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Tetrapodomorph
Tetrapodomorpha (also known as Choanata) is a clade of vertebrates consisting of tetrapods (four-limbed vertebrates) and their closest sarcopterygian relatives that are more closely related to living tetrapods than to living lungfish. Advanced forms transitional between fish and the early labyrinthodonts, such as '' Tiktaalik'', have been referred to as "fishapods" by their discoverers, being half-fish, half-tetrapods, in appearance and limb morphology. The Tetrapodomorpha contains the crown group tetrapods (the last common ancestor of living tetrapods and all of its descendants) and several groups of early stem tetrapods, which includes several groups of related lobe-finned fishes, collectively known as the osteolepiforms. The Tetrapodomorpha minus the crown group Tetrapoda are the stem Tetrapoda, a paraphyletic unit encompassing the fish to tetrapod transition. Characteristics Among the characteristics defining tetrapodomorphs are modifications to the fins, notably a hum ...
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Tetrapodomorpha
Tetrapodomorpha (also known as Choanata) is a clade of vertebrates consisting of tetrapods (four-limbed vertebrates) and their closest sarcopterygian relatives that are more closely related to living tetrapods than to living lungfish. Advanced forms transitional between fish and the early labyrinthodonts, such as '' Tiktaalik'', have been referred to as "fishapods" by their discoverers, being half-fish, half-tetrapods, in appearance and limb morphology. The Tetrapodomorpha contains the crown group tetrapods (the last common ancestor of living tetrapods and all of its descendants) and several groups of early stem tetrapods, which includes several groups of related lobe-finned fishes, collectively known as the osteolepiforms. The Tetrapodomorpha minus the crown group Tetrapoda are the stem Tetrapoda, a paraphyletic unit encompassing the fish to tetrapod transition. Characteristics Among the characteristics defining tetrapodomorphs are modifications to the fins, notably a ...
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Lobe-finned Fish
Sarcopterygii (; )—sometimes considered synonymous with Crossopterygii ()—is a clade (traditionally a class or subclass) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish. These vertebrates are characterised by prominent muscular limb buds (lobes) within their fins, which are supported by articulated appendicular skeletons. This is in contrast to the other clade of bony fish, the Actinopterygii, which have only skin-covered bony spines supporting the fins. The tetrapods, a mostly terrestrial clade of vertebrates, are now recognized as having evolved from sarcopterygian ancestors and are most closely related to lungfishes. Their paired pectoral and pelvic fins evolved into limbs, and their foregut diverticulum eventually evolved into air-breathing lungs. Cladistically, this would make the tetrapods a subgroup within Sarcopterygii and thus sarcopterygians themselves. As a result, the phrase "lobe-finned fish" normally refe ...
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Tetrapod
A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetrapoda (). Tetrapods include all Neontology#Extant taxa versus extinct taxa, extant and Extinction, extinct amphibians and amniotes, with the latter in turn Evolution, evolving into two major clades, the Sauropsida, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (extinct pelycosaur, "pelycosaurs", therapsids and all extant mammals, including Homo sapiens, humans). Hox gene mutations have resulted in some tetrapods becoming Limbless vertebrate, limbless (snakes, legless lizards, and caecilians) or two-limbed (cetaceans, sirenians, Bipedidae, some lizards, kiwi (bird), kiwis, and the extinct moa and elephant birds). Nevertheless, they still qualify as tetrapods through their ancestry, and some retain a pair of ves ...
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Kenichthys
''Kenichthys'' is a genus of sarcopterygian fish from the Devonian period, and a member of the clade Tetrapodomorpha. The only known species of the genus is ''Kenichthys campbelli'' (named for the Australian palaeontologist Ken Campbell), the first remains of which were found in China in 1993.Chang, M. and Zhu, M. (1993) A new Middle Devonian osteolepidid from Qujing, Yunnan. ''Mem. Assoc. Australas. Palaeontol.'' 15 183-198 The genus is important to the study of the evolution of tetrapods due to the unique nature of its nostrils, which provide vital evidence regarding the evolutionary transition of fish-like nostrils to the tetrapod choanae.Zhu, M. and Ahlberg, P. (2004) The origin of the internal nostril of tetrapods. ''Nature'' 432 94-97 Description ''Kenichthys'' was a small tetrapodomorph, with a skull about long. While only areas of the front of the body are known, it seems likely that ''Kenichthys'' would have been similar in general body form to other basal sarcopte ...
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Stem Tetrapod
The Stem Tetrapoda are a cladistically defined group, consisting of all animals more closely related to extant four-legged vertebrates than to their closest extant relatives (the lungfish), but excluding the crown group Tetrapoda. They are thus paraphyletic, though acceptable in phylogenetic nomenclature as the group is defined by strict reference to phylogeny rather than to traits as in traditional systematics. Thus, some finned sarcopterygians are considered to be stem tetrapods. Content of the group Stem tetrapods are members of Tetrapodomorpha, the total group and clade that also includes their descendants, the crown tetrapods: The stem Tetrapoda encompass three distinct grades successively closer to crown group Tetrapoda: * Osteolepiformes, a group of lobe-finned fishes that includes the families Tristichopteridae, Canowindridae, Megalichthyidae, and Osteolepidae * Elpistostegalia, the more advanced lobe-finned fishes (Tristichopteridae) and the " fishapods" (gener ...
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Osteolepidida
Osteolepiformes, also known as Osteolepidida, is a group of prehistoric lobe-finned fishes which first appeared during the Devonian period. The order contains the families Canowindridae, Megalichthyidae, Osteolepididae and Tristichopteridae, in addition to several monotypic families. The order is generally considered to be paraphyletic because the characters that define it are mainly attributes of stem tetrapodomorphs. The following taxonomy is based on Borgen & Nakrem, 2016: * Order Osteolepiformes ** Suborder Osteolepidoidei *** Family Osteolepididae *** Family Thursiidae *** Family Megalichthyidae ** Suborder Cyclolepidoidei *** Superfamily Eopodoidea **** Family Chrysolepididae **** Family Gyroptychiidae **** Family Panderichthyidae (incl. Elpistostegalia) **** Family Tristichopteridae *** Superfamily Parapodoidea **** Family Canowindridae **** Family Medoevididae *** Superfamily Rhizodontoidea Below is a cladogram A cladogram (from Greek language, Gre ...
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Rhizodontida
Rhizodontida is an extinct group of predatory tetrapodomorphs known from many areas of the world from the Givetian through to the Pennsylvanian - the earliest known species is about 377 million years ago (Mya), the latest around 310 Mya. Rhizodonts lived in tropical rivers and freshwater lakes and were the dominant predators of their age. They reached huge sizes - the largest known species, '' Rhizodus hibberti'' from Europe and North America, was an estimated 7 m in length, making it the largest freshwater fish known. Description The upper jaw had a marginal row of small teeth on the maxilla and premaxilla, medium-sized fangs on the ectopterygoid and dermopalatine bones, and large tusks on the vomers and premaxillae. On the lower jaw were marginal teeth on the dentary, with fangs on the three coronoids and a huge tusk at the symphysial tip of the dentary. Apparently, the left and right mandibles rotated inwards towards each other on biting. This may have been a kinetic me ...
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Labyrinthodont
"Labyrinthodontia" (Greek, 'maze-toothed') is an informal grouping of extinct predatory amphibians which were major components of ecosystems in the late Paleozoic and early Mesozoic eras (about 390 to 150 million years ago). Traditionally considered a subclass of the class Amphibia, modern classification systems recognize that labyrinthodonts are not a formal natural group (clade) exclusive of other tetrapods. Instead, they consistute an evolutionary grade (a paraphyletic group), ancestral to living tetrapods such as lissamphibians (modern amphibians) and amniotes (reptiles, mammals, and kin). "Labyrinthodont"-grade vertebrates evolved from lobe-finned fishes in the Devonian, though a formal boundary between fish and amphibian is difficult to define at this point in time. "Labyrinthodont" generally refers to extinct four-limbed tetrapods with a large body size and a crocodile-like lifestyle. The name describes the pattern of infolding of the dentin and enamel of the teeth, whi ...
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Osteolepiform
Osteolepiformes, also known as Osteolepidida, is a group of prehistoric lobe-finned fishes which first appeared during the Devonian period. The order contains the families Canowindridae, Megalichthyidae, Osteolepididae and Tristichopteridae, in addition to several monotypic families. The order is generally considered to be paraphyletic because the characters that define it are mainly attributes of stem tetrapodomorphs. The following taxonomy is based on Borgen & Nakrem, 2016: * Order Osteolepiformes ** Suborder Osteolepidoidei *** Family Osteolepididae *** Family Thursiidae *** Family Megalichthyidae ** Suborder Cyclolepidoidei *** Superfamily Eopodoidea **** Family Chrysolepididae **** Family Gyroptychiidae **** Family Panderichthyidae (incl. Elpistostegalia) **** Family Tristichopteridae *** Superfamily Parapodoidea **** Family Canowindridae **** Family Medoevididae *** Superfamily Rhizodontoidea Below is a cladogram A cladogram (from Greek language, Gre ...
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Tristichopteridae
Tristichopterids (Tristichopteridae) were a diverse and successful group of fish-like tetrapodomorphs living throughout the Middle and Late Devonian. They first appeared in the Eifelian stage of the Middle Devonian. Within the group sizes ranged from a few tens of centimeters (''Tristichopterus'') to several meters (''Hyneria'' and ''Eusthenodon''). Some tristichopterids share some of the features of the elpistostegalians, a diverse clade of tetrapodomorphs close to the origin of (and including) tetrapods. This mainly concerns the shape of the skull and a reduction in size of the posterior fins. An old and persistent notion is that ''Eusthenopteron'' was able to crawl onto land using its fins. However, there is no evidence actually supporting this idea.M. Laurin, F. J. Meunier, D. Germain, and M. Lemoine 2007A microanatomical and histological study of the paired fin skeleton of the Devonian sarcopterygian ''Eusthenopteron foordi'' ''Journal of Paleontology'' 81: 143–153. All t ...
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