Wadi Milk Formation
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Wadi Milk Formation
The Wadi Milk Formation is a geological formation in Sudan whose strata date back to the Late Cretaceous. Originally, the formation was thought to be Albian to Cenomanian, later research has provided dating to the Campanian to Maastrichtian. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, Africa)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 604-605. . It stretches from the lower Wadi Al-Malik across the Wadi Muqaddam into the Bayuda Desert.Oliver W.M.Rauhut, A Dinosaur Fauna from the Late Cretaceous (Cenomanian) of Northern Sudan. In: Palaeontologie africaine,35,61-84(1999) Vertebrate paleofauna Ornithischians Saurischians See also * List of dinosaur-bearing rock formations This list of dinosaur-bearing rock formations is a list of geologic formations in which din ...
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Geological Formation
A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exposed in a geographical region (the stratigraphic column). It is the fundamental unit of lithostratigraphy, the study of strata or rock layers. A formation must be large enough that it can be mapped at the surface or traced in the subsurface. Formations are otherwise not defined by the thickness of their rock strata, which can vary widely. They are usually, but not universally, tabular in form. They may consist of a single lithology (rock type), or of alternating beds of two or more lithologies, or even a heterogeneous mixture of lithologies, so long as this distinguishes them from adjacent bodies of rock. The concept of a geologic formation goes back to the beginnings of modern scientific geology. The term was used by Abraham Gottlob Wer ...
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Iguanodontia
Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', ''Tenontosaurus'', and the hadrosaurids or "duck-billed dinosaurs". Iguanodontians were one of the first groups of dinosaurs to be found. They are among the best known of the dinosaurs, and were among the most diverse and widespread herbivorous dinosaur groups of the Cretaceous period. Classification Iguanodontia is often listed as an infraorder within a suborder Ornithopoda, though Benton (2004) lists Ornithopoda as an infraorder and does not rank Iguanodontia. Traditionally, iguanodontians were grouped into the superfamily Iguanodontoidea and family Iguanodontidae. However, phylogenetic studies show that the traditional "iguanodontids" are a paraphyletic grade leading up to the hadrosaurs (duck-billed dinosaurs). Groups like Iguanodontoidea are sometimes still used as unranked clades ...
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Carcharodontosaurus
''Carcharodontosaurus'' (; ) is a genus of large carcharodontosaurid theropod dinosaur that existed during the Cenomanian age of the Late Cretaceous in Northern Africa. The genus ''Carcharodontosaurus'' is named after the shark genus ''Carcharodon'', itself composed of the Greek (, meaning "jagged" or "sharp") and (, "teeth"), and the suffix ' ("lizard"). It is currently known to have two species: ''C. saharicus'' and ''C. iguidensis''. History of discovery In 1924, two teeth were found in the Continental intercalaire of Algeria, showing what were at the time unique characteristics. These teeth were described by Depéret and Savornin (1925) as representing a new taxon, which they named ''Megalosaurus saharicus'' and later categorized in the subgenus ''Dryptosaurus''. Some years later, paleontologist Ernst Stromer described the remains of a partial skull and skeleton from Cenomanian aged rocks in the Bahariya Formation of Egypt (Stromer, 1931); originally excavated in 1914, ...
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Carcharodontosauridae
Carcharodontosauridae (carcharodontosaurids; from the Greek καρχαροδοντόσαυρος, ''carcharodontósauros'': "shark-toothed lizards") is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae as a family, which, in modern paleontology, indicates a clade within Carnosauria. Carcharodontosaurids include some of the largest land predators ever known: ''Giganotosaurus'', ''Mapusaurus'', ''Carcharodontosaurus'', and ''Tyrannotitan'' all rivaled or exceeded ''Tyrannosaurus'' in size. A 2015 paper by Christophe Hendrickx and colleagues gives a maximum length estimate of for the largest carcharodontosaurids, while the smallest carcharodontosaurids were estimated to have been at least long. Evolution Along with the spinosaurids, carcharodontosaurids were the largest predators in the early and middle Cretaceous throughout Gondwana, with species also present in North America (''Acrocanthosaurus''), Europe (''Concavenator'') and Asia ...
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Bahariasaurus Ingens, Like Megaraptora
''Bahariasaurus'' (meaning "Bahariya lizard") is an enigmatic genus of large theropod dinosaur. ''Bahariasaurus'' is known to have included at least 1 species, ''Bahariasaurus ingens'', which was found in North African rock layers dating to the Cenomanian and Turonian ages of the Late Cretaceous. The only fossils confidently assigned to ''Bahariasaurus'' were found in the Bahariya Formation of the Bahariya (Arabic: الواحة البحرية meaning the "northern oasis") oasis in Egypt by Ernst Stromer but were destroyed during a World War II bombing raid with the same raid taking out the holotype of ''Spinosaurus'' and '' Aegyptosaurus'' among other animals found in the Bahariya Formation. While there have been more fossils assigned to the genus such as some from the Farak Formation of Niger, these remains are referred to with much less certainty. ''Bahariasaurus'' is, by most estimations, one of the largest theropods, approaching the size of other large bodied theropods such ...
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Bahariasaurus
''Bahariasaurus'' (meaning "Bahariya lizard") is an enigmatic genus of large theropod dinosaur. ''Bahariasaurus'' is known to have included at least 1 species, ''Bahariasaurus ingens'', which was found in North African rock layers dating to the Cenomanian and Turonian ages of the Late Cretaceous. The only fossils confidently assigned to ''Bahariasaurus'' were found in the Bahariya Formation of the Bahariya (Arabic: الواحة البحرية meaning the "northern oasis") oasis in Egypt by Ernst Stromer but were destroyed during a World War II bombing raid with the same raid taking out the holotype of ''Spinosaurus'' and ''Aegyptosaurus'' among other animals found in the Bahariya Formation. While there have been more fossils assigned to the genus such as some from the Farak Formation of Niger, these remains are referred to with much less certainty. ''Bahariasaurus'' is, by most estimations, one of the largest theropods, approaching the size of other large bodied theropods such as ...
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Theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago ( Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of diets was historically considered a characteri ...
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Titanosauridae
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Dicraeosaurids BW
Dicraeosauridae is a Family (biology), family of Diplodocoidea, diplodocoid sauropods who are the sister group to Diplodocidae. Dicraeosaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as ''Amargasaurus'', ''Suuwassea'', ''Dicraeosaurus'', and ''Brachytrachelopan''. Specimens of this family have been found in North America, Asia, Africa, and South America. Their temporal range is from the Early Jurassic, Early or Middle Jurassic to the Early Cretaceous. Few dicraeosaurids survived into the Cretaceous, the youngest of which was ''Amargasaurus''. The group was first described by German paleontologist Werner Janensch in 1914 with the discovery of ''Dicraeosaurus'' in Tanzania. Dicraeosauridae are distinct from other sauropods because of their relatively short neck size and small body size. The clade is monophyletic and well-supported phylogenetically with thirteen unambiguous synapomorphies uniting it. They diverged from Diplod ...
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Dicraeosauridae
Dicraeosauridae is a family of diplodocoid sauropods who are the sister group to Diplodocidae. Dicraeosaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as ''Amargasaurus'', ''Suuwassea'', ''Dicraeosaurus'', and ''Brachytrachelopan''. Specimens of this family have been found in North America, Asia, Africa, and South America. Their temporal range is from the Early or Middle Jurassic to the Early Cretaceous. Few dicraeosaurids survived into the Cretaceous, the youngest of which was ''Amargasaurus''. The group was first described by German paleontologist Werner Janensch in 1914 with the discovery of ''Dicraeosaurus'' in Tanzania. Dicraeosauridae are distinct from other sauropods because of their relatively short neck size and small body size. The clade is monophyletic and well-supported phylogenetically with thirteen unambiguous synapomorphies uniting it. They diverged from Diplodocidae in the Mid-Jurassic, as evidenced by the ...
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Sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include ''Brachiosaurus'', ''Diplodocus'', ''Apatosaurus'' and ''Brontosaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. ''Isanosaurus'' and ''Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the group in the Late Triassic. By the Late Jurassic (150 million yea ...
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