Bentonyx Sidensis
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Bentonyx Sidensis
''Bentonyx'' (meaning "Bentons' claw") is an extinct genus of rhynchosaur from the middle Triassic epoch of Devon in England. Its fossil, a well preserved skullBRSUG 27200 was discovered in Otter Sandstone Formation (late Anisian age) and was first assigned to '' Rhynchosaurus spenceri'', that is known from 25 specimens. This species was reassigned to its own genus, ''Fodonyx'', that was first described by David W. E. Hone and Michael Benton in 2008. More recently, this skull was reassigned to this genus by Max C. Langer, Felipe C. Montefeltro, David E. Hone, Robin Whatley and Cesar L. Schultz in 2010 and the type species is ''Bentonyx sidensis''. The Cladogram below is based on work by Martin Ezcurra Martin may refer to: Places * Martin City (other) * Martin County (other) * Martin Township (other) Antarctica * Martin Peninsula, Marie Byrd Land * Port Martin, Adelie Land * Point Martin, South Orkney Islands Austra ... ''et al''. References ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic fauna Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, such as the marine reptiles (e.g. ichthyosaurs, sauropterygians, thallatosaurs), ray-finned fish and many invertebrate groups like ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Teyumbaita
''Teyumbaita'' is an extinct genus of hyperodapedontine rhynchosaur from the Upper Triassic of southern Brazil. Its fossils were recovered from the early Norian-age Caturrita Formation, one of several fossiliferous formations exposed at Paleorrota Geopark in the state of Rio Grande do Sul. ''Teyumbaita'' is likely the youngest valid genus of rhynchosaur, as other members of the group likely died out before the start of the Norian. Fossils of ''Teyumbaita'' include two nearly complete skulls and a partial skull, all of which were discovered in the lower part of the Caturrita Formation. The fossils were first named as ''Scaphonyx sulcognathus'', a species of ''Scaphonyx''. The genus ''Scaphonyx'' is now considered to be a ''nomen dubium'', so ''S. sulcognathus'' was reassigned to its own genus by Felipe Chinaglia Montefeltro, Max Cardoso Langer and Cesar Leandro Schultz in 2010. This new genus name, ''Teyumbaita'', was constructed from "lizard (''teyú'') and parrot ''(mbaitá'')" ...
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Isalorhynchus
''Isalorhynchus'' is an extinct genus of hyperodapedontine rhynchosaur from the late Triassic period (Carnian stage) of Toliara Province, southwestern Madagascar. It is known from the holotype MDE-R18, a nearly complete maxilla and from other specimens from the same locality, Malio River area. It was found in the Makay Formation (or Isalo II) of the Morondava Basin (or Isalo beds). It was first named by Eric Buffetaut in 1983 and the type species is ''Isalorhynchus genovefae''. The majority of ''Isalorhynchus'' specimens are isolated jaw bones, but two nearly complete skeletons were found in 1998. Langer ''et al.'', 2000 concluded that ''Isalorhynchus'' is a synonym of ''Hyperodapedon ''Hyperodapedon'' is a genus of rhynchosaurs (beaked, archosaur-like reptiles) from the Triassic, Late Triassic period (Carnian stage). Fossils of the genus have been found in Africa, Asia, Europe and North and South America. Its first discovery ...'' and referred it to a new species of ''Hyperod ...
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Hyperodapedontinae
Hyperodapedontinae is a subfamily of rhynchosaurs within the family Hyperodapedontidae. Fossils have been found from Argentina, Brazil, Canada, India, Madagascar, Scotland, Tanzania, United States and Zimbabwe. Phylogeny Hyperodapedontinae was erected by Sankar Chatterjee in 1969 as a coordinate name of the family Hyperodapedontidae Lydekker, 1985. Chatterjee (1969) originally named Hyperodapedontinae to include all Late Triassic rhynchosaurs known at that time, ''H. gordoni'', ''H. huxleyi'' and ''"Scaphonyx" fischeri'', and proposed a morphological diagnosis for the clade. ''Scaphonyx'' includes two additional species, ''S. africanus'' and ''S. australis'', all of which are currently believed to be dubious. As noted by Langer ''et al.'' (2000), using Chatterjee' morphological definition would exclude ''Teyumbaita'' and ''H. huenei'' from the clade, and thus it would be nested within ''Hyperodapedon''. To preserve the name, with its original stratigraphical meaning, Langer ''et a ...
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Langeronyx
''Langeronyx'' is an extinct genus of basal rhynchosaurid known from the early Middle Triassic (Anisian stage) Bromsgrove Sandstone Formation of Warwickshire, UK. It contains a single species, ''Langeronyx brodiei'', originally included in the genus ''Rhynchosaurus''. ''R. brodiei'' was first described and named by Michael Benton in 1990, but its redescription by Martín D. Ezcurra, Felipe Montefeltro and Richard J. Butler in 2016 recovered it as more closely related to the more advance hyperodapedontine than to the type species of ''Rhynchosaurus'' and thus it was moved to its own genus. The generic name ''Langeronyx'' honors the Brazilian paleontologist Max Cardoso Langer in recognition of his rhynchosaur research, combined with the Greek ''onyx'' (''óνυξ'') meaning "claw", a common suffix for rhynchosaur genera. ''L. brodiei'' is known solely from the holotype, a partial skull divided into the two specimenWARMS G6097/1and NHMUK PV R8495, housed in the Warwickshire Mu ...
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Stenaulorhynchus
''Stenaulorhynchus'' (possibly meaning "narrow tube beak") is an extinct genus of hyperodapedontid rhynchosaur known from the Middle Triassic (late Anisian stage) deposits of Tanganyika Territory, Tanzania. It was found in the Lifua Member of the Manda Formation in the Karoo Supergroup. It was named and first described by Sidney Henry Haughton in 1932. The type species is ''Stenaulorhynchus stockleyi'', a beaked herbivore measuring 1–6 meters in length. Description Dentition The teeth of ''Stenaulorhynchus'' were conical, pointed, and composed mostly of dentine, although new unworn teeth may have had a thin layer of enamel. They were set deeply into and fused with the jaw bones. They are arranged with two-to-several rows of teeth on top and only a few on bottom. The middle row of maxillary teeth only occupied the posterior third of the jaw while the other rows extended all the way forwards and sometimes down the crest of the jaw. The teeth at the front of the mouth, by th ...
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Brasinorhynchus
''Brasinorhynchus'' is an extinct genus of derived stenaulorhynchine known from the late Middle Triassic (Ladinian stage) Santa Maria Formation of Rio Grande do Sul, Brazil.''Brasinorhynchus''
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It contains a single species, ''Brasinorhynchus mariantensis''.


Description

''B. mariantensis'' is known from two specimens, including the UFRGS-PV-0168-T, a complete

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Stenaulorhynchinae
Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaurs first appeared in the Middle Triassic or possibly the Early Triassic, before becoming abundant and globally distributed during the Carnian stage of the Late Triassic. Description Rhynchosaurs were herbivores, and at times abundant (in some fossil localities accounting for 40 to 60% of specimens found), with stocky bodies and a powerful beak. Early primitive forms, like ''Mesosuchus'' and '' Howesia'', were generally small and more typically lizard-like in build, and had skulls rather similar to the early diapsid ''Youngina'', except for the beak and a few other features. Later and more advanced genera grew to medium to medium large size, up to two meters in length. The skull in these forms were short, broad, and triangular, becoming muc ...
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Hyperodapedontidae
Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaurs first appeared in the Middle Triassic or possibly the Early Triassic, before becoming abundant and globally distributed during the Carnian stage of the Late Triassic. Description Rhynchosaurs were herbivores, and at times abundant (in some fossil localities accounting for 40 to 60% of specimens found), with stocky bodies and a powerful beak. Early primitive forms, like ''Mesosuchus'' and '' Howesia'', were generally small and more typically lizard-like in build, and had skulls rather similar to the early diapsid ''Youngina'', except for the beak and a few other features. Later and more advanced genera grew to medium to medium large size, up to two meters in length. The skull in these forms were short, broad, and triangular, becoming muc ...
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Howesia
''Howesia'' is an extinct genus of basal rhynchosaur from early Middle Triassic (early Anisian stage) deposits of Eastern Cape, South Africa. It is known from the holotype SAM 5884, a partial skeleton with palate and partial lower jaws and from two paratypes, SAM 5885 and SAM 5886. It was found in the Burgersdorp Formation of the middle deposits of the Beaufort Group (Karoo Basin) and referred to Subzone B of the Cynognathus Assemblage Zone. It was first named by Robert Broom in 1905 and the type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ... is ''Howesia browni'', named after Alfred Brown. References Rhynchosaurs Middle Triassic reptiles of Africa Anisian life Fossils of South Africa Fossil taxa described in 1905 Taxa named by Robert Broom Prehistoric ...
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