Exostinus
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Exostinus
''Exostinus'' is an extinct genus of xenosaurid lizard from the Late Cretaceous to Oligocene of the western United States. It was named in 1873 by Edward Drinker Cope as ''Exostinus serratus'' for jaw and skull bones found in the White River Formation of Colorado showing distinguishing cranial scutes. Material of ''E. serratus'' is also known from the Brule Formation, also Middle Oligocene in age, but in Wyoming. A second species, ''E. lancensis'' was named in 1928 by Charles W. Gilmore and is also known from skull and jaw bones, from the Maastrichtian Hell Creek, Lance and Fort Union Formations, and the Paleocene Tongue River Formation The Tongue River Member is the uppermost Member (geology), geologic member of the Fort Union Formation in Montana, North Dakota, Wyoming. The strata are yellow or light-colored massive sandstones and numerous thick coal beds. The vertebrate fo .... Other species named by Gilmore are now considered synonyms of ''E. lancensis''. References ...
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Xenosauridae
Xenosauridae is a family of anguimorph lizards whose only living representative is the genus '' Xenosaurus'', which is native to Central America. Xenosauridae also includes the extinct genera '' Exostinus'' and '' Restes''. Also known as knob-scaled lizards, they have rounded, bumpy scales and osteoderms. Most living species prefer humid, rocky habitats, although they are widespread within their native regions, with some inhabiting semi-arid scrub environments. They are carnivorous or insectivorous, and give birth to live young. '' Shinisaurus'', the Chinese crocodile lizard, was once also regarded as a member of Xenosauridae, but most recent studies of the evolutionary relationships of anguimorphs consider ''Shinisaurus'' to be more closely related to monitor lizards and helodermatids than to ''Xenosaurus''. Xenosauridae is part of a larger clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is compose ...
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Fort Union Formation
The Fort Union Formation is a geologic unit containing sandstones, shales, and coal beds in Wyoming, Montana, and parts of adjacent states. In the Powder River Basin, it contains important economic deposits of coal, uranium, and coalbed methane. Description The Fort Union is mostly of Paleocene age and represents a time of extensive swamps as well as fluvial and Lake, lacustrine conditions. The rocks are more sandy in southwestern Wyoming and more coal-bearing in northeast Wyoming and southeast Montana, reflecting a general change from rivers and lakes in the west to swamps in the east, but all three environments were present at various times in most locations. Coal in the Fort Union in the Powder River Basin occurs mainly in the Tongue River Member, where as many as 32 coal seams total more than 300 feet in thickness. One such bed, the Wyodak Coal near Gillette, Wyoming, is as much as thick. Most of the coals in the Fort Union Formation are ranked subbituminous.''Cretaceous an ...
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Wyoming
Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the south. With an estimated population of 587,618 as of 2024, Wyoming is the List of U.S. states and territories by population, least populous state despite being the List of U.S. states and territories by area, 10th largest by area, and it has the List of U.S. states by population density, second-lowest population density after Alaska. The List of capitals in the United States, state capital and List of municipalities in Wyoming, most populous city is Cheyenne, Wyoming, Cheyenne, which had a population of 65,132 in 2020. Wyoming's western half consists mostly of the ranges and rangelands of the Rocky Mountains; its eastern half consists of high-elevation prairie, and is referred to as th ...
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Oligocene Reptiles Of North America
The Oligocene ( ) is a geologic epoch of the Paleogene Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from Ancient Greek (''olígos'') 'few' and (''kainós'') 'new', and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion of gras ...
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Tongue River Formation
The Tongue River Member is the uppermost Member (geology), geologic member of the Fort Union Formation in Montana, North Dakota, Wyoming. The strata are yellow or light-colored massive sandstones and numerous thick coal beds. The vertebrate fossil fauna includes fishes, turtles, crocodiles, and mammals. Mammalian genera known from rocks of both the Torrejonian and the Tiffanian land mammal ages (middle and late Paleocene) are present. See also * List of fossiliferous stratigraphic units in North Dakota * Paleontology in North Dakota * Paleontology in Wyoming References Paleogene Montana Paleogene geology of North Dakota Geological members {{Paleogene-stub File:Tongue River Mbr 1.jpg, Mudstones and sandstone, Tongue River Member, exposed in a roadcut, US Highway 87 ...
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Paleocene
The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by an asteroid impact (Chicxulub impact) and possibly volcanism (Deccan Traps), marked the beginning of the Paleocene and killed off 75% of species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. ...
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Lance Formation
The Lance (Creek) Formation is a division of Late Cretaceous (dating to about 69–66 Ma) rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the latest Mesozoic vertebrate faunas. The Lance Formation is Late Maastrichtian in age (Lancian NALMA, land mammal age), and shares much fauna with the Hell Creek Formation of Montana and North Dakota, the Frenchman Formation of southwest Saskatchewan, and the lower part of the Scollard Formation of Alberta. The Lance Formation occurs above the ''Baculites clinolobatus'' ammonite marine zone in Wyoming, the top of which has been dated to about 69 million years ago, and extends to the K-Pg boundary, 66 million years ago. However, the characteristic land vertebrate fauna of the Lancian age (which take its name from this formation) is only found in the upper strata of the Lance, roughly corresponding to the thinner equivalent formations such as the H ...
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Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation stretches over portions of Montana, North Dakota, South Dakota, and Wyoming. In Montana, the Hell Creek Formation overlies the Fox Hills Formation. The site of Pompeys Pillar National Monument is a small isolated section of the Hell Creek Formation. In 1966, the Hell Creek Fossil Area was designated as a National Natural Landmark by the National Park Service. It is a series of fresh and brackish-water clays, mudstones, and sandstones deposited during the Maastrichtian and Danian (respectively, the end of the Cretaceous period and the beginning of the Paleogene) by fluvial activity in fluctuating river channels and deltas and very occasional peaty swamp deposits along the low-lying eastern continental margin fronting the late Cretaceous West ...
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Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (stratigraphy), Series, the Cretaceous geologic period, Period or system (stratigraphy), System, and of the Mesozoic geologic era, Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). It is named after the city of Maastricht, the capital and largest city of the Limburg (Netherlands), Limburg province in the Netherlands. The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary period, Tertiary extinction event) occurred at the end of this age. In this extinction event, mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-kn ...
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Edward Drinker Cope
Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontology, paleontologist, comparative anatomy, comparative anatomist, herpetology, herpetologist, and ichthyology, ichthyologist. Born to a wealthy Quaker family, he distinguished himself as a child prodigy interested in science, publishing his first scientific paper at the age of 19. Though his father tried to raise Cope as a gentleman farmer, he eventually acquiesced to his son's scientific aspirations. Cope had little formal scientific training, and he eschewed a teaching position for field work. He made regular trips to the Western United States, American West, prospecting in the 1870s and 1880s, often as a member of United States Geological Survey, U.S. Geological Survey teams. A personal feud between Cope and paleontologist Othniel Charles Marsh led to a period of intense fossil-finding competition now known as the Bone Wars. Cope's financial fortunes soured after failed mining ventures i ...
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Middle Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from Ancient Greek (''olígos'') 'few' and (''kainós'') 'new', and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion of gras ...
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