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Neoaves
Neoaves is a clade that consists of all modern bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...s (Neornithes or Aves) with the exception of Palaeognathae (ratites and kin) and Galloanserae (ducks, chickens and kin). This group is defined in the '' PhyloCode'' by George Sangster and colleagues in 2022 as "the most inclusive crown clade containing '' Passer domesticus'', but not '' Gallus gallus''". Almost 95% of the roughly 10,000 known species of extant birds belong to the Neoaves. The early diversification of the various neoavian groups occurred very rapidly around the Cretaceous–Paleogene extinction event, and attempts to resolve their relationships with each other have resulted initially in much controversy. Phylogeny The early diversification of the various neoavia ...
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Bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a four-chambered heart, and a strong yet lightweight Bird skeleton, skeleton. Birds live worldwide and range in size from the bee hummingbird to the common ostrich. There are over 11,000 living species and they are split into 44 Order (biology), orders. More than half are passerine or "perching" birds. Birds have Bird wing, wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the Flightless bird, loss of flight in some birds, including ratites, penguins, and diverse endemism, endemic island species. The digestive and respiratory systems of birds are also uniquely a ...
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Passerea
Passerea is a clade of Neoaves, neoavian birds that was proposed by Jarvis ''et al''. (2014). Their genomic analysis recovered two major clades within Neoaves, Passerea and Columbea, and concluded that both clades appear to have many ecologically driven convergent traits. According to Jarvis (2014), these convergences include the foot-propelled diving trait of grebes in Columbea with loons and cormorants in Passerea; the wading-feeding trait of flamingos in Columbea with ibises and egrets in Passerea; and pigeons and sandgrouse in Columbea with shorebirds (killdeer) in Passerea. For Jarvis (2014), these long-known trait and morphological alliances suggest that some of the traditional nongenomic trait classifications are based on Polyphyly, polyphyletic assemblages. Passerea was not recovered in other studies. Phylogeny Cladogram of Passerea relationships based on Jarvis ''et al.'' (2014) with some clade names after Yuri ''et al.'' (2013) and Kimball ''et al.'' 2013. The fol ...
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Gruiformes
The Gruiformes ( ) are an order containing a considerable number of living and extinct bird families, with a widespread geographical diversity. Gruiform means "crane-like". Traditionally, a number of wading and terrestrial bird families that did not seem to belong to any other order were classified together as Gruiformes. These include 15 species of large cranes, about 145 species of smaller crakes and rails, as well as a variety of families comprising one to three species, such as the Heliornithidae, the limpkin, or the Psophiidae. Other birds have been placed in this order more out of necessity to place them ''somewhere''; this has caused the expanded Gruiformes to lack distinctive apomorphies. Recent studies indicate that these "odd Gruiformes" are if at all only loosely related to the cranes, rails, and relatives ("core Gruiformes"). Systematics There are only two suprafamilial clades (natural groups) among the birds traditionally classified as Gruiformes. Rails ( Ra ...
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Cretaceous–Paleogene Extinction Event
The Cretaceous–Paleogene (K–Pg) extinction event, also known as the K–T extinction, was the extinction event, mass extinction of three-quarters of the plant and animal species on Earth approximately 66 million years ago. The event caused the extinction of all non-avian dinosaurs. Most other tetrapods weighing more than also became extinct, with the exception of some ectothermic species such as sea turtles and crocodilians. It marked the end of the Cretaceous period, and with it the Mesozoic era, while heralding the beginning of the current era, the Cenozoic. In the geologic record, the K–Pg event is marked by a thin layer of sediment called the Cretaceous–Paleogene boundary, K–Pg boundary or K–T boundary, which can be found throughout the world in marine and terrestrial rocks. The boundary clay shows unusually high levels of the metal iridium, which is more common in asteroids than in the Earth's crust. As originally proposed in 1980 by a team of scientists le ...
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Columbimorphae
Columbimorphae is a clade/superorder discovered by genome analysis that includes birds of the orders Columbiformes (pigeons and doves), Pterocliformes (sandgrouse), and Mesitornithiformes (mesites). This group was defined in the ''PhyloCode'' by George Sangster and colleagues in 2022 as "the least inclusive crown clade containing ''Columba oenas'', ''Mesitornis variegatus'', and ''Pterocles alchata''". Previous analyses had also recovered this grouping, although the exact relationships differed. Some studies indicated a sister relationship between sandgrouse and pigeons (the traditional view) while other studies favored a sister grouping of mesites and sandgrouse instead. This sister relationship of the sandgrouses and mesites was named by George Sangster and colleagues in 2022 as the clade Pteroclimesites and defined in the ''PhyloCode'' as "the least inclusive crown clade containing ''Mesitornis variegatus'' and ''Pterocles alchata''". In 2020 Kuhl ''et al''. sequenced 3-prime ...
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Eurypygimorphae
Eurypygimorphae or Phaethontimorphae is a clade of birds that contains the orders Phaethontiformes (tropicbirds) and Eurypygiformes (kagu and sunbittern) recovered by genome analysis. The relationship was first identified in 2013 based on their nuclear genes. This group was defined in the ''PhyloCode'' by George Sangster and colleagues in 2022 as "the least inclusive crown clade containing '' Phaethon aethereus'', ''Eurypyga helias The sunbittern (''Eurypyga helias'') is a bittern-like bird of tropical regions of the Americas, and the Monotypic taxon, sole member of the family (biology), family Eurypygidae (sometimes spelled Eurypigidae) and genus ''Eurypyga''. It is found ...'', and '' Rhynochetos jubatus''". Historically these birds were placed at different parts of the tree, with tropicbirds in Pelecaniformes and the kagu and sunbittern in Gruiformes. Some genetic analyses have placed the eurypygimorph taxa in the controversial and obsolete clade Metaves, with uncertai ...
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Columbea
Columbea is a clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ... suggested by genome analysis that contains Columbiformes (pigeons and doves), Pteroclidae (sandgrouse), Mesitornithidae (mesites) and Mirandornithes (flamingos and grebes). Until their recent placement as the sister taxon to Passerea, in the last decade various genetic analysis found them to be in the obsolete clade Metaves. References Neognathae Tetrapod unranked clades {{bird-stub ...
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Mirandornithes
Mirandornithes () is a clade that consists of flamingos and grebes. Many scholars use the term Phoenicopterimorphae for the superorder containing flamingoes and grebes. Determining the relationships of both groups has been problematic. Flamingos had been placed with numerous branches within Neognathae, such as ducks and storks. The grebes had been placed with the loons. However more recent genomic studies have confirmed these two branches as sister groups. Both primitive phoenicopteriformes and their closest relatives, the grebes, were highly aquatic.Mayr, G. (2014) The Eocene ''Juncitarsus'' – its phylogenetic position and significance for the evolution and higher-level affinities of flamingos and grebes. ''Comptes Rendus Palevol''. 13(1):9-18. https://doi.org/10.1016/j.crpv.2013.07.005 This indicates that the entire mirandornithe group evolved from aquatic, probably swimming ancestors. Etymology The term was coined by Sangster in 2005, in order to properly describe the n ...
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Opisthocomiformes
Opisthocomidae is a family of birds, the only named family within the order Opisthocomiformes. The only living representative is the hoatzin (''Opisthocomus hoazin'') which lives in the Amazon and the Orinoco delta in South America. Several fossil species have been identified, including one from Africa and one from Europe. Phylogeny The phylogeny below is based on the work of Hughes & Baker 1999 and Mayr & De Pietri 2014. Traditionally classified among the fowl-like birds (Galliformes), recent studies have favored Opisthocomidae's placement within the Neoaves. Taxonomy * Family Opisthocomidae Swainson 1837Mikko's Phylogeny Archiv ** Genus ?†'' Foro (bird), Foro'' Olson 1992 (mid-Eocene, USA) - cuculiform? *** Species †'' Foro panarium'' Olson 1992 ** Genus ?†'' Onychopteryx'' Cracraft 1971 (Early Eocene of Argentina) – falconid? A ''nomen dubium'' *** Species †'' Onychopteryx simpsoni'' Cracraft 1971 ** Genus †'' Protoazin'' Mayr & De Pietri 2014 (late Eocene of Fr ...
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Otidimorphae
Otidimorphae is a clade of birds that contains the orders Cuculiformes (cuckoos), Musophagiformes (turacos), and Otidiformes (bustards) identified in 2014 by genome analysis. George Sangster and colleagues in 2022 named the clade uniting turacos and bustards as Musophagotides, defining it in the ''PhyloCode The ''International Code of Phylogenetic Nomenclature'', known as the ''PhyloCode'' for short, is a formal set of rules governing phylogenetic nomenclature. Its current version is specifically designed to regulate the naming of clades, leaving the ...'' as "the least inclusive crown clade containing '' Otis tarda'' and '' Musophaga violacea'', but not '' Grus grus'' or '' Mesitornis variegatus''". While the bustards seem to be related to the turacos, other genetic studies have found the cuckoos to be closer to the bustards than the turacos are. References Neognathae {{bird-stub ...
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Phaethoquornithes
Phaethoquornithes is a clade of birds that contains Eurypygimorphae and Aequornithes, which was first recovered by genome analysis in 2014. Members of Eurypygimorphae were originally classified in the obsolete group Metaves, and Aequornithes were classified as the sister taxon to Musophagiformes or Gruiformes. This group has also been informally called Ardeae. Older classifications have used Ardeae in a different sense, as a suborder of Ciconiiformes containing herons and related species. George Sangster and colleagues in 2022 named and defined this clade in the ''PhyloCode'' as the least inclusive crown clade containing '' Phaethon aethereus'' and '' Pelecanus onocrotalus'', but not '' Apus apus'', '' Charadrius hiaticula'', '' Musophaga violacea'', or ''Passer domesticus''. Cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an P ...
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Sunbittern
The sunbittern (''Eurypyga helias'') is a bittern-like bird of tropical regions of the Americas, and the sole member of the family Eurypygidae (sometimes spelled Eurypigidae) and genus ''Eurypyga''. It is found in Central and South America, and has three subspecies. The sunbittern shows both morphological and molecular similarities with the kagu (''Rhynochetos jubatus'') of New Caledonia, indicating a Gondwanan origin, both species being placed in the clade Eurypygiformes.del Hoyo, J. Elliott, A. & Sargatal, J. (editors). (1996) ''Handbook of the Birds of the World. Volume 3: Hoatzin to Auks''. Lynx Edicions. Taxonomy The sunbittern was traditionally placed in the Gruiformes, but this was always considered preliminary. Altogether, the bird is most similar to another bird that was provisionally placed in the Gruiformes, the kagu (''Rhynochetos jubatus''). Molecular studies seem to confirm that the kagu and sunbittern are each other's closest living relatives and have a simil ...
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