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Hesperornithoides
''Hesperornithoides'' (meaning "western bird form"; nicknamed "Lori") is a genus of troodontid theropod dinosaur that lived in North America during the Late Jurassic period. Discovery Although several Troodontid teeth were found, with the troodontid '' Koparion'' named in 1994, a well preserved skeleton wasn't found until in 2001, a field crew from the Tate Museum supervised by William Wahl unexpectedly discovered the type skeleton ''Hesperornithoides'' in Jimbo Quarry of the Morrison Formation, overlying the excavation site of '' Supersaurus vivianae'', near Douglas, Wyoming. The muddy sandstone layers of the Jimbo Quarry from which ''Hesperornithoides'' came from the middle Morrison Formation dating to the between the Oxfordian and Tithonian ages of the Upper Jurassic.Trujillo, K., Chamberlain, K., & Strickland, A. (2006). Oxfordian Park: U/Pb ages from SHRIMP analysis for the Upper Jurassic Morrison Formation of southeastern Wyoming with implications for biostratigraphic corr ...
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Troodontid
Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens (including specimens which preserve feathers, eggs, embryos, and complete juveniles), have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like '' Sinovenator'', demonstrate striking anatomical similarities with ''Archaeopteryx'' and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves. Evolution The oldest definitive troodontid known is '' Hesperornithoides'' from the Late Jurassic of Wyoming. The slightly older '' Koparion'' of Utah is only represented by a single tooth, and small maniraptoran teeth from the Middle Jurassic ...
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Koparion
''Koparion'' is a genus of small coelurosaurian theropod dinosaur, from the late Jurassic Period ( Kimmeridgian stage), of Utah. It contains the single named species ''Koparion douglassi'' which is known only from a single isolated tooth. Discovery In 1993, Daniel Chure and Brooks Britt reported the discovery of small theropod remains, found by screenwashing large amounts of earth in the Rainbow Park near Dinosaur National Monument in Uintah County, Utah. In 1994, Chure named and described a unique tooth as the type specimen of the new species ''Koparion douglassi''. The generic name ''Koparion'' comes from the Ancient Greek κοπάριον, "small surgical knife", in reference to the small size of the tooth. The specific name honors Earl Douglass, who, in the early twentieth century, excavated the Dinosaur National Monument quarry. The holotype specimen, DINO 3353, was found in the Brushy Basin Member of the Morrison Formation dating from the late Kimmeridgian, about ...
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Phylogenetic
In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data and observed heritable traits of DNA sequences, protein amino acid sequences, and morphology. The results are a phylogenetic tree—a diagram depicting the hypothetical relationships among the organisms, reflecting their inferred evolutionary history. The tips of a phylogenetic tree represent the observed entities, which can be living taxa or fossils. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the taxa represented on the tree. An unrooted tree diagram (a network) makes no assumption about directionality of character state transformation, and does not show the origin or "root" of the taxa in question. In addition to their use for inferring phylogenetic pa ...
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Humerus
The humerus (; : humeri) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius (bone), radius and ulna, and consists of three sections. The humeral upper extremity of humerus, upper extremity consists of a rounded head, a narrow neck, and two short processes (tubercles, sometimes called tuberosities). The body of humerus, body is cylindrical in its upper portion, and more prism (geometry), prismatic below. The lower extremity of humerus, lower extremity consists of 2 epicondyles, 2 processes (trochlea of the humerus, trochlea and capitulum of the humerus, capitulum), and 3 fossae (radial fossa, coronoid fossa, and olecranon fossa). As well as its true anatomical neck, the constriction below the greater and lesser tubercles of the humerus is referred to as its Surgical neck of the humerus, surgical neck due to its tendency to fracture, thus often becoming the focus of surgeons. Etymology The word ...
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Quadrate Bone
The quadrate bone is a skull bone in most tetrapods, including amphibians, sauropsids ( reptiles, birds), and early synapsids. In most tetrapods, the quadrate bone connects to the quadratojugal and squamosal bones in the skull, and forms upper part of the jaw joint. The lower jaw articulates at the articular bone, located at the rear end of the lower jaw. The quadrate bone forms the lower jaw articulation in all classes except mammals. Evolutionarily, it is derived from the hindmost part of the primitive cartilaginous upper jaw. Function in reptiles In certain extinct reptiles, the variation and stability of the morphology of the quadrate bone has helped paleontologists in the species-level taxonomy and identification of mosasaur squamates and spinosaurine dinosaurs. In some lizards and dinosaurs, the quadrate is articulated at both ends and movable. In snakes, the quadrate bone has become elongated and very mobile, and contributes greatly to their ability to swallow ...
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Lacrimal Bone
The lacrimal bones are two small and fragile bones of the facial skeleton; they are roughly the size of the little fingernail and situated at the front part of the medial wall of the orbit. They each have two surfaces and four borders. Several bony landmarks of the lacrimal bones function in the process of lacrimation. Specifically, the lacrimal bones help form the nasolacrimal canal necessary for tear translocation. A depression on the anterior inferior portion of one bone, the lacrimal fossa, houses the membranous lacrimal sac. Tears, from the lacrimal glands, collect in this sac during excessive lacrimation. The fluid then flows through the nasolacrimal duct and into the nasopharynx. This drainage results in what is commonly referred to a runny nose during excessive crying or tear production. Injury or fracture of the lacrimal bone can result in posttraumatic obstruction of the lacrimal pathways. Structure Lateral or orbital surface The lateral or orbital surface is divided b ...
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Jugal Bone
The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic bone, zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy The jugal bone is located on either side of the skull in the Ocular scales, circumorbital region. It is the origin of several masticatory muscles in the skull. The jugal and Lacrimal bone, lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. During development, the jugal bone originates from dermal bone. In dinosaurs This bone is considered key in the determination of general traits in cases in which the entire skull has not been found intact (for instance, as with dinosaurs in paleontology). In some dinosaur genera the jugal also forms part of the lower margin of either the antorbital fenestra or the infratemporal fenestr ...
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Paraves
Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosauridae, dromaeosaurids, troodontidae, troodontids, Anchiornithidae, anchiornithids, and possibly the scansoriopterygidae, scansoriopterygids, the group also contains the Avialae, avialans, which include diverse extinct taxa as well as the over 11,000 species of living birds. Basal members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. A number of differing scientific interpretations of the relationships between paravian taxa exist. New fossil discoveries and analyses make the classification of Paraves an active subject of research. Description Like other theropods, all paravians are bipedal, walking on their two hind legs. The teeth of basal paravians were curved and serrated, but not blade-like except in some specialized species, suc ...
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Apomorphic
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Determining apom ...
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