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Conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class (biology), class, Pinopsida. All Neontology, extant conifers are perennial plant, perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include Cedrus, cedars, Pseudotsuga, Douglas-firs, Cupressaceae, cypresses, firs, junipers, Agathis, kauri, larches, pines, Tsuga, hemlocks, Sequoioideae, redwoods, spruces, and Taxaceae, yews.Campbell, Reece, "Phylum Coniferophyta". Biology. 7th. 2005. Print. P. 595 As of 1998, the division Pinophyta was estimated to contain eight families, 68 genera, and 629 living species. Although the total number of species is relatively small, conifers are ecology, ecologically important. They are the dominant plants over large areas of land, most ...
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Pinales
The order Pinales in the division Pinophyta, class Pinopsida, comprises all the extant conifers. The distinguishing characteristic is the reproductive structure known as a cone produced by all Pinales. All of the extant conifers, such as cedar, celery-pine, cypress, fir, juniper, larch, pine, redwood, spruce, yew and ''Araucaria araucana'' ("Monkey tail tree" or "Monkey puzzle tree") are included here. Some fossil conifers, however, belong to other distinct orders within the division Pinophyta. Multiple molecular studies indicate this order being paraphyletic with respect to Gnetales, with studies recovering Gnetales as either a sister group to Pinaceae or being more derived than Pinaceae but sister to the rest of the group. Taxonomy History Brown (1825) first discerned that there were two groups of seed plants, distinguished by the form of seed development, based on whether the ovules were exposed, receiving pollen directly, or enclosed, which do not. Shortly afterwar ...
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Gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνόσπερμος ( el, γυμνός, translit=gymnos, lit=naked, label=none and el, σπέρμα, translit=sperma, lit=seed, label=none), literally meaning 'naked seeds'. The name is based on the unenclosed condition of their seeds (called ovules in their unfertilized state). The non-encased condition of their seeds contrasts with the seeds and ovules of flowering plants (angiosperms), which are enclosed within an ovary. Gymnosperm seeds develop either on the surface of scales or leaves, which are often modified to form cones, or solitary as in yew, ''Torreya'', ''Ginkgo''. Gymnosperm lifecycles involve alternation of generations. They have a dominant diploid sporophyte phase and a reduced haploid gametophyte phase which is dependent on ...
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Cupressales
Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecious trees and shrubs up to tall. The bark of mature trees is commonly orange- to red- brown and of stringy texture, often flaking or peeling in vertical strips, but smooth, scaly or hard and square-cracked in some species. Description The leaves are arranged either spirally, in decussate pairs (opposite pairs, each pair at 90° to the previous pair) or in decussate whorls of three or four, depending on the genus. On young plants, the leaves are needle-like, becoming small and scale-like on mature plants of many genera; some genera and species retain needle-like leaves throughout their lives. Old leaves are mostly not shed individually, but in small sprays of foliage (cladoptosis); exceptions are leaves on the shoot ...
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Conifer Cone
A conifer cone (in formal botany, botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers and cycads. The cone of Pinophyta (conifer clade) contains the plant sexuality, reproductive structures. The woody cone is the female cone, which produces seeds. The male cone, which produces pollen, is usually herbaceous plant, herbaceous and much less conspicuous even at full maturity. The name "cone" derives from Greek ''konos'' (pine cone), which also gave name to the cone (geometry), geometric cone. The individual plates of a cone are known as ''scales''. The ''umbo'' of a conifer cone refers to the first year's growth of a seed scale on the cone, showing up as a protuberance at the end of the two-year-old scale. The male cone (microstrobilus or pollen cone) is structurally similar across all conifers, differing only in small wa ...
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Cupressaceae
Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecious trees and shrubs up to tall. The bark of mature trees is commonly orange- to red- brown and of stringy texture, often flaking or peeling in vertical strips, but smooth, scaly or hard and square-cracked in some species. Description The leaves are arranged either spirally, in decussate pairs (opposite pairs, each pair at 90° to the previous pair) or in decussate whorls of three or four, depending on the genus. On young plants, the leaves are needle-like, becoming small and scale-like on mature plants of many genera; some genera and species retain needle-like leaves throughout their lives. Old leaves are mostly not shed individually, but in small sprays of foliage (cladoptosis); exceptions are leaves on the s ...
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Araucariales
Araucariaceae – also known as araucarians – is an extremely ancient family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods and the early Cenozoic, when it was distributed almost worldwide. Most of the Araucariaceae in the Northern Hemisphere vanished in the Cretaceous–Paleogene extinction event, and they are now largely confined to the Southern Hemisphere, except for a few species of '' Agathis'' in Southeast Asia. Description Members of Araucariaceae are typically extremely tall evergreen trees, reaching heights of or more. They can also grow very large stem diameters; a New Zealand kauri tree (''Agathis australis'') named ''Tāne Mahuta'' ("The Lord of the Forest") has been measured at tall with a diameter at breast height of . Its total wood volume is calculated to be , making it the third-largest conifer after '' Sequoia'' and ''Sequoiadendron'' (both from the Cupressaceae subfamily Sequoioideae). The trun ...
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Araucariaceae
Araucariaceae – also known as araucarians – is an extremely ancient family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods and the early Cenozoic, when it was distributed almost worldwide. Most of the Araucariaceae in the Northern Hemisphere vanished in the Cretaceous–Paleogene extinction event, and they are now largely confined to the Southern Hemisphere, except for a few species of ''Agathis'' in Southeast Asia. Description Members of Araucariaceae are typically extremely tall evergreen trees, reaching heights of or more. They can also grow very large stem diameters; a New Zealand kauri tree (''Agathis australis'') named ''Tāne Mahuta'' ("The Lord of the Forest") has been measured at tall with a diameter at breast height of . Its total wood volume is calculated to be , making it the third-largest conifer after '' Sequoia'' and ''Sequoiadendron'' (both from the Cupressaceae subfamily Sequoioideae). The trunks ...
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Podocarpaceae
Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Press. . It contains 19 genera if ''Phyllocladus'' is included and ''Manoao'' and ''Sundacarpus'' are recognized. The family is a classic member of the Antarctic flora, with its main centres of diversity in Australasia, particularly New Caledonia, Tasmania, and New Zealand, and to a slightly lesser extent Malesia and South America (primarily in the Andes Mountains). Several genera extend north of the equator into Indochina and the Philippines. ''Podocarpus'' reaches as far north as southern Japan and southern China in Asia, and Mexico in the Americas, and ''Nageia'' into southern China and southern India. Two genera also occur in sub-Saharan Africa, the widespread ''Podocarpus'' and the endemic '' Afrocarpus''. ''Parasitaxus usta'' is uniqu ...
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Taxaceae
Taxaceae (), commonly called the yew family, is a coniferous family which includes six extant and two extinct genera, and about 30 species of plants, or in older interpretations three genera and 7 to 12 species. Description They are many-branched, small trees and shrubs. The leaves are evergreen, spirally arranged, often twisted at the base to appear 2-ranked. They are linear to lanceolate, and have pale green or white stomatal bands on the undersides. The plants are dioecious, or rarely monoecious. The catkin like male cones are long, and shed pollen in the early spring. They are sometimes externally only slightly differentiated from the branches. The fertile bracts have 2-8 pollen sacs. The female 'cones' are highly reduced. Only the upper or uppermost bracts are fertile and bear one or rarely two seeds. The ovule usually exceeds the scale, although ovules are sometimes rarely enclosed by it. They may be found on the ends of branches or on the branches. They may grow sin ...
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Pinaceae
The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly known as Coniferales. Pinaceae are supported as monophyletic by their protein-type sieve cell plastids, pattern of proembryogeny, and lack of bioflavonoids. They are the largest extant conifer family in species diversity, with between 220 and 250 species (depending on Taxonomy (biology), taxonomic opinion) in 11 genera, and the second-largest (after Cupressaceae) in geographical range, found in most of the Northern Hemisphere, with the majority of the species in temperate climates, but ranging from subarctic to tropical. The family often forms the dominant component of Boreal forest, boreal, coastal, and montane forests. One species, ''Pinus merkusii'', grows just south of the equator in Southeast Asia. Major centre of diversity, c ...
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Sciadopityaceae
''Sciadopitys verticillata'', the or Japanese umbrella-pine, is a unique conifer endemic to Japan. It is the sole member of the family Sciadopityaceae and genus ''Sciadopitys'', a living fossil with no close relatives. The oldest fossils of ''Sciadopitys'' are from the Late Cretaceous of Japan, and the genus was widespread in Laurasia during most of the Cenozoic, especially in Europe until the Pliocene. Taxonomy Molecular evidence indicates that Sciadopityaceae is the sister group to a clade comprising Taxaceae and Cupressaceae, and has an extremely ancient divergence, having diverged from the rest of the conifers during the early mid-Permian; this would also make it a survivor of the Permian–Triassic extinction event. There is inconsistent evidence regarding the plant family which produced Baltic amber. Both macrofossil and microfossil evidence suggest a ''Pinus'' relative, whereas chemical and infrared microspectroscopy evidence suggest relatives of either '' Agathis'' ...
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Palissyales
Palissyales are an extinct order of conifers, known from the Mesozoic. They are best known from the genus '' Palissya'', which is found in Laurasia and Eastern Gondwana dating from the Late Triassic to Early Cretaceous. The only other confirmed genus of the family, '' Stachyotaxus'' known from the Late Triassic of the Northern Hemisphere. The genus '' Knezourocarpon'' from the Jurassic of Australia has also been tentatively considered a member of the order. The cone of the best known genus ''Palissya'' is noted for its unusual construction, which is borne on a large bract (modified leaf), and consists of two parallel rows of ovules that run along the midline of the adaxial surface of the bract which are encased in cup-like structures formed by scales. The bracts are helically arranged around an axis, forming a compound catkin-like structure. The seeds are thin-walled were likely only viable for a short period of time, and were likely adapted to wind dispersal. ''Palissya'' has been ...
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