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Procolophonia
The Procolophonia are a suborder of herbivorous reptiles that lived from the Middle Permian till the end of the Triassic period. They were originally included as a suborder of the Cotylosauria (later renamed Captorhinida Carroll 1988) but are now considered a clade of Parareptilia. They are closely related to other generally lizard-like Permian reptiles such as the Millerettidae, Bolosauridae, Acleistorhinidae, and Lanthanosuchidae, all of which are included under the Anapsida or "Parareptiles" (as opposed to the Eureptilia). Classification There are two main groups of Procolophonia, the small, lizard-like Procolophonoidea, and the Pareiasauroidea, which include the large, armoured Pareiasauridae. According to the traditional classification of Carroll 1988 as well as recent phylogenetic analyses, smaller groups like Rhipaeosauridae (now a synonym of Nycteroleteridae) and Sclerosauridae are classified with the pareiasaurs and with the procolophonids, respectively. The ...
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Nyctiphruretidae
Nyctiphruretidae is an extinct family (biology), family of hallucicranian parareptiles known from the late Early Permian, Early to the late Middle Permian of European Russia and south-central United States. Nyctiphruretidae was named by Efremov in 1938. The type genus is ''Nyctiphruretus'' known from several Middle Permian locations in European Russia. Michael S. Y. Lee in 1997 placed the family in the larger clade Nyctiphruretia. The order was defined to include both nyctiphruretids and nycteroleterids. However, recent cladistic analyses suggest that nycteroleterids are more closely related to the Pareiasauria instead, thus making ''Nyctiphruretus'' the only genus of its order. In 2014, MacDougall & Reisz described and named a second genus of Nyctiphruretidae, ''Abyssomedon'', from the middle Leonardian stage of the late Early Permian of Comanche County, Oklahoma, south-central United States. It contains a single species, ''A. williamsi'', which represents the first nyctiphrureti ...
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Pareiasauromorpha
Pareiasauromorpha is a group of parareptilian amniotes from the Permian. It includes genera found all over the world, with many genera from Asia and South Africa. The clade was first used as a group by Linda A. Tsuji in 2011, in order to group the family Nycteroleteridae (nycteroleters) and the superfamily Pareiasauroidea (pareiasaurs). Pareiasauromorpha is considered to be a monophyletic node, the sister group to procolophonoids. Classification ''Pareiasauromorpha'' was first used to define a group of parareptilians in 2011 by Linda A. Tsuji. The next year, Tsuji and her colleagues used Pareiasauromorpha as a node inside Procolophonia. In their 2012 publication, Tsuji ''et al.'' defined it as a monophyletic node containing "nycteroleters" (the family Nycteroleteridae) and "pareiasaurs" (in the superfamily Pareiasauroidea). Nycteroleteridae Nycteroleteridae is a family, commonly called "nycteroleters", classified in Pareiasauromorpha. The group includes the genera '' ...
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Parareptilia
Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles (bolosaurids such as '' Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by t ...
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Procolophonoidea
Procolophonoidea is an extinct superfamily of procolophonian parareptiles. Members were characteristically small, stocky, and lizard-like in appearance. Fossils have been found worldwide from many continents including Antarctica. The first members appeared during the Late Permian in the Karoo Basin of South Africa. Taxonomy Procolophonoidea includes the families Owenettidae and Procolophonidae. ''Sclerosaurus'', which is placed within its own family Sclerosauridae, may be a member of the superfamily as well. In 1997, De Braga and Rieppel defined this same taxon (the oldest common ancestor of Procolophonidae and Owenettidae and all its descendants) using the name Procolophoniformes.Jalil, N. E., & Janvier, P. (2005). Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc. Geodiversitas, 27(1), 35-132. When the superfamily was constructed in 1956, it was thought to be within the anthracosaur suborder Diadectomorpha. Since then it has been pla ...
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Reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchocephalians ( tuatara). As of March 2022, the Reptile Database includes about 11,700 species. In the traditional Linnaean classification system, birds are considered a separate class to reptiles. However, crocodilians are more closely related to birds than they are to other living reptiles, and so modern cladistic classification systems include birds within Reptilia, redefining the term as a clade. Other cladistic definitions abandon the term reptile altogether in favor of the clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals. The study of the traditional reptile orders, historically combined with that of modern amphibians, is called herpetology. The earliest known proto-reptiles originated ...
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Procolophonoid
Procolophonoidea is an extinct superfamily of procolophonian parareptiles. Members were characteristically small, stocky, and lizard-like in appearance. Fossils have been found worldwide from many continents including Antarctica. The first members appeared during the Late Permian in the Karoo Basin of South Africa. Taxonomy Procolophonoidea includes the families Owenettidae and Procolophonidae. ''Sclerosaurus'', which is placed within its own family Sclerosauridae, may be a member of the superfamily as well. In 1997, De Braga and Rieppel defined this same taxon (the oldest common ancestor of Procolophonidae and Owenettidae and all its descendants) using the name Procolophoniformes.Jalil, N. E., & Janvier, P. (2005). Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc. Geodiversitas, 27(1), 35-132. When the superfamily was constructed in 1956, it was thought to be within the anthracosaur suborder Diadectomorpha. Since then it has been plac ...
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Acleistorhinidae
Acleistorhinidae is an extinct family of Late Carboniferous and Early Permian-aged ( Moscovian to Kungurian stage) parareptiles. Acleistorhinids are most diverse from the Richards Spur locality of the Early Permian of Oklahoma. Richards Spur acleistorhinids include '' Acleistorhinus'', '' Colobomycter'', and possibly '' Delorhynchus'' and '' Feeserpeton''. Other taxa include ''Carbonodraco'' from the Late Carboniferous of Ohio and '' Karutia'' from the Early Permian of Brazil. Acleistorhinidae is commonly considered a subgroup of lanthanosuchoids, related to taxa such as '' Chalcosaurus'', '' Lanthaniscus'' and '' Lanthanosuchus''. However, a re-examination of parareptile phylogeny conducted by Cisneros ''et al.'' (2021) argued that lanthanosuchids were not closely related to acleistorhinids. The phylogenetic analysis conducted by these authors recovered acleistorhinids as the sister group of the clade Procolophonia, while lanthanosuchids were recovered within the procolophon ...
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Owenettidae
Owenettidae is an extinct family of procolophonian parareptiles. Fossils have been found primarily from Africa and Madagascar, with one genus present from South America. It is the sister taxon to the family Procolophonidae. The family was constructed in 1939 for the type genus ''Owenetta''. Since then several other genera have been assigned to Owenettidae, including '' Barasaurus'' and '' Saurodektes''. The oldest known owenettid, ''Owenetta rubidgei'', dates back to the Wuchiapingian stage of the Late Permian. It and ''Saurodektes'', have been found from the Beaufort Group in the Karoo Basin of South Africa, while '' Barasaurus'' is known from the Late Permian and the Early Triassic of Sakamena Group in Madagascar. These strata span the Permo-Triassic boundary and provide evidence for the great faunal change that occurred as a result of the Permian–Triassic extinction event. Although most owenettids lived during the Permian, remains of some taxa such as ''"Owenetta" kitching ...
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Order (biology)
Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may fol ...
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Captorhinida
Fossil of '' Labidosaurus hamatus'' Captorhinida (older name: Cotylosauria) is a doubly paraphyletic grouping of early reptiles. Robert L. Carroll (1988) ranked it as an order in the subclass Anapsida, composed of the following suborders:R. L. Carroll (1988), '' Vertebrate Paleontology and Evolution'', W. H. Freeman and Company, New York * A paraphyletic Captorhinomorpha, containing the families Protorothyrididae, Captorhinidae, Bolosauridae, Acleistorhinidae and possibly also Batropetidae * Procolophonia, containing families Nyctiphruretidae, Procolophonidae and Sclerosauridae * Pareiasauroidea, with families Rhipaeosauridae and Pareiasauridae * Millerosauroidea, with a single family Millerettidae. While they all share primitive features and resemble the ancestors of all modern reptiles, some of these families are more closely related to (or belong to) the clade Parareptilia, while others are further along the line leading to diapsid Diapsids ("two arches") are a cla ...
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Cotylosauria
Fossil of '' Labidosaurus hamatus'' Captorhinida (older name: Cotylosauria) is a doubly paraphyletic grouping of early reptiles. Robert L. Carroll (1988) ranked it as an order in the subclass Anapsida, composed of the following suborders:R. L. Carroll (1988), '' Vertebrate Paleontology and Evolution'', W. H. Freeman and Company, New York * A paraphyletic Captorhinomorpha, containing the families Protorothyrididae, Captorhinidae, Bolosauridae, Acleistorhinidae and possibly also Batropetidae * Procolophonia, containing families Nyctiphruretidae, Procolophonidae and Sclerosauridae * Pareiasauroidea, with families Rhipaeosauridae and Pareiasauridae * Millerosauroidea, with a single family Millerettidae. While they all share primitive features and resemble the ancestors of all modern reptiles, some of these families are more closely related to (or belong to) the clade Parareptilia, while others are further along the line leading to diapsid Diapsids ("two arches") are a cla ...
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Phylogenetic Analyses
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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