Rhynchonellata
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Rhynchonellata
The Rhynchonellata is a class of Lower Cambrian to Holocene, Recent Articulata (Brachiopoda), articulate brachiopods that combines orders from within the Rhynchonelliformea (Articulata revised) with well developed pedicle attachment. Shell forms vary from those with wide hinge lines to beaked forms with virtually no hinge line and from generally smooth to strongly plicate. Most all are biconvex. Lophophores vary and include both looped and spiraled forms. Although morphologically distinct, included orders follow a consistent phylogenetic sequence. Orders Orders assigned to the Rhynchonellata, in temporal sequence, include: * †Protorthida (Lower Cambrian – Upper Devonian) * †Orthida (Lower Cambrian – Upper Permian) * †Pentamerida (Middle Cambrian – Upper Devonian) * †Atrypida (Lower Ordovician – Upper Devonian) * Rhynchonellida (Lower Ordovician – Recent) * †Spiriferida (Upper Ordovician – Lower Jurassic) * †Athyridida (Upper Ordovician – Lower Juras ...
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Articulata (Brachiopoda)
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations (hence the name) and separate sets of simple opening and closing muscles. The Rhynchonelliformea (as described in the Treatise Part H, revised 1997–2007) is divided into five classes: Obolellata, Kutorginata, Chileata, Strophomenata, and Rhynchonellata. The Rhynchonellata are found living today, as the major constituent of modern brachiopod faunas. The other classes are all extinct: the Obolellata and Kutorginata are restricted to the Cambrian, while the Chileata and Strophomenata range through most of the Paleozoic. Anatomy Like all bra ...
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Pentamerida
Pentamerida is an order of biconvex, impunctate shelled, articulate brachiopods that are found in marine sedimentary rocks that range from the Middle Cambrian through the Devonian.Moore, Lalcker and Fischer, 1952, Invertebrate Fossils, McGraw-Hill Pentamerids are characterized by a short hinge line where the two valves articulate, inner areas above the hinge line that slope inwardly from the beak of each valve, and a well-developed spondylium on the pedicle valve. The spondylium is a raised platform for muscle attachment found in the middle of the interior pedicle valve, generally toward the hinge and beak. The pedicle valve is the one that the pedicle, or hold-fast stalk, attaches to. The brachidia, which hold the lophophore, the ciliated feeding arms, are looped, as in the Orthida. The Short hinge line helps distinguish the pentamerids from the ancestral orthids from which they are obviously derived. The hinge line is not as short as found in the Rhynchonellida The taxono ...
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Athyridida
Athyridida is an order of Paleozoic brachiopods included in the Rhynchonellata, which makes up part of the articulate brachiopods. The Athyridida are the Rostrospracea of R.C. Moore, 1952,Moore, Lalcker and Fischer, 1952, Invertebrate Fossils, McGraw-Hill considered at that time to be a suborder of the Spiriferida. As with the Spiriferida, the Athyridida have outwardly directed spiral brachidia that support the lophophores on either side, but instead have non-plicate shells with rounded outlines and prominent beaks but almost no inner areas on the pedicle valve. Athyridids began early in the Silurian, reached their greatest diversity in the following Devonian, and from then declined steadily until almost becoming extinct at the end of the Permian The order rejuvenated somewhat during the Triassic, only to decline again until becoming extinct in the Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (st ...
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Thecideida
Thecideida is an order of cryptic articulate brachiopods characterized by their small size and habit of cementing their ventral valves to hard substrates such as shells, rocks and carbonate hardgrounds. Thecideides first appeared in the Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ... and are common today. Taxonomy Order Thecideida *Superfamily Thecideoidea Gray, 1840 *Superfamily Thecospiroidea Bittner, 1890 References Brachiopod orders Rhynchonellata {{Brachiopod-stub ...
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Atrypida
Atrypida is an extinct order of rhynchonelliform brachiopods. They first appeared in middle Ordovician and survived the Ordovician-Silurian extinction, becoming the dominant brachiopods of the Silurian alongside the order Pentamerida. They would survive into the Late Devonian before going extinct at the end of the Frasnian (Late Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago ( Ma), to the beginning of the succeeding ...). References Brachiopod orders Rhynchonellata Silurian brachiopods Ordovician brachiopods Devonian brachiopods Frasnian extinctions Middle Ordovician first appearances {{Brachiopod-stub ...
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Protorthida
Protorthida is an extinct order of Rhynchonellate brachiopods containing the taxa: * Superfamily Protorthoidea ** Family Protorthidae ** Family Arctohedridae ** Family Leioriidae * Superfamily Skenidioidea ** Family Skenidiidae References Rhynchonellata {{brachiopod-stub ...
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Brachiopod
Brachiopods (), phylum (biology), phylum Brachiopoda, are a phylum of animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachio ...
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Orthida
Orthida is an extinct order of brachiopods which appeared during the Early Cambrian period and became very diverse by the Ordovician, living in shallow-shelf seas. Orthids are the oldest member of the subphylum Rhynchonelliformea (Articulate Brachiopods), and is the order from which all other brachiopods of this group stem. Physically they are usually strophic, with well-developed interareas. They also commonly have radiating ribs, sulcus, and fold structures. Typically one valve, often the brachial valve, is flatter than the other. The interior structure of the brachial valves are usually simple. In shape they are sub-circular to elliptical, with typically biconvex valves. There is some debate over the forms that first appeared of this order as to how they should be classified. However, they began to differentiate themselves by the late Early Cambrian period, and by the late Cambrian period had diversified into numerous varieties and reach 2 to 5 cm in width. Spec ...
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Lower Cambrian
The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovician Period 486.85 Ma. Most of the continents lay in the southern hemisphere surrounded by the vast Panthalassa Ocean. The assembly of Gondwana during the Ediacaran and early Cambrian led to the development of new convergent plate boundaries and continental-margin arc magmatism along its margins that helped drive up global temperatures. Laurentia lay across the equator, separated from Gondwana by the opening Iapetus Ocean. The Cambrian marked a profound change in life on Earth; prior to the Period, the majority of living organisms were small, unicellular and poorly preserved. Complex, multicellular organisms gradually became more common during the Ediacaran, but it was not until the Cambrian that fossil diversity seems to rapidly incre ...
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Rhynchonellida
The taxonomic order Rhynchonellida is one of the two main groups of living articulate brachiopods, the other being the order Terebratulida. They are recognized by their strongly ribbed wedge-shaped or nut-like shells, and the very short hinge line. The hinges come to a point, a superficial resemblance to many (phylogenetically unrelated) bivalve mollusk shells. The loss of the hinge line was an important evolutionary innovation, rhynchonellids being the first truly non-strophic shells with a purely internal articulation (teeth-sockets). Strong radiating ribs are common in this group; and there are generally very strong plications or accordion-like folds on the sulcus (the long middle section) of the shell. This probably helps regulate the flow of water in and out of the shell. All rhynchonellids are biconvex (have a bulbous shell), and have a fold located in the brachial valve. This means that the commissure, the line between the two valves or shells, is zigzagged, a dist ...
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