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Dennis Robert Hoagland
Dennis Robert Hoagland (April 2, 1884 – September 5, 1949) was an American chemist and plant scientist working in the fields of plant nutrition, agricultural chemistry, and physiology. He was Professor of Plant Nutrition at the University of California at Berkeley from 1927 until his death in 1949. Hoagland is commonly known for discovering the active transport of nutrients in plants, using innovative model systems under controlled experimental conditions, such as solution culture. He and his associates formulated an artificial, complete inorganic nutrient medium, universally known as Hoagland solution, that is still widely used for culturing plants hydroponically. Biography Private life Dennis Hoagland was the son of Charles Breckinridge Hoagland (1859 – 1934) and Lillian May Hoagland (1863 – 1951). He spent his first eight years in Golden and during his later childhood he lived in Denver. He attended the Denver public schools and in 1903 entered Stanford U ...
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Hoagland Solution
The Hoagland solution is a hydroponic nutrient solution that was newly developed by Hoagland and Snyder in 1933, modified by Hoagland and Arnon in 1938, and revised by Arnon in 1950. It is one of the most popular artificial solution compositions for growing plants, in the scientific world at least, with more than 17,000 citations listed by Google Scholar. The Hoagland solution provides all essential elements for plant nutrition and is appropriate for supporting normal growth of a large variety of plant species. The solution described by Dennis Hoagland in 1933, known as Hoagland solution (0), has been modified several times, mainly to add ferric chelates to keep iron effectively in solution, and to optimize the composition and concentration of other trace elements, some of which are not generally credited with a function in plant nutrition. In Hoagland's nutrient recipes of 1938, referred to as Hoagland solution (1, 2), the number of trace elements was subsequently reduced to ...
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Dennis Robert Hoagland
Dennis Robert Hoagland (April 2, 1884 – September 5, 1949) was an American chemist and plant scientist working in the fields of plant nutrition, agricultural chemistry, and physiology. He was Professor of Plant Nutrition at the University of California at Berkeley from 1927 until his death in 1949. Hoagland is commonly known for discovering the active transport of nutrients in plants, using innovative model systems under controlled experimental conditions, such as solution culture. He and his associates formulated an artificial, complete inorganic nutrient medium, universally known as Hoagland solution, that is still widely used for culturing plants hydroponically. Biography Private life Dennis Hoagland was the son of Charles Breckinridge Hoagland (1859 – 1934) and Lillian May Hoagland (1863 – 1951). He spent his first eight years in Golden and during his later childhood he lived in Denver. He attended the Denver public schools and in 1903 entered Stanford U ...
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Hydroponics
Hydroponics is a type of horticulture and a subset of hydroculture which involves growing plants, usually crops or medicinal plants, without soil, by using water-based mineral nutrient solutions in aqueous solvents. Terrestrial or aquatic plants may grow with their roots exposed to the nutritious liquid or in addition, the roots may be mechanically supported by an inert medium such as perlite, gravel, or other substrates. Despite inert media, roots can cause changes of the rhizosphere pH and root exudates can affect rhizosphere biology and physiological balance of the nutrient solution by secondary metabolites. Transgenic plants grown hydroponically allow the release of pharmaceutical proteins as part of the root exudate into the hydroponic medium. The nutrients used in hydroponic systems can come from many different organic or inorganic sources, including fish excrement, duck manure, purchased chemical fertilizers, or artificial nutrient solutions. Plants are commonl ...
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Julius Von Sachs
Julius von Sachs (; 2 October 1832 – 29 May 1897) was a German botanist from Breslau, Prussian Silesia. He is considered the founder of experimental plant physiology and co-founder of modern water culture. Julius von Sachs and Wilhelm Knop are monumental figures in the history of botany by first demonstrating the importance of water culture for the study of plant nutrition and plant physiology in the 19th century. Early life Sachs was born at Breslau on 2 October 1832. His father, Graveur Sachs, was an engraver by trade, and father taught son delineation and accuracy of line and color. From earliest boyhood Julius was fascinated with plants, making collections of them on many field excursions with his father. He gave much of his time between the ages of thirteen and sixteen to drawing and painting the flowers, fungi, and other specimens which he collected. At the Gymnasium from 1845 to 1850 he was most interested in the natural sciences, collecting skulls, writing a monogr ...
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Golden, Colorado
Golden is a List of municipalities in Colorado#Home rule municipality, home rule city that is the county seat of Jefferson County, Colorado, Jefferson County, Colorado, United States. The city population was 20,399 at the 2020 United States Census. Golden lies along Clear Creek (Colorado), Clear Creek at the base of the Front Range of the Rocky Mountains. Founded during the Pike's Peak Gold Rush on June 16, 1859, the mining camp was originally named Golden City in honor of Thomas L. Golden. Golden City served as the capital of the provisional Territory of Jefferson from 1860 to 1861, and capital of the official Territory of Colorado from 1862 to 1867. In 1867, the territorial capital was moved about east to Denver#History, Denver City. Golden is now a part of the Denver–Aurora–Lakewood, CO Metropolitan Statistical Area and the Front Range Urban Corridor. The Colorado School of Mines, offering programs in engineering and science, is located in Golden. In addition, it is also h ...
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Plant Nutrition
Plant nutrition is the study of the chemical elements and compounds necessary for plant growth and reproduction, plant metabolism and their external supply. In its absence the plant is unable to complete a normal life cycle, or that the element is part of some essential plant constituent or metabolite. This is in accordance with Justus von Liebig’s law of the minimum. The total essential plant nutrients include seventeen different elements: carbon, oxygen and hydrogen which are absorbed from the air, whereas other nutrients including nitrogen are typically obtained from the soil (exceptions include some parasitic or carnivorous plants). Plants must obtain the following mineral nutrients from their growing medium: * the macronutrients: nitrogen (N), phosphorus (P), potassium (K), calcium (Ca), sulfur (S), magnesium (Mg), carbon (C), oxygen (O), hydrogen (H) * the micronutrients (or trace minerals): iron (Fe), boron (B), chlorine (Cl), manganese (Mn), zinc (Zn), copper (Cu ...
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Pneumonia
Pneumonia is an inflammatory condition of the lung primarily affecting the small air sacs known as alveoli. Symptoms typically include some combination of productive or dry cough, chest pain, fever, and difficulty breathing. The severity of the condition is variable. Pneumonia is usually caused by infection with viruses or bacteria, and less commonly by other microorganisms. Identifying the responsible pathogen can be difficult. Diagnosis is often based on symptoms and physical examination. Chest X-rays, blood tests, and culture of the sputum may help confirm the diagnosis. The disease may be classified by where it was acquired, such as community- or hospital-acquired or healthcare-associated pneumonia. Risk factors for pneumonia include cystic fibrosis, chronic obstructive pulmonary disease (COPD), sickle cell disease, asthma, diabetes, heart failure, a history of smoking, a poor ability to cough (such as following a stroke), and a weak immune system. Vaccines to ...
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Denver
Denver () is a consolidated city and county, the capital, and most populous city of the U.S. state of Colorado. Its population was 715,522 at the 2020 census, a 19.22% increase since 2010. It is the 19th-most populous city in the United States and the fifth most populous state capital. It is the principal city of the Denver–Aurora–Lakewood, CO Metropolitan Statistical Area and the first city of the Front Range Urban Corridor. Denver is located in the Western United States, in the South Platte River Valley on the western edge of the High Plains just east of the Front Range of the Rocky Mountains. Its downtown district is immediately east of the confluence of Cherry Creek and the South Platte River, approximately east of the foothills of the Rocky Mountains. It is named after James W. Denver, a governor of the Kansas Territory. It is nicknamed the ''Mile High City'' because its official elevation is exactly one mile () above sea level. The 105th meridian we ...
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Artificiality
Artificiality (the state of being artificial or manmade) is the state of being the product of intentional human manufacture, rather than occurring naturally through processes not involving or requiring human activity. Connotations Artificiality often carries with it the implication of being false, counterfeit, or deceptive. The philosopher Aristotle wrote in his '' Rhetoric'': However, artificiality does not necessarily have a negative connotation, as it may also reflect the ability of humans to replicate forms or functions arising in nature, as with an artificial heart or artificial intelligence. Political scientist and artificial intelligence expert Herbert A. Simon observes that "some artificial things are imitations of things in nature, and the imitation may use either the same basic materials as those in the natural object or quite different materials.Herbert A. Simon, ''The Sciences of the Artificial'' (1996), p. 4. Simon distinguishes between the artificial and the synt ...
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Model Organism
A model organism (often shortened to model) is a non-human species that is extensively studied to understand particular biological phenomena, with the expectation that discoveries made in the model organism will provide insight into the workings of other organisms. Model organisms are widely used to research human disease when human experimentation would be unfeasible or unethical. This strategy is made possible by the common descent of all living organisms, and the conservation of metabolic and developmental pathways and genetic material over the course of evolution. Studying model organisms can be informative, but care must be taken when generalizing from one organism to another. In researching human disease, model organisms allow for better understanding the disease process without the added risk of harming an actual human. The species chosen will usually meet a determined taxonomic equivalency to humans, so as to react to disease or its treatment in a way that resembles ...
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Nutrient
A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excreted by cells to create non-cellular structures, such as hair, scales, feathers, or exoskeletons. Some nutrients can be metabolically converted to smaller molecules in the process of releasing energy, such as for carbohydrates, lipids, proteins, and fermentation products (ethanol or vinegar), leading to end-products of water and carbon dioxide. All organisms require water. Essential nutrients for animals are the energy sources, some of the amino acids that are combined to create proteins, a subset of fatty acids, vitamins and certain minerals. Plants require more diverse minerals absorbed through roots, plus carbon dioxide and oxygen absorbed through leaves. Fungi live on dead or living organic matter and meet nutrient needs from their host. ...
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Active Transport
In cellular biology, ''active transport'' is the movement of molecules or ions across a cell membrane from a region of lower concentration to a region of higher concentration—against the concentration gradient. Active transport requires cellular energy to achieve this movement. There are two types of active transport: primary active transport that uses adenosine triphosphate ( ATP), and secondary active transport that uses an electrochemical gradient. Some examples of active transport include: * Phagocytosis of bacteria by macrophages * Movement of calcium ions out of cardiac muscle cells * Transportation of amino acids across the intestinal lining in the human gut * Secretion of proteins such as enzymes, peptide hormones, and antibodies from various cells * Functioning of white blood cells to defend invading diseases Active cellular transportation (ACT) Unlike passive transport, which uses the kinetic energy and natural entropy of molecules moving down a gradient, active ...
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