus of amphibious reptile, known from the Cenozoic of Europe. It is the youngest known member of Choristodera, an extinct order of aquatic reptiles that first appeared in the Middle Jurassic. Fossils have been found in Late Paleocene, Late Oligocene, Early Miocene and Late Miocene deposits (~56-11.6 million years ago) in France, Germany, and the Czech Republic. Two species have been named: the type species ''L. inexpectatus'' ("unexpected") (Hecht, 1992) from the late Oligocene of France. and ''L. dvoraki'' from the early Miocene of the Czech Republic. It was not a large animal; the skull of ''L. inexpectatus'' was only about 4.53 centimeters long (1.78 in), with the total preserved body and tail length being just over 30 centimetres. A complete specimen of ''Lazarussuchus'' with preserved soft tissue was found from the Late Paleocene of France, but has not been assigned to a species. Discovery The firs ...
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Shokawa
''Shokawa'' is an extinct genus of choristoderan diapsid reptile, known from the Lower Cretaceous of Japan. It is only known from one species, ''Shokawa ikoi.'' The only known remains are a postcranial specimen lacking the skull, discovered at the KO2 locality in sediments belonging to the Okurodani Formation near the village of Shokawa in Gifu Prefecture. ''Shokawa'' possessed a long neck with at least 16 cervical vertebrae, and closely resembles and is closely related to the smaller choristoderan, ''Hyphalosaurus ''Hyphalosaurus'' (meaning "submerged lizard") is a genus of freshwater aquatic reptiles, belonging to the extinct order Choristodera. They lived during the early Cretaceous period (Aptian age), about 122 million years ago. The genus contains ...''.The dentary of a Choristodere (Reptilia: Archosauromorpha) from the Okurodani Formation, Tetori Group (Lower Cretaceous) of Japa/ref> The generic name refers to the village near where it was found, while the specifi ...
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Pachystropheus
''Pachystropheus'' (After Greek ''Pachys'', "Thick" and ''Strophaios'', Vertebrae) is a genus of prehistoric reptile, possibly a choristodere (champsosaur), from the Rhaetian (Late Triassic) of southwestern England. It was named by Erika von Huene in 1935; Huene described ''Pachystropheus'' as a champsosaur, but this was overlooked for decades until its redescription by Storrs and Gower in 1993. This reevaluation would extend the fossil record of champsosaurs back 45 million years. However, other authors consider attribution of ''Pachystropheus'' to Choristodera problematic, stating that it depends on vertebral and girdle characters that are also found in the skeletons of aquatic reptiles other than choristoderes; most of the diagnostic features of choristoderes are skull features, but the presence of these cannot be confirmed in ''Pachystropheus'', as there is no confirmed skull material for this taxon. Silvio Renesto (2005) found similarities in the postcranial skeleton ...
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Champsosaurus
''Champsosaurus'' is an extinct genus of crocodile-like choristodere reptile, known from the Late Cretaceous and early Paleogene periods of North America and Europe (Campanian-Paleocene). The name ''Champsosaurus'' is thought to come from , () said in an Ancient Greek source to be an Egyptian word for "crocodiles", and , () Greek for "lizard". The morphology of ''Champsosaurus'' resembles that of gharials, with a long, elongated snout. It was native to freshwater environments where it likely preyed on fish, similar to living gharials. History of research ''Champsosaurus'' was the first member of the Choristodera to be described. ''Champsosaurus'' was named by Edward Drinker Cope in 1876, from isolated vertebrae found in Late Cretaceous strata of the Judith River Formation on the banks of the Judith River in Fergus County, Montana. Cope designated ''C. annectens'' as the type species rather than the first named ''C. profundus'' due to the larger number of vertebrae he attributed ...
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Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic period (), and the Classical period (). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Epic and Classical periods of the language. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regarded as a separate historical stage, although its earliest form closely resembles Attic Greek and its latest form approaches Medieval Greek. There were several regional dialects of Ancient Greek, of which Attic Greek developed into Koi ...
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