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Perissodactyl
Perissodactyla (, ), or odd-toed ungulates, is an order of Ungulate, ungulates. The order includes about 17 living species divided into three Family (biology), families: Equidae (wild horse, horses, Asinus, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, Vestigiality, vestigial, or positioned posteriorly. By contrast, Artiodactyl, artiodactyls (even-toed ungulates) bear most of their weight equally on four or two (an even number) of the five toes: their third and fourth toes. Another difference between the two is that perissodactyls digest plant cellulose in their intestines, rather than in one or more stomach chambers as artiodactyls, with the exception of Suina, do. The order was considerably more diverse in the past, with notable extinct groups inclu ...
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Ungulate
Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including Equidae, equines, rhinoceroses, and tapirs; and Artiodactyla including Bos, cattle, antelope, Sus (genus), pigs, giraffes, camels, Ovis, sheep, deer, and Hippopotamidae, hippopotamuses, among others. Cetaceans such as Whale, whales, Dolphin, dolphins, and Porpoise, porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes ...
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Tapiridae
Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South and Central America and Southeast Asia. They are one of three extant branches of Perissodactyla (odd-toed ungulates), alongside equines and rhinoceroses. Only a single genus, ''Tapirus'', is currently extant. Tapirs migrated into South America during the Pleistocene epoch from North America after the formation of the Isthmus of Panama as part of the Great American Interchange. Tapirs were formerly present across North America, but became extinct in the region at the end of the Late Pleistocene, around 12,000 years ago. Name The term ''tapir'' comes from the Portuguese-language words , , which themselves trace their origins back to Old Tupi, specifically the term . This word, according to Eduardo de Almeida Navarro, referred in a more precise manner to the species '' ...
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Chalicotheriidae
Chalicotheriidae (from Ancient Greek ''khálix'', "gravel", and ''theríon'', "beast") is an extinct family of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene to the Early Pleistocene. They are often called chalicotheres, a term which is also applied to the broader grouping of Chalicotherioidea. They are noted for their unusual morphology compared to other ungulates, such as their clawed forelimbs. Members of the subfamily Chalicotheriinae developed elongate gorilla-like forelimbs that are thought to have been used to grasp vegetation. They are thought to have been browsers on foliage as well as possibly bark and fruit. History of discovery The first chalicotheres remains discovered were ungual phalanges found near Eppelsheim, Germany in the early 19th century. These remains were considered to belong to gigantic pangolins by Georges Cuvier in 1822 while Johann Jakob Kaup in 1833 alternatively attr ...
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Rhinocerotidae
A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant taxon, extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family (biology), family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South Asia, South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh over half a tonne in adulthood. They have a herbivore, herbivorous diet, small brains for mammals of their size, one or two horns, and a thick , protective skin formed from layers of collagen positioned in a crystal structure, lattice structure. They generally eat leafy material, although their ability to ferment food in their colon (anatomy), hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African ...
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Rhinoceros Male 2003
A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh over half a tonne in adulthood. They have a herbivorous diet, small brains for mammals of their size, one or two horns, and a thick , protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their ...
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Palaeotheriidae
Palaeotheriidae is an extinct family of herbivorous perissodactyl mammals that inhabited Europe, with less abundant remains also known from Asia, from the mid-Eocene to the early Oligocene. They are classified in Equoidea, along with the living family Equidae (which includes zebras, horses and asses). Morphology Palaeotheres ranged widely in size, from small species like '' Palaeotherium lautricense,'' which is estimated to have only weighed to large species like '' Palaeotherium magnum'', which are comparable in size to living equines, with body masses over . Their teeth are brachydont (low crowned). According to Danilo et al. 2013., paleotheriids are distinguished from other equoids by one unambiguous synapomorphy "the nasal notch opening distally to the canine, above the postcanine diastema" and two unambiguous character state changes "an average metaconule on he fourth premolar and "an oblique metastyle on he first and second molars. Taxonomy Palaeotheriidae is ge ...
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Eggysodontidae
Eggysodontidae is a family of perissodactyls, closely related to rhinoceroses. Fossils have been found in Oligocene deposits in Europe, the Caucasus, Central Asia, China, and Mongolia. Taxonomy The family Eggysodontidae has been at times considered a subfamily of Hyracodontidae The Hyracodontidae are an extinction, extinct Family (biology), family of rhinocerotoids endemic to North America, Europe, and Asia during the Eocene through early Oligocene, living from 48.6 to 26.3 million years ago (Mya), existing about . The ..., as Eggysodontinae, by several authors. However, recent analyses have leaned towards recognizing eggysodonts as a distinct family more closely related to crown Rhinocerotoidea than to hyracodonts or paraceratheres.B. Bai and Y.-Q. Wang. 2012. Proeggysodon gen. nov., a primitive Eocene eggysodontine (Mammalia, Perissodactyla) from Erden Obo, Siziwangqi, Nei Mongol, China. Vertebrata PalAsiatica 50(3):204-218 Biology Eggysodonts were ground-dwelling browser ...
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Brontotheriidae
Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos with some developing bony nose horns, and were some of the earliest mammals to have evolved large body sizes of several tonnes. They lived around 56–34 million years ago, until the very close of the Eocene. Brontotheres had a Holarctic distribution, with the exception of Western Europe: they occupied North America, Asia, and Eastern Europe. They were the first fossilized mammals to be discovered west of the Mississippi, and were first discovered in South Dakota. Characteristics and evolution This group has also been referred to as "Titanotheres." "Titan" refers to the mythological Greek gods who were symbols of strength and large size, and "theros" is Greek for "wild animal." "Bronto" is Greek for "thunder," which may be how this group got the nickname "thunder beasts." Brontotheres retai ...
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Amynodontidae
Amynodontidae ("defensive tooth") is a family of extinct perissodactyls related to true rhinoceroses. They are commonly portrayed as semiaquatic hippo-like rhinos but this description only fits members of the Metamynodontini; other groups of amynodonts like the cadurcodontines had more typical ungulate proportions and convergently evolved a tapir-like proboscis. The Greek name of the family describes their tusks, derived from enlarged canine teeth. Odd-toed ungulates are herbivores, so these tusks would have been used either to deter or defend against predators (as suggested by the name) or perhaps in fights among males. Their fossils have been found in North America, and Eurasia ranging in age from the Middle Eocene to the Early Oligocene, with a single genus (''Cadurcotherium'') surviving into the Late Oligocene in South Asia (Pakistan). Taxonomy †Amynodontidae *Subfamily Amynodontinae **Tribe Cadurcodontini ***Genus '' Cadurcodon'' ***Genus '' Lushiamynodon'' ***Genu ...
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Equidae
Equidae (commonly known as the horse family) is the Taxonomy (biology), taxonomic Family (biology), family of Wild horse, horses and related animals, including Asinus, asses, zebra, zebras, and many extinct species known only from fossils. The family evolved more than 50 million years ago, in the Eocene epoch, from a small, multi-toed ungulate into larger, single-toed animals. All Extant taxon, extant species are in the genus ''Equus (genus), Equus'', which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families. It is more specifically grouped within the superfamily (taxonomy), superfamily Equoidea, the only other family being the extinct Palaeotheriidae. The term equid refers to any member of this family, including any equinae, equine. Evolution The oldest known fossils assigned to Equidae were found in North America, and date from the early Eocene epoch, 54 million years ago. Th ...
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Hyracodontidae
The Hyracodontidae are an extinction, extinct Family (biology), family of rhinocerotoids endemic to North America, Europe, and Asia during the Eocene through early Oligocene, living from 48.6 to 26.3 million years ago (Mya), existing about . The Hyracodontidae thrived in the rainforests of Kazakhstan, Pakistan, and southwest China, a former coastal region. Fossil evidence also extends their geographical range to Germany, as well as to Mongolia. References Further reading

* Lucas, S. G. & Sobus, J. C., (1989), The Systematics of Indricotheres. 358–378 in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls, Oxford University Press, New York, New York & Oxford, England. Rhinocerotoidea Prehistoric rhinoceroses Oligocene Perissodactyla Rupelian extinctions Prehistoric mammals of North America Prehistoric mammals of Europe Prehistoric mammals of Asia Eocene first appearances Prehistoric mammal families Taxa named by Edward Drinker Cope {{paleo- ...
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Tapiroidea
Tapiroidea is a superfamily of Perissodactyla, perissodactyls which includes the modern tapir, tapirs and their extinct relatives. Taxonomically, they are placed in suborder Ceratomorpha along with the rhino superfamily, Rhinocerotoidea. The first members of Tapiroidea appeared during the Early Eocene, 55 million years ago, and were present in North America and Asia during the Eocene. Tapiridae first appeared during the early Oligocene in Europe, and are thought to have originated from the tapiroid family Helaletidae. Taxonomy *Superfamily Tapiroidea **Family †Deperetellidae ***Genus †''Bahinolophus'' ***Genus †''Deperetella'' ***Genus †''Irenolophus'' ***Genus †''Teleolophus'' **Family Tapiridae ***Genus †''Eotapirus'' ***Genus †''Miotapirus'' ***Genus †''Nexuotapirus'' ***Genus †''Paratapirus'' ***Genus †''Plesiotapirus'' ***Genus †''Protapirus'' (syn. ''Tanyops'') ***Genus †''Tapiravus'' ***Genus ''Tapirus'' **Family †Helaletidae ***Genus †''Colod ...
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