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Shaochilong
''Shaochilong'' (meaning "shark toothed dragon") is an extinct genus of carcharodontosaurid dinosaur from the mid-Cretaceous (Late Cenomanian to the end of the Turonian stage) Ulansuhai Formation of China (about 92 million years ago). The type species, ''S. maortuensis'', was originally named '' Chilantaisaurus maortuensis'', but was re-described and reclassified in 2009. It was one of the last known carcharodontosaurids to walk the earth. Alongside '' Ulughbegsaurus'' from Uzbekistan and ''Mapusaurus'' from Argentina, they were the only members of the family to live until the end of the Turonian epoch. History The material referred to ''Shaochilong'', IVPP V.2885.1-7, consisted of skull fragments (a braincase, partial skull roof, quadrates, and a right maxilla), axis and six caudal vertebrae. A fragmentary left maxilla was also referred to the species, although it has apparently gone missing as of 2009. Although these are believed to belong to a single individual, a lectotype ...
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Shaochilong Scale
''Shaochilong'' (meaning "shark toothed dragon") is an extinct genus of carcharodontosaurid dinosaur from the mid-Cretaceous (Late Cenomanian to the end of the Turonian stage) Ulansuhai Formation of China (about 92 million years ago). The type species, ''S. maortuensis'', was originally named ''Chilantaisaurus maortuensis'', but was re-described and reclassified in 2009. It was one of the last known carcharodontosaurids to walk the earth. Alongside '' Ulughbegsaurus'' from Uzbekistan and ''Mapusaurus'' from Argentina, they were the only members of the family to live until the end of the Turonian epoch. History The material referred to ''Shaochilong'', IVPP V.2885.1-7, consisted of skull fragments (a braincase, partial skull roof, quadrates, and a right maxilla), axis and six caudal vertebrae. A fragmentary left maxilla was also referred to the species, although it has apparently gone missing as of 2009. Although these are believed to belong to a single individual, a lectotype ...
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Carcharodontosauridae
Carcharodontosauridae (carcharodontosaurids; from the Greek καρχαροδοντόσαυρος, ''carcharodontósauros'': "shark-toothed lizards") is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae as a family, which, in modern paleontology, indicates a clade within Carnosauria. Carcharodontosaurids include some of the largest land predators ever known: ''Giganotosaurus'', ''Mapusaurus'', ''Carcharodontosaurus'', and ''Tyrannotitan'' all rivaled or exceeded ''Tyrannosaurus'' in size. A 2015 paper by Christophe Hendrickx and colleagues gives a maximum length estimate of for the largest carcharodontosaurids, while the smallest carcharodontosaurids were estimated to have been at least long. Evolution Along with the spinosaurids, carcharodontosaurids were the largest predators in the early and middle Cretaceous throughout Gondwana, with species also present in North America (''Acrocanthosaurus''), Europe (''Concavenator'') and Asia ...
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Chilantaisaurus
''Chilantaisaurus'' (" lizard") is a genus of large theropod dinosaur, possibly a neovenatorid or a primitive coelurosaur, from the Late Cretaceous Ulansuhai Formation of China (Turonian age, about 92 million years ago). The type species, ''C. tashuikouensis'', was described by Hu in 1964. Description ''Chilantaisaurus'' was a large theropod, estimated as weighing between and . In 2010, Brusatte et al. estimated it to weigh , based on femur length measurements. However, considering that greater femoral circumference indicates the greater capacity to withstand greater locomotor loads (not greater body mass), the 2020 study moderated the body mass of the holotype at . It is estimated to be around to long. Classification Hu considered ''Chilantaisaurus'' to be a carnosaur related to ''Allosaurus'', though some subsequent studies suggested that it may be a spinosauroid, possibly a primitive member of the spinosaurid family (Sereno, 1998; Chure, 2000; Rauhut, 2001) because i ...
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Ulansuhai Formation
The Ulansuhai Formation () is a geological formation in Inner Mongolia, north China. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Cretaceous, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 563-570. . The Ulansuhai Formation has traditionally been considered to date to the Aptian-Albian stages of the Lower Cretaceous, due to similarities between the Ulansuhai fauna and known Aptian formations. However, radiometric dating done on underlying formations has shown this to be incorrect. Due to the age of underlying rocks, the Ulansuhai Formation cannot be older than the Turonian stage of the Late Cretaceous, about 92 Ma.Kobayashi, Y., and Lu, J.-C. (2003). "A new ornithomimid dinosaurian with gregarious habits from the Late Cretaceous of China." ''Acta Palaeontol. Pol.'', 48: 235–259.Be ...
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Mapusaurus
''Mapusaurus'' () was a giant carcharodontosaurid carnosaurian dinosaur from the early Late Cretaceous (early Turonian stage), approximately 93.9 to 89.6 million years ago, of what is now Argentina. Discovery ''Mapusaurus'' was excavated between 1997 and 2001, by the Argentinian-Canadian Dinosaur Project, from an exposure of the Huincul Formation (Rio Limay Subgroup, Cenomanian) at Cañadón del Gato. It was described and named by paleontologists Rodolfo Coria and Phil Currie in 2006. The name ''Mapusaurus'' is derived from the Mapuche word ''Mapu'', meaning 'of the Land' or 'of the Earth' and the Greek ''sauros'', meaning 'lizard'. The type species, ''Mapusaurus roseae'', is named for both the rose-colored rocks, in which the fossils were found and for Rose Letwin, who sponsored the expeditions which recovered these fossils. The designated holotype for the genus and type species, ''Mapusaurus roseae'', is an isolated right nasal (MCF-PVPH-108.1, Museo Carmen Funes, Pale ...
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Ulughbegsaurus
''Ulughbegsaurus'' (meaning "Ulugh Beg's lizard") is a dubious genus of theropod dinosaur from the Late Cretaceous aged Bissekty Formation, Uzbekistan. The type species is ''Ulughbegsaurus uzbekistanensis''. Discovery and naming ''Ulughbegsaurus'' was initially discovered within the Bissekty Formation, Uzbekistan in the 1980s based on the holotype UzSGM 11-01-02, consisting of a partial left maxilla. The holotype, despite its significance, would remain in the collection of the State Geological Museum of the State Committee of the Republic of Uzbekistan on Geology and Mineral Resources, Tashkent, Uzbekistan, until 2019, when it was rediscovered. Two additional referred specimens have been attributed to the genus, including CCMGE 600/12457, an jugal ramus of a left maxilla that was previously referred to the dromaeosaurid ''Itemirus'' and ZIN PH 357/16, the posterior end of a right maxilla. Isolated teeth from the Bissekty Formation show similarities with carcharodontosaurian mor ...
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Braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skullcap. The remainder of the skull is the facial skeleton. In comparative anatomy, neurocranium is sometimes used synonymously with endocranium or chondrocranium. Structure The neurocranium is divided into two portions: * the membranous part, consisting of flat bones, which surround the brain; and * the cartilaginous part, or chondrocranium, which forms bones of the base of the skull. In humans, the neurocranium is usually considered to include the following eight bones: * 1 ethmoid bone * 1 frontal bone * 1 occipital bone * 2 parietal bones * 1 sphenoid bone * 2 temporal bones The ossicles (three on each side) are usually not included as bones of the neurocranium. There may variably also be extra sutural bones present. Below the ne ...
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Femur
The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with the tibia (shinbone) and patella (kneecap), forming the knee joint. By most measures the two (left and right) femurs are the strongest bones of the body, and in humans, the largest and thickest. Structure The femur is the only bone in the upper leg. The two femurs converge medially toward the knees, where they articulate with the proximal ends of the tibiae. The angle of convergence of the femora is a major factor in determining the femoral-tibial angle. Human females have thicker pelvic bones, causing their femora to converge more than in males. In the condition ''genu valgum'' (knock knee) the femurs converge so much that the knees touch one another. The opposite extreme is ''genu varum'' (bow-leggedness). In the general popu ...
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Carnosauria
Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as '' Giganotosaurus'', '' Mapusaurus'', '' Carcharodontosaurus'' and '' Tyrannotitan'' which are among the largest known predatory dinosaurs. While it originally contained a wide assortment of giant theropods that were not closely related, the group has since been defined to encompass only the allosaurs and their closest kin. However, with the description and publication in 2019 of ''Asfaltovenator vialidadi'', a basal allosauroid curiously displaying both primitive and derived features seen in Tetanurae, the new phylogenetic analysis has found Megalosauroidea to be a basal grade of carnosaurs in respect to Allosauroidea; thus significantly expanding Carnosauria's inclusiveness towards its original context. Distinctive characteristics of ca ...
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Coelurosaur
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrannosaurs, ornithomimosaurs, and maniraptorans; Maniraptora includes birds, the only known dinosaur group alive today. Most feathered dinosaurs discovered so far have been coelurosaurs. Philip J. Currie had considered it likely and probable that all coelurosaurs were feathered. However, several skin impressions found for some members of this group show pebbly, scaly skin, indicating that feathers did not completely replace scales in all taxa. In the past, Coelurosauria was used to refer to all small theropods, but this classification has since been abolished. Anatomy Bodyplan The studying of anatomical traits in coelurosaurs indicates that the last common ancestor had evolved the ability to eat and digest plant matter, adapting to an omni ...
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Nomen Nudum
In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate description. This makes it a "bare" or "naked" name, which cannot be accepted as it stands. A largely equivalent but much less frequently used term is ''nomen tantum'' ("name only"). In zoology According to the rules of zoological nomenclature a ''nomen nudum'' is unavailable; the glossary of the ''International Code of Zoological Nomenclature'' gives this definition: And among the rules of that same Zoological Code: In botany According to the rules of botanical nomenclature a ''nomen nudum'' is not validly published. The glossary of the ''International Code of Nomenclature for algae, fungi, and plants'' gives this definition: The requirements for the diagnosis or description are covered by articles 32, 36, 41, 42, and 44. ''Nomina nu ...
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Lectotype
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms, a set that includes some organisms and excludes others, based on a detailed published description (for example a species description) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. Type specimen According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and the International Code of Nomenclature for algae, fungi, and plants (ICN), the scientific name of every taxon is almost al ...
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