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Nitrogen-fixing Bacteria
Diazotrophs are organisms capable of nitrogen fixation, i.e. converting the relatively inert diatomic nitrogen (N2) in Earth's atmosphere into bioavailable compound forms such as ammonia. Diazotrophs are typically microorganisms such as bacteria and archaea, with examples being rhizobia and '' Frankia'' and ''Azospirillum''. All diazotrophs contain iron-molybdenum or iron-vanadium nitrogenase systems, and two of the most studied systems are those of '' Klebsiella pneumoniae'' and '' Azotobacter vinelandii'' due to their genetic tractability and their fast growth. Etymology The word diazotroph is derived from the words ''diazo'' ("di" = two + "azo" = nitrogen) meaning "dinitrogen (N2)" and ''troph'' meaning "pertaining to food or nourishment", in summary dinitrogen utilizing. The word ''azote'' means nitrogen in French and was named by French chemist and biologist Antoine Lavoisier, who saw it as the part of air which cannot sustain life. Types Diazotrophs are scattered across Ba ...
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Organism
An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have been proposed to define what an organism is. Among the most common is that an organism has autonomous reproduction, Cell growth, growth, and metabolism. This would exclude viruses, despite the fact that they evolution, evolve like organisms. Other problematic cases include colonial organisms; a colony of eusocial insects is organised adaptively, and has Germ-Soma Differentiation, germ-soma specialisation, with some insects reproducing, others not, like cells in an animal's body. The body of a siphonophore, a jelly-like marine animal, is composed of organism-like zooids, but the whole structure looks and functions much like an animal such as a jellyfish, the parts collaborating to provide the functions of the colonial organism. The evolutiona ...
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Bacterial
Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth, and are present in most of its habitats. Bacteria inhabit the air, soil, water, acidic hot springs, radioactive waste, and the deep biosphere of Earth's crust. Bacteria play a vital role in many stages of the nutrient cycle by recycling nutrients and the fixation of nitrogen from the atmosphere. The nutrient cycle includes the decomposition of dead bodies; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps, extremophile bacteria provide the nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane, to energy. Bacteria also live in mutualistic, commensal an ...
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Nostoc Commune
''Nostoc commune'' is a species of cyanobacterium in the family Nostocaceae. Common names include star jelly, witch's butter, mare's eggs. It is the type species of the genus ''Nostoc'' and is cosmopolitan in distribution. Description ''Nostoc commune'' is a colonial species of cyanobacterium. It initially forms a small, hollow gelatinous globule which grows and becomes leathery, flattened and convoluted, forming a gelatinous mass with other colonies growing nearby. Inside the thin sheath are numerous unbranched hair-like structures called trichomes formed of short cells in a string. The cells are bacteria and thus have no nucleus nor internal membrane system. To multiply, they form two new cells when they divide by binary fission. Along the trichomes, larger specialist nitrogen-fixing cells called heterocysts occur between the ordinary cells. When wet, ''Nostoc commune'' is bluish-green, olive green or brown but in dry conditions it becomes an inconspicuous, crisp brownish m ...
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Anabaena Cylindrica
''Anabaena'' is a genus of filamentous cyanobacteria that exist as plankton. They are known for nitrogen-fixing abilities, and they form symbiotic relationships with certain plants, such as the mosquito fern. They are one of four genera of cyanobacteria that produce neurotoxins, which are harmful to local wildlife, as well as farm animals and pets. Production of these neurotoxins is assumed to be an input into its symbiotic relationships, protecting the plant from grazing pressure. A DNA sequencing project was undertaken by the United States Department of Energy between 1999 and 2005. This project mapped the complete genome of model organism ''Anabaena variabilis'' ATCC 29413, which is 7.2 million base pairs long. A paper detailing the process was published in 2014. The study focused on heterocysts, which convert nitrogen into ammonia. Certain species of ''Anabaena'' have been used on rice paddy fields, proving to be an effective natural fertilizer. Species List of current ...
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Heterocyst
Heterocysts or heterocytes are specialized nitrogen-fixing cells formed during nitrogen starvation by some filamentous cyanobacteria, such as ''Nostoc'', ''Cylindrospermum'', and '' Anabaena''. They fix nitrogen from dinitrogen (N2) in the air using the enzyme nitrogenase, in order to provide the cells in the filament with nitrogen for biosynthesis. Nitrogenase is inactivated by oxygen, so the heterocyst must create a microanaerobic environment. The heterocysts' unique structure and physiology require a global change in gene expression. For example, heterocysts: * produce three additional cell walls, including one of glycolipid that forms a hydrophobic barrier to oxygen * produce nitrogenase and other proteins involved in nitrogen fixation * degrade photosystem II, which produces oxygen * up-regulate glycolytic enzymes * produce proteins that scavenge any remaining oxygen * contain polar plugs composed of cyanophycin which slows down cell-to-cell diffusion Cyanobacteria usu ...
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Photosynthesis
Photosynthesis ( ) is a system of biological processes by which photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical energy necessary to fuel their metabolism. ''Photosynthesis'' usually refers to oxygenic photosynthesis, a process that produces oxygen. Photosynthetic organisms store the chemical energy so produced within intracellular organic compounds (compounds containing carbon) like sugars, glycogen, cellulose and starches. To use this stored chemical energy, an organism's cells metabolize the organic compounds through cellular respiration. Photosynthesis plays a critical role in producing and maintaining the oxygen content of the Earth's atmosphere, and it supplies most of the biological energy necessary for complex life on Earth. Some bacteria also perform anoxygenic photosynthesis, which uses bacteriochlorophyll to split hydrogen sulfide as a reductant instead of water, p ...
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Cyanobacteria
Cyanobacteria ( ) are a group of autotrophic gram-negative bacteria that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" () refers to their bluish green (cyan) color, which forms the basis of cyanobacteria's informal common name, blue-green algae. Cyanobacteria are probably the most numerous taxon to have ever existed on Earth and the first organisms known to have produced oxygen, having appeared in the middle Archean eon and apparently originated in a freshwater or terrestrial environment. Their photopigments can absorb the red- and blue-spectrum frequencies of sunlight (thus reflecting a greenish color) to split water molecules into hydrogen ions and oxygen. The hydrogen ions are used to react with carbon dioxide to produce complex organic compounds such as carbohydrates (a process known as carbon fixation), and the oxygen is released as a byproduct. By continuously producing and releasing oxygen over billions of years, cyanobacte ...
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Escherichia Intermedia
''Escherichia'' ( ) is a genus of Gram-negative, non-spore-forming, facultatively anaerobic, rod-shaped bacteria from the family Enterobacteriaceae. In those species which are inhabitants of the gastrointestinal tracts of warm-blooded animals, ''Escherichia'' species provide a portion of the microbially derived vitamin K for their host. A number of the species of ''Escherichia'' are pathogenic. The genus is named after Theodor Escherich, the discoverer of ''Escherichia coli''. ''Escherichia'' are facultative aerobes, with both aerobic and anaerobic growth, and an optimum temperature of 37 °C. ''Escherichia'' are usually motile by flagella, produce gas from fermentable carbohydrates, and do not decarboxylate lysine or hydrolyze arginine. Species include '' E. albertii'', '' E. fergusonii'', '' E. hermannii'', '' E. ruysiae'', '' E. marmotae'' and most notably, the model organism and clinically relevant ''E. coli''. Formerly, ''Shimwellia blattae'' and Pseudescherichia vulneris ...
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Bacillus Macerans
''Bacillus'', from Latin "bacillus", meaning "little staff, wand", is a genus of Gram-positive, rod-shaped bacteria, a member of the phylum ''Bacillota'', with 266 named species. The term is also used to describe the shape (rod) of other so-shaped bacteria; and the plural ''Bacilli'' is the name of the class of bacteria to which this genus belongs. ''Bacillus'' species can be either obligate aerobes which are dependent on oxygen, or facultative anaerobes which can survive in the absence of oxygen. Cultured ''Bacillus'' species test positive for the enzyme catalase if oxygen has been used or is present. ''Bacillus'' can reduce themselves to oval endospores and can remain in this dormant state for years. The endospore of one species from Morocco is reported to have survived being heated to 420 °C. Endospore formation is usually triggered by a lack of nutrients: the bacterium divides within its cell wall, and one side then engulfs the other. They are not true spores (i.e., ...
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Paenibacillus Polymyxa
''Paenibacillus polymyxa'', also known as ''Bacillus polymyxa'', is a Gram-positive bacterium capable of fixing nitrogen. It is found in soil, plant tissues, marine sediments and hot springs. It may have a role in forest ecosystems and potential future applications as a biofertilizer and biocontrol agent in agriculture. Growth conditions ''P. polymyxa'' can be grown in the laboratory on trypticase soy agar medium. It can also be grown on brain heart infusion agar medium. Applications Agricultural use ''P. polymyxa'' might have possible future applications as a soil inoculant in agriculture and horticulture. Biofilms of ''P. polymyxa'' growing on plant roots have been shown to produce exopolysaccharides which protect the plants from pathogens. The interactions between this bacterial species and plant roots also cause the root hairs to undergo physical changes. Antibiotics Some strains of ''P. polymyxa'' produce antibiotics including fusaricidin and polymyxins. ''P. polymyxa' ...
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Methanococcus
''Methanococcus'' is a genus of coccoid methanogens of the family Methanococcaceae. They are all mesophiles, except the thermophilic '' M. thermolithotrophicus'' and the hyperthermophilic '' M. jannaschii''. The latter was discovered at the base of a "white smoker" chimney at 21° N on the East Pacific Rise and it was the first archaeal genome to be completely sequenced, revealing many novel and eukaryote-like elements. Phylogeny The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI). See also * List of Archaea genera This article lists the genera of the Archaea. The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI). However, in the List provided bel ... References Further reading * * * * * * * * * Archaea genera Euryarchaeota ...
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