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Procolophonines
Procolophoninae is an Extinction, extinct subfamily of Procolophonidae, procolophonid Parareptilia, parareptiles from the late Early Triassic to the early Middle Triassic (Olenekian and Anisian stages) of Africa, Antarctica, Asia, Europe and South America. Currently, the oldest-known procolophonine is ''Procolophon'' from the earliest Olenekian stage. Phylogeny Procolophoninae was named in 1890 by Richard Lydekker. It is a Phylogenetic nomenclature, stem-based taxon defined Phylogenetics, phylogenetically for the first time by Modesto ''et al.'' (2002) as "all taxa more related to ''Procolophon trigoniceps'' Owen, 1876 than to ''Leptopleuron lacertinum'' Owen, 1851". The cladogram below follows Ruta ''et al.'' 2011. Below are two cladograms that follow phylogenetic analyses by Butler ''et al''. (2023): Analyses 1 and 3: Strict consensus of 760 and 18 most parsimonious trees (MPTs). Analysis 2: Single MPT. References

Procolophoninae Early Triassic reptiles Midd ...
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Timanophon
''Timanophon'' is an extinct genus of procolophonine procolophonid parareptile from early Triassic deposits of Arkhangelsk, Russia. It is known from the holotype PIN 3359/11, a partial skeleton including nearly complete skull and lower jaw which was previously referred to ''Burtensia'' sp. by Ivakhnenko in 1975. It was collected in the Mezenskaya Pizhma and Lower Syamzhen'ga localities from the Pizhmomezenskoi Formation. Ten additional specimens from the same localities are PIN 3359/1-3, 3359/63-65 and 4364/35-38. The fragmentary dentary, dentaries PIN 3360/1-3 were collected in the Vybor River locality, from the same formation. All specimens came from the Ustmylian Gorizont, dating to the early Olenekian faunal stage of the Early Triassic, about 249-247 million years ago. It was first named by I. V. Novikov in 1991 in paleontology, 1991 and the type species is ''Timanophon raridentatus''. References

Procolophoninae Early Triassic reptiles of Asia Fos ...
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Thelephon
''Thelephon'' is an extinct genus of procolophonine procolophonid parareptile from middle Triassic (early Anisian stage) deposits of Free State Province, South Africa. It is known from the holotype BP/1/3512, a partial skull lacking the snout and anterior third of the mandible. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Winnaarsbaken and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group (Karoo Basin). It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is ''Thelephon contritus''. It was first assigned to a species of '' Thelegnathus'' (now considered to be a ''nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...''), ''Thelegnathus ...
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Richard Lydekker
Richard Lydekker (; 25 July 1849 – 16 April 1915) was a British naturalist, geologist and writer of numerous books on natural history. He was known for his contributions to zoology, paleontology, and biogeography. He worked extensively in cataloging fossil vertebrates and describing new species, particularly from India, where he spent several years studying the region’s prehistoric fauna. Lydekker was a key figure in the field of vertebrate paleontology, authoring numerous scientific papers and books that helped classify extinct and extant species. Biography Richard Lydekker was born at Tavistock Square in London. His father was Gerard Wolfe Lydekker, a barrister-at-law with Dutch ancestry. The family moved to Harpenden Lodge soon after Richard's birth. He was educated at Trinity College, Cambridge, where he took a first-class in the Natural Science tripos (1872). In 1874 he joined the Geological Survey of India and made studies of the vertebrate palaeontology of norther ...
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South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion of the Americas. South America is bordered on the west by the Pacific Ocean, on the north and east by the Atlantic Ocean, and to the south by the Drake Passage; North America and the Caribbean Sea lie to the northwest. The continent includes twelve sovereign states: Argentina, Bolivia, Brazil, Chile, Colombia, Ecuador, Guyana, Paraguay, Peru, Suriname, Uruguay, and Venezuela; two dependent territory, dependent territories: the Falkland Islands and South Georgia and the South Sandwich Islands; and one administrative division, internal territory: French Guiana. The Dutch Caribbean ABC islands (Leeward Antilles), ABC islands (Aruba, Bonaire, and Curaçao) and Trinidad and Tobago are geologically located on the South-American continental shel ...
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Europe
Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east. Europe shares the landmass of Eurasia with Asia, and of Afro-Eurasia with both Africa and Asia. Europe is commonly considered to be Boundaries between the continents#Asia and Europe, separated from Asia by the Drainage divide, watershed of the Ural Mountains, the Ural (river), Ural River, the Caspian Sea, the Greater Caucasus, the Black Sea, and the waterway of the Bosporus, Bosporus Strait. "Europe" (pp. 68–69); "Asia" (pp. 90–91): "A commonly accepted division between Asia and Europe ... is formed by the Ural Mountains, Ural River, Caspian Sea, Caucasus Mountains, and the Black Sea with its outlets, the Bosporus and Dardanelles." Europe covers approx. , or 2% of Earth#Surface, Earth's surface (6.8% of Earth's land area), making it ...
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Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which has long been home to the majority of the human population, was the site of many of the first civilisations. Its 4.7 billion people constitute roughly 60% of the world's population. Asia shares the landmass of Eurasia with Europe, and of Afro-Eurasia with both Europe and Africa. In general terms, it is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, and on the north by the Arctic Ocean. The border of Asia with Europe is a social constructionism, historical and cultural construct, as there is no clear physical and geographical separation between them. A commonly accepted division places Asia to the east of the Suez Canal separating it from Africa; and to the east of the Turkish straits, the Ural Mountains an ...
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Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. Antarctica is the fifth-largest continent, being about 40% larger than Europe, and has an area of . Most of Antarctica is covered by the Antarctic ice sheet, with an average thickness of . Antarctica is, on average, the coldest, driest, and windiest of the continents, and it has the highest average elevation. It is mainly a polar desert, with annual Climate of Antarctica#Precipitation, precipitation of over along the coast and far less inland. About 70% of the world's freshwater reserves are frozen in Antarctica, which, if melted, would raise global sea levels by almost . Antarctica holds the record for the Lowest temperature recorded on Earth, lowest measured temperature on Earth, . The coastal regions can reach temperatures over in the ...
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Africa
Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surface area.Sayre, April Pulley (1999), ''Africa'', Twenty-First Century Books. . With nearly billion people as of , it accounts for about of the world's human population. Demographics of Africa, Africa's population is the youngest among all the continents; the median age in 2012 was 19.7, when the worldwide median age was 30.4. Based on 2024 projections, Africa's population will exceed 3.8 billion people by 2100. Africa is the least wealthy inhabited continent per capita and second-least wealthy by total wealth, ahead of Oceania. Scholars have attributed this to different factors including Geography of Africa, geography, Climate of Africa, climate, corruption, Scramble for Africa, colonialism, the Cold War, and neocolonialism. Despite this lo ...
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Anisian
In the geologic timescale, the Anisian is the lower stage (stratigraphy), stage or earliest geologic age, age of the Middle Triassic series (stratigraphy), series or geologic epoch, epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic Epoch) and precedes the Ladinian Age. Stratigraphic definitions The stage and its name were established by Austrian geologists Wilhelm Heinrich Waagen and Carl Diener in 1895. The name comes from ''Anisus'', the Latin name of the river Enns (river), Enns. The original type locality (geology), type locality is at Großreifling in the states of Austria, Austrian state of Styria. The base of the Anisian Stage (also the base of the Middle Triassic series) is sometimes laid at the first appearance of conodont species ''Chiosella, Chiosella timorensis'' in the stratigraphic record. Other stratigraphers prefer to use the base of magnetic chronozone MT1n. There is no accept ...
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Olenekian
In the geologic timescale, the Olenekian is an age (geology), age in the Early Triassic epoch (geology), epoch; in chronostratigraphy, it is a stage (stratigraphy), stage in the Lower Triassic series (stratigraphy), series. It spans the time between annum, Ma and Ma (million years ago). The Olenekian is sometimes divided into the Smithian and the Spathian subages or substages. The Olenekian follows the Induan and is followed by the Anisian (Middle Triassic). The Olenekian saw the deposition of a large part of the Buntsandstein in Europe. The Olenekian is roughly coeval with the regional Yongningzhenian Stage used in China. Stratigraphic definitions The Olenekian Stage was introduced into scientific literature by Russian stratigraphers in 1956. The stage is named after Olenëk in Siberia. Before the subdivision in Olenekian and Induan became established, both stages formed the Scythian Stage, which has since disappeared from the official timescale. The base of the Olenekian is ...
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Parareptilia
Parareptilia ("near-reptiles") is an extinct group of basal sauropsids (" reptiles"), traditionally considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as '' Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by t ...
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Procolophonidae
Procolophonidae is an extinct family (biology), family of small, lizard-like parareptiles known from the Late Permian to Late Triassic that were distributed across Pangaea, having been reported from Europe, North America, China, South Africa, South America, Antarctica and Australia. The most primitive procolophonids were likely insectivorous or omnivorous, more Derived (phylogenetics), derived members of the clade developed Cusp (anatomy), bicusped molars, and were likely herbivorous feeding on high fiber vegetation or Durophagy, durophagous omnivores. Many members of the group are noted for spines projecting from the quadratojugal bone of the skull, which likely served a defensive purpose as well as possibly also for display. At least some taxa were likely fossorial burrowers. While diverse during the Early and Middle Triassic, they had very low diversity during the Late Triassic, and were extinct by the beginning of the Jurassic. Phylogeny The family is defined as all taxa more ...
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