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Self-fertilisation
Autogamy or self-fertilization refers to the fusion of two gametes that come from one individual. Autogamy is predominantly observed in the form of self-pollination, a reproductive mechanism employed by many flowering plants. However, species of protists have also been observed using autogamy as a means of reproduction. Flowering plants engage in autogamy regularly, while the protists that engage in autogamy only do so in stressful environments. Occurrence Protists ''Paramecium aurelia'' '' Paramecium aurelia'' is the most commonly studied protozoan for autogamy. Similar to other unicellular organisms, ''Paramecium aurelia'' typically reproduce asexually via binary fission or sexually via cross-fertilization. However, studies have shown that when put under nutritional stress, ''Paramecium aurelia'' will undergo meiosis and subsequent fusion of gametic-like nuclei. This process, defined as hemixis, a chromosomal rearrangement process, takes place in a number of steps. First, the ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta. Angiosperms are by far the most diverse group of Embryophyte, land plants with 64 Order (biology), orders, 416 Family (biology), families, approximately 13,000 known Genus, genera and 300,000 known species. They include all forbs (flowering plants without a woody Plant stem, stem), grasses and grass-like plants, a vast majority of broad-leaved trees, shrubs and vines, and most aquatic plants. Angiosperms are distinguished from the other major seed plant clade, the gymnosperms, by having flowers, xylem consisting of vessel elements instead of tracheids, endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from the commo ...
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Cell Fusion
Cell fusion is an important cellular process in which several uninucleate cells (cells with a single nucleus) combine to form a multinucleate cell, known as a syncytium. Cell fusion occurs during differentiation of myoblasts, osteoclasts and trophoblasts, during embryogenesis, and morphogenesis. Cell fusion is a necessary event in the maturation of cells so that they maintain their specific functions throughout growth. History In 1839, Theodor Schwann, in his '' Microscopical Researches,'' expanded upon the theory that all living organisms are composed of cells when he added that discrete cells are the basis of life. Schwann observed that in certain cells the walls and cavities of the cells coalesce ('' verschmelzen'') together. This observation provided the first hint that cells fuse. It was not until 1960 that cell biologists deliberately fused cells for the first time. To fuse the cells, biologists combined isolated mouse cells and induced fusion of their outer membrane ...
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Ciliate
The ciliates are a group of alveolates characterized by the presence of hair-like organelles called cilia, which are identical in structure to flagellum, eukaryotic flagella, but are in general shorter and present in much larger numbers, with a different wikt:undulating, undulating pattern than flagella. Cilia occur in all members of the group (although the peculiar Suctoria only have them for part of their biological life cycle, life cycle) and are variously used in swimming, crawling, attachment, feeding, and sensation. Ciliates are an important group of protists, common almost anywhere there is water—in lakes, ponds, oceans, rivers, and soils, including anoxic and oxygen-depleted habitats. About 4,500 unique free-living species have been described, and the potential number of extant species is estimated at 27,000–40,000. Included in this number are many Ectosymbiosis, ectosymbiotic and endosymbiotic species, as well as some Obligate parasite, obligate and Facultative paras ...
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Self-pollination
Self-pollination is a form of pollination in which pollen arrives at the stigma of a flower (in flowering plants) or at the ovule (in gymnosperms) of the same plant. The term cross-pollination is used for the opposite case, where pollen from one plant moves to a different plant. There are two types of self-pollination: in autogamy, pollen is transferred to the stigma of the same flower; in geitonogamy, pollen is transferred from the anther of one flower to the stigma of another flower on the same flowering plant, or from microsporangium to ovule within a single ( monoecious) gymnosperm. Some plants have mechanisms that ensure autogamy, such as flowers that do not open ( cleistogamy), or stamens that move to come into contact with the stigma. The term selfing that is often used as a synonym is not limited to self-pollination, but also applies to other types of self-fertilization. Occurrence Few plants self-pollinate without the aid of pollen vectors (such as wind or ...
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Geitonogamy
Geitonogamy (from Greek ''geiton'' (γείτων) = neighbor + ''gamein'' (γαμεῖν) = to marry) is a type of self-pollination. Geitonogamous pollination is sometimes distinguished from the fertilizations that can result from it, geitonogamy. If a plant is self-incompatible, geitonogamy can reduce seed production. Geitonogamy is when pollen is exported using a vector (pollinator or wind) out of one flower but only to another flower on the same plant. It is a form of self-fertilization. In flowering plants, pollen is transferred from a flower to another flower on the same plant, and in animal pollinated systems this is accomplished by a pollinator visiting multiple flowers on the same plant. Geitonogamy is also possible within species that are wind-pollinated, and may actually be a quite common source of self-fertilized seeds in self-compatible species. It also occurs in monoecious gymnosperms. Although geitonogamy is functionally cross-pollination involving a pollinating ag ...
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Gynoecium
Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists of (one or more) ''#Pistil, pistils'' and is typically surrounded by the pollen-producing plant reproductive morphology, reproductive organs, the stamens, collectively called the androecium. The gynoecium is often referred to as the "female" portion of the flower, although rather than directly producing female gametes (i.e. egg cells), the gynoecium produces megaspores, each of which develops into a female gametophyte which then produces egg cells. The term gynoecium is also used by botanists to refer to a cluster of archegonia and any associated modified leaves or stems present on a gametophyte shoot in mosses, Marchantiophyta, liverworts, and hornworts. The corresponding terms for the male parts of those plants are clusters of antheridiu ...
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Stamen
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament and an anther which contains sporangium, microsporangia. Most commonly, anthers are two-lobed (each lobe is termed a locule) and are attached to the filament either at the base or in the middle area of the anther. The sterile (i.e. nonreproductive) tissue between the lobes is called the Connective (botany), connective, an extension of the filament containing conducting strands. It can be seen as an extension on the dorsal side of the anther. A pollen grain develops from a microspore in the microsporangium and contains the male gametophyte. The size of anthers differs greatly, from a tiny fraction of a millimeter in ''Wolfia'' spp up to five inches (13 centimeters) in ''Canna iridiflora'' and ''Strelitzia nicolai''. The stamens in a flower ...
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Pollen
Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male Conifer cone, cone to the female cone of gymnosperms. If pollen lands on a compatible pistil or female cone, it Germination, germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require magnification to see detail. The study of pollen is called palynology and is highly useful in paleoecology, paleontology, archaeology, and Forensic science, forensics. Pollen in plants is used for transferring Ploidy#Haploid and monoploid, haploid male genetic ma ...
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Sperm
Sperm (: sperm or sperms) is the male reproductive Cell (biology), cell, or gamete, in anisogamous forms of sexual reproduction (forms in which there is a larger, female reproductive cell and a smaller, male one). Animals produce motile sperm with a tail known as a flagellum, which are known as spermatozoa, while some red algae and fungi produce non-motile sperm cells, known as spermatia. Flowering plants contain non-motile sperm inside pollen, while some more basal plants like ferns and some gymnosperms have motile sperm. Sperm cells form during the process known as spermatogenesis, which in amniotes (reptiles and mammals) takes place in the seminiferous tubules of the testicles. This process involves the production of several successive sperm cell precursors, starting with spermatogonia, which Cellular differentiation, differentiate into spermatocytes. The spermatocytes then undergo meiosis, reducing their Ploidy, chromosome number by half, which produces spermatids. The sper ...
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Ploidy
Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, respectively, in each homologous chromosome pair—the form in which chromosomes naturally exist. Somatic cells, tissues, and individual organisms can be described according to the number of sets of chromosomes present (the "ploidy level"): monoploid (1 set), diploid (2 sets), triploid (3 sets), tetraploid (4 sets), pentaploid (5 sets), hexaploid (6 sets), heptaploid or septaploid (7 sets), etc. The generic term polyploid is often used to describe cells with three or more sets of chromosomes. Virtually all sexually reproducing organisms are made up of somatic cells that are diploid or greater, but ploidy level may vary widely between different organisms, between different tissues within the same organism, and at different stages in an o ...
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Macronucleus
A macronucleus (formerly also meganucleus) is the larger type of nucleus in ciliates. Macronuclei are polyploid and undergo direct division without mitosis. It controls the non-reproductive cell functions, such as metabolism Metabolism (, from ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cellular processes; the co .... During conjugation, the macronucleus disintegrates, and a new one is formed by karyogamy of the micronuclei. Macronuclei contain hundreds to thousands of chromosomes, each present in many copies. There is no mechanism to precisely partition this complex genome equally during nuclear division; thus, how the cell manages to maintain a balanced genome after generations of divisions is unknown. See also * Micronucleus References . Cell nucleus Ciliate biology {{Ciliate-stub ...
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Nuclear Dimorphism
Nuclear dimorphism is a term referred to the special characteristic of having two different kinds of nuclei in a cell. There are many differences between the types of nuclei. This feature is observed in protozoan ciliates, like ''Tetrahymena'', and some foraminifera. Ciliates contain two nucleus types: a macronucleus that is primarily used to control metabolism, and a micronucleus which performs reproductive functions and generates the macronucleus. The compositions of the nuclear pore complexes help determine the properties of the macronucleus and micronucleus. Nuclear dimorphism is subject to complex epigenetic controls. Nuclear dimorphism is continuously being studied to understand exactly how the mechanism works and how it is beneficial to cells. Learning about nuclear dimorphism is beneficial to understanding old eukaryotic mechanisms that have been preserved within these unicellular organisms but did not evolve into multicellular eukaryotes. Key components The ciliated ...
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