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Moradi Formation
The Moradi Formation is a geological formation in Niger. It is of Late Permian age. It is informally divided into three subunits. The lower portion of the formation consists of red mudstone, with muddy calcareous sandstone and quartz-granlule conglomerate present as lenses. The middle portion consists of muddy siltstone in thick beds interbedded with red argillaceous sandstone. The lower two thirds of the upper portion of the formation consist of red siltstone intercalated with channel lag intraformational conglomerates, while the upper third consists of barchanoid shaped lenses of conglomeratic sandstone with ventifacts. These facies are indicatived of deposition under arid conditions, with less than of annual rainfall in the Central Pangean desert, with annual temperatures of , but with ephemeral water presence including lakes. Fossil content The formation is known for its fossils, including the temnospondyls '' Nigerpeton'' and '' Saharastega, ''the pareiasaur ''Bunosteg ...
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Lopingian
The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic. The Lopingian is often synonymous with the informal terms late Permian or upper Permian. The name was introduced by Amadeus William Grabau in 1931 and derives from Leping, Jiangxi in China. It consists of two stages/ ages. The earlier is the Wuchiapingian and the later is the Changhsingian. The International Chronostratigraphic Chart (v2018/07) provides a numerical age of 259.1 ±0.5 Ma. If a Global Boundary Stratotype Section and Point (GSSP) has been approved, the lower boundary of the earliest stage determines numerical age of an epoch. The GSSP for the Wuchiapingian has a numerical age of 259.8 ± 0.4 Ma. Evidence from Milankovitch cycles suggests that the length of an Earth day during this epoch was approximately 22 hours. The Lopingian ended with the Permian–Triassic extinction event. ...
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Interbedding
In geology, interbedding occurs when beds (layers of rock) of a particular lithology lie between or alternate with beds of a different lithology. For example, sedimentary rocks may be interbedded if there were sea level variations in their sedimentary depositional environment. Intercalation is a special case of interbedding where a layer is variably inserted into an already existing sequence; or where two separate depositional environments in close spatial proximity migrate alternately across the contact. While interbedding has layers that are horizontally flat (or aligned with the angle of the entire stratum), intercalated rock on the other hand has slanted layers that streak through each other (even when it aligns with the stratum). For example intercalated conglomerate and sandstone looks like ripples of different material networked through each other somewhat off the horizontal, as the beds are deposited in a gradient. This is likely due to differing fluvial conditions and ...
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Captorhinid
Captorhinidae (also known as cotylosaurs) is an extinct family of tetrapods, traditionally considered primitive reptiles, known from the late Carboniferous to the Late Permian. They had a cosmopolitan distribution across Pangea. Description Captorhinids are a clade of small to very large lizard-like reptiles that date from the late Carboniferous through the Permian. Their skulls were much stronger than those of their relatives, the Protorothyrididae, and had teeth that were better able to deal with tough plant material. The postcranial skeleton is very similar to that of advanced reptiliomorph amphibians, so much in fact that the amphibian Seymouriamorpha and Diadectomorpha were thought to be reptiles and grouped together in "Cotylosauria" as the first reptiles in the early 20th century. Captorhinids have broad, robust skulls that are generally triangular in shape when seen in dorsal view. The premaxillae are characteristically downturned. The largest captorhinid, the herbivorous ...
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Bunostegos
''Bunostegos'' ("knobbly kullroof") is an extinct genus of pareiasaur parareptile from the Late Permian of the Agadez Region in Niger. The type species, ''Bunostegos akokanensis'', was named from the Moradi Formation in 2003. It was a cow-sized animal with a distinctive skull that had large bony knobs, similar in form to those of other pareiasaurs but far larger. The species appears to have lived in a desert in the centre of the supercontinent of Pangaea. Analysis of the limb bones (including the scapulocoracoid, humerus, radius, ulna, pelvis, and femur) was published in 2015, and revealed that ''Bunostegos'' walked upright on four limbs, with the body held above ground. This new information directly suggests that it could be the first tetrapod with a fully erect gait. Description The animal has been described as about the size of a modern cow with a knobbly skull and bony plate armor on its back." Its teeth show it to have been a plant eater. It lived in an isolated desert ...
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Pareiasaur
Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct at the end of the Permian during the Permian-Triassic extinction event. Description Pareiasaurs ranged in size from long, and may have weighed up to . They were stocky, with short tails, small heads, robust limbs, and broad feet. The cow-sized species '' Bunostegos'', which lived 260 million years ago, is the earliest known example of a tetrapod with a fully erect posture as its legs were positioned directly under its body. Pareiasaurs were protected by bony scutes called osteoderms that were set into the skin. Their heavy skulls wer ...
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Journal Of Vertebrate Paleontology
The ''Journal of Vertebrate Paleontology'' is a bimonthly peer-reviewed scientific journal that was established in 1980 by Jiri Zidek (University of Oklahoma). It covers all aspects of vertebrate paleontology, including vertebrate origins, evolution, functional morphology, taxonomy, biostratigraphy, paleoecology, paleobiogeography, and paleoanthropology. The journal is published by Taylor & Francis on behalf of the Society of Vertebrate Paleontology. According to ''Journal Citation Reports'', the journal has a 2017 impact factor The impact factor (IF) or journal impact factor (JIF) of an academic journal is a scientometric index calculated by Clarivate that reflects the yearly mean number of citations of articles published in the last two years in a given journal, as i ... of 2.190. References External links * Paleontology journals Publications established in 1980 Quarterly journals English-language journals Taylor & Francis academic journals {{paleontology-jou ...
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Saharastega
''Saharastega'' is an extinct genus of basal temnospondyl which lived during the Late Permian period, around 251 to 260 million years ago. Remains of ''Saharastega'', discovered by paleontologist Christian A. Sidor at the Moradi Formation in Niger, were described briefly in 2005 and more comprehensively in 2006. The description is based on a skull lacking the lower jaws. Description Skull roof The skull is somewhat wide and gradually narrows towards the snout, with the portion of the skull in front of the orbits (eye sockets) longer than that behind them. It was 36.8 centimeters (14.5 inches) in length and 30 centimeters (11.8 inches) in width at its widest point. The cranial bones are somewhat eroded, but preserved portions are finely textured with pits and ridges as in most adult temnospondyls, but lacking the unornamented areas adjacent to the midline which characterize edopoids. The skull lacks lateral line grooves, which may indicate terrestriality, but the overall siz ...
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Nigerpeton
''Nigerpeton'' is a genus of temnospondyl amphibian which lived during the late Permian (Changhsingian) some 250 million years ago in Niger, in what was then central Pangaea. Specimens of ''Nigerpeton'' were first collected during field work in the Moradi Formation in 2000 and 2003. It is the youngest member of Cochleosauridae, a family of temnospondyls otherwise known from the late Carboniferous and early Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ... of Europe and North America. External links *Steyer, J. S., Damiani, R., Sidor, C. A., O'Keefe, R., Larsson, H. C. E., Maga, A. & Ide, O. (2006The vertebrate fauna of the Upper Permian of Niger. IV. ''Nigerpeton ricqlesi'' (Temnospondyli: Cochleosauridae), and the edopoid colonization of Gondwana ''Journal of Vertebrate ...
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Temnospondyl
Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order (biology), order of small to giant tetrapods—often considered Labyrinthodontia, primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had cha ...
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Pangaea
Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million years ago, and began to break apart about 200 million years ago, at the end of the Triassic and beginning of the Jurassic. In contrast to the present Earth and its distribution of continental mass, Pangaea was centred on the equator and surrounded by the superocean Panthalassa and the Paleo-Tethys Ocean, Paleo-Tethys and subsequent Tethys Ocean, Tethys Oceans. Pangaea is the most recent supercontinent to have existed and the first to be reconstructed by geologists. Origin of the concept The name "Pangaea" is derived from Ancient Greek ''pan'' (, "all, entire, whole") and ''Gaia (mythology), Gaia'' or Gaea (, "Mother goddess, Mother Earth, land"). The concept that the continents once formed a contiguous land mass was hypothesised, with ...
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Arid
A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most arid climates straddle the Equator; these regions include parts of Africa, Asia, South America, North America, and Australia. Change over time The distribution of aridity at any time is largely the result of the general circulation of the atmosphere. The latter does change significantly over time through climate change. For example, temperature increase by 1.5–2.1 percent across the Nile Basin over the next 30–40 years could change the region from semi-arid to arid, significantly reducing the land usable for agriculture. In addition, changes in land use can increase demands on soil water and thereby increase aridity. See also * Arid Forest Research Institute * Aridity index * Desert climate * Desiccation tolerance * Drought * Hu ...
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