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Symphyta
Sawflies are wasp-like insects that are in the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay their eggs. The name is associated especially with the Tenthredinoidea, by far the largest Taxonomic rank#Ranks in zoology, superfamily in the suborder, with about 7,000 known species; in the entire suborder, there are 8,000 described species in more than 800 genera. Symphyta is paraphyly, paraphyletic, consisting of several Basal (phylogenetics), basal groups within the order Hymenoptera, each one rooted inside the previous group, ending with the Apocrita which are not sawflies. The primary distinction between sawflies and the Apocrita – the ants, bees, and wasps – is that the adults lack a "wasp waist", and instead have a broad connection between the abdomen and the Thorax (insect anatomy), thorax. Some sawflies are Ba ...
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Xyeloidea
The Xyelidae are a comparatively species-poor family of sawflies, comprising about 80 Extant taxon, extant species in five genera worldwide, and is the only family in the superfamily Xyeloidea. The fossil record of the family is extensive, comprising more than 120 species and including the oldest fossil Hymenoptera species dating back to the Triassic, between 245 and 208 million years ago. Xyelidae are the most basal contemporary lineage of Hymenoptera and have many ancestral morphological features. The extant species occur in the Northern Hemisphere, especially in boreal ecosystem, boreal regions of the Holarctic, though there are a few Oriental species. Two genera and about 15 species occur in Europe.Taeger, A., Blank, S.M. & Liston, A. D. 2006: European Sawflies (Hymenoptera: Symphyta) – A Species Checklist for the Countries. Pp. 399-504. In: Blank, S.M., Schmidt, S. & Taeger, A. (eds): Recent Sawfly Research: Synthesis and Prospects. Goecke & Evers, Keltern.Blank, S.M. 2002: ...
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Hymenoptera
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they reach adulthood. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek πτερόν (''pteron'') for wing. The Ancient Greek ὑμήν (''hymen'') for membrane provides a plausible etymology for the term because species in this order have membranous wings. However, a key characteristic of this order is that the hi ...
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Paraphyly
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancest ...
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Wasp
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey. The most commonly known wasps, such as yellowjackets and hornets, are in the family Vespidae and are eusocial, living together in a nest with an egg-laying queen and non-reproducing workers. Eusociality is favoured by the unusual haplodiploid system of sex determination in Hymenoptera, as it makes sisters exceptionally closely related to each other. However, the majority of wasp species are solitary, with each adult female living and breeding independently. Females typically have an oviposit ...
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Apocrita
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" ( petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma (or gaster) rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host (plant or animal) or in a nest cell prov ...
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Crochet (insect Anatomy)
A proleg is a small, fleshy, stub structure found on the ventral surface of the abdomen of most larval forms of insects of the order Lepidoptera, though they can also be found on larvae of insects such as sawflies. In all the orders in which they appear, mainly Hymenoptera and Lepidoptera, prolegs of any form evolved independently of each other by convergent evolution. They appear to have evolved from ancient inactive genes that have since been activated again. Prolegs of lepidopteran larvae have a small circle of gripping hooks, called "crochets". The arrangement of the crochets can be helpful in identification to family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ... level. Although the point has been debated, prolegs are not widely regarded as true legs, derived from the p ...
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Proleg
A proleg is a small, fleshy, stub structure found on the ventral surface of the abdomen of most larval forms of insects of the Order (biology), order Lepidoptera, though they can also be found on larvae of insects such as symphyta, sawflies. In all the orders in which they appear, mainly Hymenoptera and Lepidoptera, prolegs of any form evolved independently of each other by convergent evolution. They appear to have evolved from ancient inactive genes that have since been activated again. Prolegs of lepidopteran larvae have a small circle of gripping hooks, called "crochets". The arrangement of the crochets can be helpful in identification to Rank (zoology), family level. Although the point has been debated, prolegs are not widely regarded as true arthropod leg, legs, derived from the primitive Arthropod leg#Biramous and uniramous, uniramous limbs. Certainly in their morphology they are not jointed, and so lack the five segments (coxa, trochanter, femur, tibia, tarsus) of thoraci ...
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Tenthredinoidea
The Tenthredinoidea are the dominant superfamily of sawflies within the Symphyta, containing some 8,400 species worldwide, primarily in the family Tenthredinidae. All known larvae are phytophagous, and a number are considered pests. The included extant families share the distinctive features of a medially narrowed pronotum, paired protibial spurs, and the loss of the transverse mesonotal groove. The superfamily also includes two extinct families. Meicai and Haiyan (1998) identified 66 extant tribes and 17 subfamilies. Taxonomy Families * Argidae Konow, 1890 (58 genera, 897 spp.) * Blasticotomidae Thomson, 1871 (3 genera, 13 spp.) * Cimbicidae W. Kirby, 1837 (16 genera, 182 spp.) * Diprionidae Rohwer, 1910 (11 genera, 136 spp.) * Pergidae Rohwer, 1911 (60 genera, 442 spp.) *Tenthredinidae Tenthredinidae is the largest family of sawflies, with well over 7,500 species worldwide, divided into 430 genera. Larvae are herbivores and typically feed on the foliage of tree ...
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Pamphilioidea
The Pamphilioidea are a small superfamily within the Symphyta (the sawflies), containing some 250 living species restricted to the temperate regions of Eurasia and North America. These hymenopterans share the distinctive feature of a very large, almost prognathous head, which is widest ventrally. The superfamily contains two extant families. The Pamphiliidae are the leaf-rolling or web-spinning sawflies such as '' Acantholyda'', '' Neurotoma'', and '' Pamphilius'' whose larvae eat plants such as conifers; the adults have simple filiform antennae. The Megalodontesidae include genera such as '' Megalodontes'' and several fossil groups. Their larvae eat herbaceous plants, while the adults have serrate or pectinate antennae. References Bibliography * , in Zhang, Z.-Q. (ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013) * Rasnitsyn, Alexandr P.; Zhang, Haichun & Wang, Bo (2006): Bizarre fossil insects: web-spinn ...
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Cephoidea
Cephoidea is a small superfamily within the Symphyta, commonly referred to as stem sawflies, containing some 100 species in 10 genera in the living family, Cephidae, plus another 17 genera in the extinct family Sepulcidae. They first appeared around 212 million years ago in the Norian Age, and are diurnal. Most species occur in the Northern Hemisphere, especially in Eurasia. The larvae are stem borers in various plants, especially grasses, but sometimes other herbaceous plants, shrubs, or trees. A few are pests of cereal grains (e.g. '' Cephus cinctus'', which attacks wheat). They are exceptionally slender for symphytans, often resembling other types of wasps, and they are the only Symphyta which lack cenchri. They are sometimes postulated to be the sister taxon to the Apocrita Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Sy ...
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Xiphydrioidea
Xiphydriidae are a family of wood wasps that includes around 150 species. They are located all over the world including North and South America, Australia, Europe, and others. Xiphydriidae larvae are wood borers in dead trees or branches of a range of trees. They are characterized as having long and skinny necks with dome-shaped heads. The oldest fossils of the group are from the mid Cretaceous. Genera These 29 genera belong to the family Xiphydriidae: * '' Alloxiphia'' Wei, 2002 * '' Austrocyrta'' Riek, 1955 * '' Austroxiphyda'' Jennings, Macdonald, Schiff & Parslow, 2021 * '' Brachyxiphus'' Philippi, 1871 * '' Calexiphyda'' Smith, 2008 * '' Carinoxiphia'' Wei, 1999 * '' Derecyrta'' Smith, 1860 * '' Eoxiphia'' Maa, 1949 * '' Euxiphydria'' Semenov-Tian-Shanskii & Gussakovskii, 1935 * '' Genaxiphia'' Maa, 1949 * '' Gryponeura'' Benson, 1954 * '' Heteroxiphia'' Saini & Singh, 1987 * '' Hyperxiphia'' Maa, 1949 * '' Indoxiphia'' Maa, 1949 * '' Lataxiphyda'' Smith, 2008 * '' Lissoxiph ...
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