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Ecdysis
Ecdysis is the moulting of the cuticle in many invertebrates of the clade Ecdysozoa. Since the cuticle of these animals typically forms a largely inelastic exoskeleton, it is shed during growth and a new, larger covering is formed. The remnants of the old, empty exoskeleton are called exuviae. After moulting, an arthropod is described as ''teneral'', a ''callow''; it is "fresh", pale and soft-bodied. Within one or two hours, the cuticle hardens and darkens following a Tanning (leather), tanning process analogous to the production of leather. During this short phase the animal expands, since growth is otherwise constrained by the rigidity of the exoskeleton. Growth of the limbs and other parts normally covered by the hard exoskeleton is achieved by transfer of body fluids from soft parts before the new skin hardens. A spider with a small abdomen may be undernourished but more probably has recently undergone ecdysis. Some arthropods, especially large insects with tracheal respira ...
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Procuticle
Arthropods are covered with a tough, resilient integument, cuticle or exoskeleton of chitin. Generally the exoskeleton will have thickened areas in which the chitin is reinforced or stiffened by materials such as minerals or hardened proteins. This happens in parts of the body where there is a need for rigidity or elasticity. Typically the mineral crystals, mainly calcium carbonate, are deposited among the chitin and protein molecules in a process called biomineralization. The crystals and fibres interpenetrate and reinforce each other, the minerals supplying the hardness and resistance to compression, while the chitin supplies the tensile strength. Biomineralization occurs mainly in crustaceans. In insects and arachnids, the main reinforcing materials are various proteins hardened by linking the fibres in processes called sclerotin, sclerotisation and the hardened proteins are called sclerotin. The dorsal tergum, ventral sternum (arthropod anatomy), sternum, and the lateral pleur ...
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Moulting
In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is a process by which an animal casts off parts of its body to serve some beneficial purpose, either at specific times of the year, or at specific points in its life cycle. In medieval times, it was also known as "mewing" (from the French verb "muer", to moult), a term that lives on in the name of Britain's Royal Mews where the King's hawks used to be kept during moulting time before becoming horse stables after Tudor times. Moulting can involve shedding the Epidermis (skin), epidermis (skin), pelage (hair, feathers, fur, wool), or other external layer. In some groups, other body parts may be shed, for example, the entire exoskeleton in arthropods, including the wings in some insects. Examples In birds In birds, moulting is the periodic replacement of feathers by shedding old feathers while producing new ones. Feathers are dead struct ...
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Ecdysozoa
Ecdysozoa () is a group of protostome animals, including Arthropoda (insects, chelicerates (including arachnids), crustaceans, and myriapods), Nematoda, and several smaller phylum (biology), phyla. The grouping of these animal phyla into a single clade was first proposed by Eernisse ''et al.'' (1992) based on a phylogenetic analysis of 141 morphological characters of ultrastructural and embryological phenotypes. This clade, that is, a group consisting of a common ancestor and all its descendants, was formally named by Aguinaldo ''et al.'' in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA genes. A large study in 2008 by Dunn ''et al.'' strongly supported the monophyly of Ecdysozoa. The group Ecdysozoa is supported by many Morphology (biology), morphological characters, including growth by ecdysis, with moulting of the cuticle – without mitosis in the epidermis – under control of the prohormone ecdysone, and internal fertilization. The group was i ...
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Instar
An instar (, from the Latin '' īnstar'' 'form, likeness') is a developmental stage of arthropods, such as insects, which occurs between each moult (''ecdysis'') until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or assume a new form. Differences between instars can often be seen in altered body proportions, colors, patterns, changes in the number of body segments or head width. After shedding their exoskeleton (moulting), the juvenile arthropods continue in their life cycle until they either pupate or moult again. The instar period of growth is fixed; however, in some insects, like the salvinia stem-borer moth, the number of instars depends on early larval nutrition. Some arthropods can continue to moult after sexual maturity, but the stages between these subsequent moults are generally not called instars. For most insect species, an ''instar'' is the developmental stage of the larval forms of holometabolous (complete metamorphism) or ny ...
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Cicada Molting Animated-2
The cicadas () are a superfamily, the Cicadoidea, of insects in the order Hemiptera (true bugs). They are in the suborder Auchenorrhyncha, along with smaller jumping bugs such as leafhoppers and froghoppers. The superfamily is divided into two families, the Tettigarctidae, with two species in Australia, and the Cicadidae, with more than 3,000 species described from around the world; many species remain undescribed. Nearly all cicada species are annual cicadas with the exception of the few North American periodical cicada species, genus ''Magicicada'', which in a given region emerge en masse every 13 or 17 years. Cicadas have prominent eyes set wide apart, short antennae, and membranous front wings. They have an exceptionally loud song, produced in most species by the rapid buckling and unbuckling of drum-like tymbals. The earliest known fossil Cicadomorpha appeared in the Upper Permian period; extant species occur all around the world in temperate to tropical climates. Th ...
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Ecdysone
Ecdysone is a prohormone of the major insect molting hormone 20-hydroxyecdysone, secreted from the prothoracic glands. It is of steroidal structure. Insect molting hormones (ecdysone and its homologues) are generally called ecdysteroids. Ecdysteroids act as moulting hormones of arthropods but also occur in other related phyla where they can play different roles. In ''Drosophila melanogaster'', an increase in ecdysone concentration induces the expression of genes coding for proteins that the larva requires. It causes chromosome puffs (sites of high expression) to form in polytene chromosomes. Recent findings in the laboratory of Chris Q. Doe have found a novel role of this hormone in regulating temporal gene transitions within neural stem cells of the fruit fly. Ecdysone and other ecdysteroids also appear in many plants mostly as a protection agent ( toxins or antifeedants) against herbivorous insects. These phytoecdysteroids have been reputed to have medicinal value. Th ...
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Exopterygota
The Exopterygota (Ancient Greek ''ἔξω'' (éxō, "outside") + ''πτερόν'' (pterón, "wing") + Neo-Latin ''-ota'' ("having")), also known as Hemimetabola, are a superorder of insects of the subclass Pterygota in the infraclass Neoptera, in which the young resemble adults but have externally developing wings. They undergo a modest change between immature and adult, without going through a pupal stage. The nymphs develop gradually into adults through a process of moulting. The Exopterygota are a highly diverse insect superorder, with at least 130,000 living species divided between 15 orders. They include cockroaches, termites, grasshoppers, thrips, lice and stick insects, among many other types of insects. Exopterygota are distinguished from the Endopterygota (or Holometabola) by the way their wings develop. Endopterygota (meaning literally "internal winged forms") develop wings inside the body and undergo an elaborate metamorphosis involving a pupal stage. Exopterygota ...
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Invertebrate Trachea
The trachea (: tracheae or tracheas), also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all animals' lungs. The trachea extends from the larynx and branches into the two primary bronchi. At the top of the trachea, the cricoid cartilage attaches it to the larynx. The trachea is formed by a number of horseshoe-shaped rings, joined together vertically by overlying annular ligaments of trachea, ligaments, and by the trachealis muscle at their ends. The epiglottis closes the opening to the larynx during swallowing. The trachea begins to form in the second month of embryo development, becoming longer and more fixed in its position over time. Its epithelium is lined with columnar epithelium, column-shaped cells that have hair-like extensions called cilia, with scattered goblet cells that produce protective mucins. The trachea can be affected by inflammation or infection, usua ...
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Apolysis
Apolysis ( "discharge, lit. absolution") is the separation of the cuticle from the epidermis in arthropods and related groups (Ecdysozoa). Since the cuticle of these animals is also the skeletal support of the body and is inelastic, it is shed during growth and a new covering of larger dimensions is formed. During this process, an arthropod becomes dormant for a period of time. Enzymes are secreted to digest the inner layers of the existing cuticle, detaching the animal from the outer cuticle. This allows the new cuticle to develop without being exposed to the environmental elements. After apolysis, ecdysis Ecdysis is the moulting of the cuticle in many invertebrates of the clade Ecdysozoa. Since the cuticle of these animals typically forms a largely inelastic exoskeleton, it is shed during growth and a new, larger covering is formed. The remnant ... occurs. Ecdysis is the actual emergence of the arthropod into the environment and always occurs directly after apolysis. The n ...
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Exoskeleton
An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that of a human) which is enclosed underneath other soft tissues. Some large, hard and non-flexible protective exoskeletons are known as mollusc shell, shell or armour (anatomy), armour. Examples of exoskeletons in animals include the arthropod exoskeleton, cuticle skeletons shared by arthropods (insects, chelicerates, myriapods and crustaceans) and tardigrades, as well as the corallite, skeletal cups formed by hardened secretion of stony corals, the test (biology), test/tunic of sea squirts and sea urchins, and the prominent mollusc shell shared by snails, bivalvia, clams, tusk shells, chitons and nautilus. Some vertebrate animals, such as the turtle, have both an endoskeleton and a turtle shell, protective exoskeleton. Role Exoskeletons c ...
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Exuviae
In biology, exuviae are the remains of an exoskeleton and related structures that are left after ecdysozoans (including insects, crustaceans and arachnids) have molted. The exuviae of an animal can be important to biologists as they can often be used to identify the species of the animal and even its sex. As studying insects, crustaceans, or arachnids directly is not always possible, and because exuviae can be collected fairly easily, they can play an important part in helping to determine some general aspects of a species' overall life cycle such as distribution, sex ratio, production, and proof of breeding in a habitat. Exuviae have been suggested as a "gold standard" for insect monitoring. For instance, when monitoring dragonfly populations, the presence of exuviae of a species demonstrates that the species has completed its full life cycle from egg to adult in a habitat. However, it has also been suggested that the fact that exuviae can be hard to find could lead to an under ...
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