Dipnomorph
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Dipnomorph
Lungfish are freshwater vertebrates belonging to the class Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, including the presence of lobed fins with a well-developed internal skeleton. Lungfish represent the closest living relatives of the tetrapods (which includes living amphibians, reptiles, birds and mammals). The mouths of lungfish typically bear tooth plates, which are used to crush hard shelled organisms. Today there are only six known species of lungfish, living in Africa, South America, and Australia, though they were formerly globally distributed. The fossil record of the group extends into the Early Devonian, over 410 million years ago. The earliest known members of the group were marine, while almost all post-Carboniferous representatives inhabit freshwater environments. Etymology Dipnoi is Modern Latin derived from the Greek δίπν ...
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Sarcopterygii
Sarcopterygii (; )—sometimes considered synonymous with Crossopterygii ()—is a clade (traditionally a class (biology), class or subclass) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish. These vertebrates are characterised by prominent muscular limb buds (lobes) within their fish fin, fins, which are supported by articulated appendicular skeletons. This is in contrast to the other clade of bony fish, the Actinopterygii, which have only skin-covered lepidotrichia, bony spines supporting the fins. The tetrapods, a mostly terrestrial animal, terrestrial clade of vertebrates, are now recognized as having evolved from sarcopterygian ancestors and are most closely related to lungfishes. Their paired pectoral fins, pectoral and pelvic fins evolved into limb (anatomy), limbs, and their lung bud, foregut diverticulum eventually evolved into air-breathing lungs. Cladistics, Cladistically, this would make the tetrapods a subgroup within ...
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Tetrapods
A tetrapod (; from Ancient Greek τετρα- ''(tetra-)'' 'four' and πούς ''(poús)'' 'foot') is any four- limbed vertebrate animal of the clade Tetrapoda (). Tetrapods include all extant and extinct amphibians and amniotes, with the latter in turn evolving into two major clades, the sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (extinct "pelycosaurs", therapsids and all extant mammals, including humans). Hox gene mutations have resulted in some tetrapods becoming limbless (snakes, legless lizards, and caecilians) or two-limbed (cetaceans, sirenians, some lizards, kiwis, and the extinct moa and elephant birds). Nevertheless, they still qualify as tetrapods through their ancestry, and some retain a pair of vestigial spurs that are remnants of the hindlimbs. Tetrapods evolved from a group of primitive semiaquatic animals known as the tetrapodomorphs which, in turn, evolved from ancient lobe-finned fish ( sarcopterygians) aro ...
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Tetrapod
A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetrapoda (). Tetrapods include all Neontology#Extant taxa versus extinct taxa, extant and Extinction, extinct amphibians and amniotes, with the latter in turn Evolution, evolving into two major clades, the Sauropsida, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (extinct pelycosaur, "pelycosaurs", therapsids and all extant mammals, including Homo sapiens, humans). Hox gene mutations have resulted in some tetrapods becoming Limbless vertebrate, limbless (snakes, legless lizards, and caecilians) or two-limbed (cetaceans, sirenians, Bipedidae, some lizards, kiwi (bird), kiwis, and the extinct moa and elephant birds). Nevertheless, they still qualify as tetrapods through their ancestry, and some retain a pair of ves ...
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Queensland Lungfish
The Australian lungfish (''Neoceratodus forsteri''), also known as the Queensland lungfish, Burnett salmon and barramunda, is the only surviving member of the family Neoceratodontidae. It is one of only six extant lungfish species in the world. Endemic to Australia, the Neoceratodontidae are an ancient family belonging to the class Sarcopterygii, or lobe-finned fishes. Fossil records of this group date back 380 million years, around the time when the higher vertebrate classes were beginning to evolve. Fossils of lungfish almost identical to this species have been uncovered in northern New South Wales, indicating that ''Neoceratodus'' has remained virtually unchanged for well over 100 million years, making it a living fossil and one of the oldest living vertebrate genera on the planet. It is one of six extant representatives of the ancient air-breathing Dipnoi (lungfishes) that flourished during the Devonian period (about 413–365 million years ago) and is the outgroup to all ...
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Notochord
The notochord is an elastic, rod-like structure found in chordates. In vertebrates the notochord is an embryonic structure that disintegrates, as the vertebrae develop, to become the nucleus pulposus in the intervertebral discs of the vertebral column. In non-vertebrate chordates a notochord persists. The notochord is derived from the embryonic mesoderm and consists of an inner core of vacuolated cells filled with glycoproteins, covered by two helical collagen-elastin sheaths. It lies longitudinally along the rostral-caudal (head to tail) axis of the body, dorsal to the gut tube, and ventral to the dorsal nerve cord. Some chordate invertebrates, such as tunicates, develop a notochord during the larval stage but lose it through subsequent stages into adulthood. The notochord is important for signaling the dorso-ventral patterning of cells coming from the mesodermal progenitors. This helps form the precursors needed for certain organs and the embryo to develop ...
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Choana
The choanae (: choana), posterior nasal apertures or internal nostrils are two openings found at the back of the nasal passage between the nasal cavity and the pharynx, in humans and other mammals (as well as crocodilians and most skinks). They are considered one of the most important synapomorphies of tetrapodomorphs, that allowed the passage from water to land. In animals with secondary palates, they allow breathing when the mouth is closed. Janvier, Philippe (2004) "Wandering nostrils". ''Nature'', 432 (7013): 23–24. In tetrapods without secondary palates their function relates primarily to olfaction (sense of smell). The choanae are separated in two by the vomer. Boundaries A choana is the opening between the nasal cavity and the nasopharynx. It is therefore not a structure but a space bounded as follows: * anteriorly and inferiorly by the horizontal plate of palatine bone, * superiorly and posteriorly by the sphenoid bone * laterally by the medial pterygoid pla ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is Cladogram#Homoplasies, homoplasy. The recurrent evolution of flight is a classic example, as flying pterygota, insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas ''homology (biology), homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different trai ...
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Greenwood Press (publisher)
Greenwood Publishing Group, Inc. (GPG) was an educational and academic publisher (middle school through university level) which was part of ABC-Clio. Since 2021, ABC-Clio and its suite of imprints, including GPG, are collectively imprints of British publishing house Bloomsbury Publishing. The Greenwood name stopped being used for new books in 2023. Established in 1967 as Greenwood Press, Inc., and based in Westport, Connecticut, GPG published reference works under its Greenwood Press imprint; and scholarly, professional, and general-interest books under its related imprint, Praeger Publishers (). Also part of GPG was Libraries Unlimited, which published professional works for librarians and teachers. Both of the latter became stand-alone imprints of ABC-Clio, in 2008–2009, after its purchase of GPG. History 1967–1999 The company was founded as Greenwood Press, Inc. (GPI) in 1967 by Harold Mason, a librarian and antiquarian bookseller, and Harold Schwartz, who had a backg ...
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South American Lungfish
The South American lungfish (''Lepidosiren paradoxa''), also known as the American mud-fish and scaly salamanderfish, is the single species of lungfish found in swamps and slow-moving waters of the Amazon Basin, Amazon, Paraguay River, Paraguay, and lower Paraná River drainage basin, basins in South America. Notable as an obligate air-breather, it is the sole member of its family Lepidosirenidae, although some authors also place ''Protopterus'' in the family. In Brazil, it is known by the indigenous language Tupi language, Tupi name , which means "snake-fish" (), and synonyms (), (), and (). Taxonomy The South American lungfish is most closely related to the African lungfishes (family Protopterus, Protopteridae), and both families are thought to have diverged during the Early Cretaceous. Some papers suggest classifying both ''Lepidosiren'' and ''Protopterus'' within Lepidosirenidae, though authorities continue to classify both as distinct families. Description The immature lu ...
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Ray-finned Fish
Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of skin supported by radially extended thin bony spines called '' lepidotrichia'', as opposed to the bulkier, fleshy lobed fins of the sister clade Sarcopterygii (lobe-finned fish). Resembling folding fans, the actinopterygian fins can easily change shape and wetted area, providing superior thrust-to-weight ratios per movement compared to sarcopterygian and chondrichthyian fins. The fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the articulation between these fins and the internal skeleton (e.g., pelvic and pectoral girdles). The vast majority of actinopterygians are teleosts. By species count, they dominate the subphylum Vertebrata, and constitute nearly 99% of the over 30,000 extant ...
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Homology (biology)
In biology, homology is similarity in anatomical structures or genes between organisms of different taxa due to shared ancestry, ''regardless'' of current functional differences. Evolutionary biology explains homologous structures as retained heredity from a common descent, common ancestor after having been subjected to adaptation (biology), adaptive modifications for different purposes as the result of natural selection. The term was first applied to biology in a non-evolutionary context by the anatomist Richard Owen in 1843. Homology was later explained by Charles Darwin's theory of evolution in 1859, but had been observed before this from Aristotle's biology onwards, and it was explicitly analysed by Pierre Belon in 1555. A common example of homologous structures is the forelimbs of vertebrates, where the bat wing development, wings of bats and origin of avian flight, birds, the arms of primates, the front flipper (anatomy), flippers of whales, and the forelegs of quadrupedalis ...
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Cosmine
Cosmine is a spongy, bony material that makes up the dentine-like layers in the scales of the lobe-finned fishes of the class Sarcopterygii. Fish scales that include layers of cosmine are known as cosmoid scales. Description As traditionally described, cosmine consists of a layer of dentine covered by a continuous sheet of enamel. Pulp cavities, which secrete dentine tubules, are surrounded by a complex polygonal network of 'pore cavities' which pierce the overlying enamel layer, giving cosmine its characteristic dotted appearance. The pulp cavities and pore chambers are connected by a complex, reticulated pore canal network which continues into a layer of vascular bone beneath the dentine. The exact configuration of the pore canal network and shape of the pore chambers differs between various taxa, although the general organization into a single layer of enamel over dentine with pore canals with vascular bone underneath remains consistent, at least within the Sarcopterygii. ...
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