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Tortonian Genera
The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Tortonian roughly overlaps with the regional Pannonian Stage of the Paratethys timescale of Central Europe. It also overlaps the upper Astaracian, Vallesian and lower Turolian European land mammal ages, the upper Clarendonian and lower Hemphillian North American land mammal ages and the upper Chasicoan and lower Huayquerian South American land mammal ages. Definition The Tortonian was introduced by Swiss stratigrapher Karl Mayer-Eymar in 1858. It was named after the Italian city of Tortona in the Piedmont region. The base of the Tortonian Stage is at the last common appearance of calcareous nanoplankton '' Discoaster kugleri'' and planktonic foram '' Globigerinoides subquadratus''. It is also associated with the short normal polarized magnetic ...
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Tertiary Period
The Tertiary ( ) is an obsolete Period (geology), geologic period spanning 66 million to 2.6 or 1.8 million years ago. The period began with the extinction of the non-bird, avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start of the Cenozoic, Cenozoic Era, and extended to the beginning of the Quaternary glaciation at the end of the Pliocene, Pliocene Epoch. The Tertiary has not been recognised by the International Commission on Stratigraphy (ICS) since the late 1980s, with the timespan of the Tertiary now being split in to the earlier Paleogene and the more recent Neogene periods, though the Tertiary continues to be used in some scientific publications. Historical use of the term The term Tertiary was first used by Giovanni Arduino (geologist), Giovanni Arduino during the mid-18th century. He classified geologic time into primitive (or primary), secondary, and tertiary periods based on observations of geology in Northern Italy. Later a fourth period, t ...
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Late Miocene
The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma. The evolution of ''Homo'' The gibbons (family Hylobatidae) and orangutans (genus ''Pongo'') were the first groups to split from the line leading to the hominins, including humans, then gorillas (genus ''Gorilla''), and finally chimpanzees and bonobos (genus ''Pan (genus), Pan''). The splitting date between hominin and chimpanzee lineages is placed by some between 4 and 8 million years ago, that is, during the Late Miocene. References External links GeoWhen Database - Late Miocene
Miocene, .03 Miocene geochronology, 03 Messinian, * Tortonian, * {{geochronology-stub ...
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Huayquerian
The Huayquerian () age is a period of geologic time (9.0–6.8 Ma) within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification. It follows the Chasicoan and precedes the Montehermosan age. Etymology The age is named after the Huayquerías Formation in the western Cuyo Basin of northwestern Argentina that was later dated to the Montehermosan The Montehermosan age is a period of geologic time (6.8–4.0 Mya (unit), Ma) within the Miocene and Pliocene epochs of the Neogene used more specifically with South American Land Mammal Ages. It follows the Huayquerian and precedes the Chapadmalal .... The most complete Huayquerian fauna is found in the Cerro Azul Formation, in Buenos Aires Province also referred to as Epecuén Formation. Formations Fossil content Correlations Notes and references Notes References Bibliography ;Huayquerías Formation * * ;Andalhuala Formation * ;Camacho Formation * * * ;Cerro A ...
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Chasicoan
The Chasicoan ( or ) age is a period of geologic time from 10–9 Ma within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Mayoan and precedes the Huayquerian age.Chasicoan
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Etymology

The Chasicoan is named after the
Arroyo Chasicó Formation Arroyo often refers to: * Arroyo (watercourse), an intermittently dry creek Arroyo may also refer to: Places United States * Arroyo, Pennsylvania * Arroyo, Puerto Rico, a muni ...
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North American Land Mammal Age
The North American land mammal ages (NALMA) establishes a geologic timescale for North American fauna beginning during the Late Cretaceous and continuing through to the present. These periods are referred to as ages or intervals (or stages when referring to the rock strata of that age) and were established using geographic place names where fossil materials were obtained. System The North American land-mammal-age system was formalized in 1941 as a series of provincial land-mammal ages. The system was the standard for correlations in the terrestrial Cenozoic record of North America and was the source for similar time scales dealing with other continents. The system was revised into a formal chronostratigraphic system. This approach is nominally justified by international stratigraphic codes; it holds that first appearances of individual species in particular sections are the only valid basis for naming and defining the land-mammal ages. The basic unit of measure is the first/last ...
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Hemphillian
The Hemphillian North American Stage on the geologic timescale is a North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 10,300,000 to 4,900,000 years BP. It is usually considered to overlap the Tortonian age of the Late Miocene and Zanclean age of the Early Pliocene. The Hemphillian is preceded by the Clarendonian and followed by the Blancan The Blancan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 4,750,000 to 1,806,000 years BP, a period of .
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Clarendonian
The Clarendonian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 13,600,000 to 9,000,000 years BP, a period of . It is usually considered to overlap the Serravallian age of the Middle Miocene and the Tortonian age of the Late Miocene. The Clarendonian is preceded by the Barstovian and followed by the Hemphillian The Hemphillian North American Stage on the geologic timescale is a North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 10,300,000 to 4,900,000 years BP. It is usually considered t ... NALMA stages. Subdivisions The Clarendonian can be further divided into the substages of: * Late/Upper Clarendonian: Lower boundary source of the base of the Clarendonian (approximate) * Early/Lower Clarendonian (shares lower boundary) References Miocene life Miocene animals of North ...
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European Land Mammal Age
The European Land Mammal Mega Zones (abbreviation: ELMMZ, more commonly known as European land mammal ages or ELMA) are zones in rock layers that have a specific assemblage of fossils ( biozones) based on occurrences of fossil assemblages of European land mammals. These biozones cover most of the Cenozoic, with particular focus having been paid to the Neogene and Paleogene systems (i.e. rock layers which are 65.5 to 2.588 million years old), the Quaternary has several competing systems. In cases when fossils of mammals are abundant, stratigraphers and paleontologists can use these biozones as a more practical regional alternative to the stages of the official ICS geologic timescale. European Land Mammal Mega Zones are often also confusingly referred to as ages, stages, or intervals. Biostratigraphic methods Mammal zones were, like all biozones, established using geographic place names where fossil materials were obtained. The basic unit of measure is the first/last boundary sta ...
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Turolian
The Turolian age is a period of geologic time (9.0–5.3 Ma) within the Miocene used more specifically with European Land Mammal Ages. It precedes the Ruscinian age and follows the Vallesian age. The Turolian overlaps the Tortonian and Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fir ... ages. ;References Miocene {{geochronology-stub ...
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Vallesian
The Vallesian age is a period of geologic time (11.6–9.0 Ma) within the Miocene used more specifically with European Land Mammal Ages. It precedes the Turolian age and follows the Astaracian age. The so-called Vallesian Crisis resulted in the extinction of several mammalian taxa characteristic of the Middle Miocene. The term "Vallesian" was introduced by Catalan palaeontologist Miquel Crusafont in 1950 to mark the arrival of the equid '' Hipparion'' in Europe. The remaining European palaeofaunas, however, had been around since the Middle Miocene, including the moschid '' Micromeryx'' (a musk deer), the cervid '' Euprox'', the suid '' Listriodon'', and the felids '' Sansanosmilus'' and '' Pseudaelurus'', and the Aragonian-Vallesian"Aragonian" is a Spanish term for a continental stage, roughly equivalent to the Middle Miocene or Astaracian (16–11 Ma). boundary does not represent a major shift in the European mammalian record. In contrast, the transition between Lower and U ...
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Astaracian
The Astaracian age is a period of geologic time (), equivalent with the Middle Miocene and used more specifically with European Land Mammal Ages. It precedes the Vallesian age and follows the Orleanian age. The Astaracian overlaps the Langhian and Serravallian ages. During the Late Orleanian and Astaracian (), oscillating sea levels resulted in a succession of palaeogeographic changes in the Eastern Mediterranean; the opening and closing of the Tethys seaway resulted in temporary land-bridges between Africa and Eurasia. Three short periods of faunal migrations between the continents can be distinguished: During the Late Orleanian, MN 5 (), a first wave of migrations from Africa correspond to fossil fauna from Greece. The Antonios locality (MN 4/5, ) on the Chalkidiki peninsula includes the small tragulid '' Dorcatherium'', the giraffid '' Palaeomeryx'', and the suiform '' Sanitheres''. The Thymiana locality (MN 5, ) on Chios includes both the giraffid '' Georgiomeryx' ...
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Paratethys
The Paratethys sea, Paratethys ocean, Paratethys realm or just Paratethys (meaning "beside Tethys"), was a large shallow inland sea that covered much of mainland Europe and parts of western Asia during the middle to late Cenozoic, from the late Paleogene to the late Neogene, and is regarded as the largest inland sea in history. At its greatest extent, it stretched from the region north of the Alps over Central Europe to the Aral Sea in Central Asia. Paratethys formed about 34 Mya (million years ago) at the beginning of the Oligocene epoch, when the northern region of the Tethys Ocean (Peri-Tethys) was separated from the Mediterranean region of the Tethys realm due to the formation of the Alps, Carpathians, Dinarides, Taurus and Elburz mountains. Paratethys was at times reconnected with the Tethys or its successors (the Mediterranean Sea or the Indian Ocean) during the Oligocene and the early and middle Miocene times, but at the onset of the late Miocene epoch, the tecton ...
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