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Apoidea
The superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the " sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional " Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.Manuela Sann, Oliver Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn and Michael Ohl (2018) Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. ''BMC Evolutionary Biology'' 18:71. doi:10.1186/s12862-018-1155-8 Nomenclature Bees appear in recent classifications to be a specialized lineage of " crabronid" wasps that switched to the use of pollen and nectar as larval food, rather than insect prey; this makes the traditional "Crabronidae" a paraphyletic group. Accordingly, bees and sphecoids are now all grouped t ...
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Stenotritidae
The Stenotritidae is the smallest of all formally recognised bee families, with only 21 species in two genera, all of them restricted to Australia. Historically, they were generally considered to belong in the family Colletidae, but the stenotritids are presently considered their sister taxon, and deserving of family status. Of prime importance is that the stenotritids have unmodified mouthparts, whereas colletids are separated from all other bees by having bilobed glossae. The American entomologist Ronald J. McGinley proposed their position as an independent family based on the morphology of the glossae in 1980. This view quickly became established. They are large, densely hairy, fast-flying bees, which make simple burrows in the ground and firm, ovoid provision masses in cells lined with a waterproof secretion. The nests of some species can reach a depth of more than three metres.
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Spheciformes
The Spheciformes is a paraphyletic assemblage of insect families which collectively comprise the "sphecoid wasps". Larvae are carnivorous. These are all the members of the superfamily Apoidea, which are not bees and which in older classifications were called the "Sphecoidea". Some are also described as mud dauber Mud dauber (or "mud wasp" or "dirt dauber") is a name commonly applied to a number of wasps from either the family Sphecidae or Crabronidae which build their nests from mud; this excludes members of the family Vespidae (especially the subfamily E ...s. The group is paraphyletic because the bees are believed to have arisen from a subgroup within the family Ammoplanidae, thus Spheciformes does not include all of the descendants of its common ancestor.Manuela Sann, Oliver Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn and Michael Ohl (2018) Phylogenomic analysis of Apoidea s ...
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Melittidae
Melittidae is a small bee family, with over 200 described species in three subfamilies. The family has a limited distribution, with all described species restricted to Africa and the northern temperate zone. Fossil melittids have been found occasionally in Eocene amber deposits, including those of Oise, France and the Baltic amber. Evolution Early molecular work suggested that the family Melittidae was sister to all other bees, and also that it was paraphyletic. Because of this finding, it was suggested that the three subfamilies of Melittidae should be elevated to family status. Neither study included many melittids, due to their rarity. Later studies suggested that the family could still be monophyletic and a 2013 investigation including a greater number of melittid bees further supports this. Recent research has shown that Melittids have a lower extinction rate compared to other hymenopterans, yet this family is considered species-poor. This is attributed to a significantl ...
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Halictidae
Halictidae is the second-largest family of bees (clade Anthophila) with nearly 4,500 species. Halictid species are an extremely diverse group that can vary greatly in appearance. These bees occur all over the world and are found on every continent except Antarctica. Usually dark-colored (frequently brown or black) and often metallic, halictids are found in various sizes, colors and patterns. Several species are all or partly green and a few are red, purple, or blue. A number of them have yellow markings, especially the males, which commonly have yellow faces, a pattern widespread among the various families of bees. The family is one of many with short tongues and is best distinguished by the arcuate (strongly curved) basal vein found on the wing. Females in this family tend to be larger than the males. They are commonly referred to as "sweat bees" (especially the smaller species), as they are often attracted to perspiration. Ecology Most halictids nest in the ground, often in hab ...
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Astatinae
Astatinae are a cosmopolitan group of solitary wasps, peculiar for their males having very large compound eye A compound eye is a visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidia, which are tiny independent photoreception units that consist of a cornea, lens, and photoreceptor cells which disti ...s that broadly meet at the top of the head. The largest genus in this subfamily is '' Astata'', with about half of more than 160 species in the subfamily. Phylogenomic analysis of Apoidea published in 2018 suggested that Astatinae, along with several other subfamilies and a subtribe, should be promoted to family rank: Ammoplanina (= Ammoplanidae), Astatinae (= Astatidae), Bembicinae (= Bembicidae), Mellininae (= Mellinidae), Pemphredoninae (= Pemphredonidae), Philanthinae (= Philanthidae), and Pseninae (= Psenidae). References Crabronidae Apocrita subfamilies Biological pest control wasps {{Apoidea-stub ...
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Crabronidae
The Crabronidae are a large paraphyletic group (nominally a family) of wasps, including nearly all of the species formerly comprising the now-defunct superfamily Sphecoidea. It collectively includes well over 200 genera, containing well over 9000 species. Crabronids were originally a part of Sphecidae, but the latter name is now restricted to a separate family based on what was once the subfamily Sphecinae. Several of the subfamilies of Crabronidae are often treated as families in their own right, as is true of the most recent phylogenies (example below). Phylogeny This phylogenetic tree is based on Sann ''et al.'', 2018, which used phylogenomics to demonstrate that both the bees (Anthophila) and the Sphecidae arose from within the former Crabronidae, which is therefore paraphyletic, and which they suggested should be split into several families; the former family Heterogynaidae nests within the Bembicidae, as here defined. These findings differ in several details from studie ...
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Apidae
Apidae is the largest family within the superfamily Apoidea, containing at least 5700 species of bees. The family includes some of the most commonly seen bees, including bumblebees and honey bees, but also includes stingless bees (also used for honey production), carpenter bees, orchid bees, cuckoo bees, and a number of other less widely known groups. Taxonomy In addition to its historical classification (honey bees, bumble bees, stingless bees and orchid bees), the family Apidae presently includes all the genera formerly placed in the families Anthophoridae and Ctenoplectridae. Although the most visible members of Apidae are social, the vast majority of apid bees are solitary, including a number of cleptoparasitic species. The old family Apidae contained four tribes (Apinae: Apini, Euglossini and Bombinae: Bombini, Meliponini) which have been reclassified as tribes of the subfamily Apinae, along with all of the former tribes and subfamilies of Anthophoridae and the former fam ...
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Sphecidae
The Sphecidae are a cosmopolitan family of wasps of the suborder Apocrita that includes sand wasps, mud daubers, and other thread-waisted wasps. The name Sphecidae was formerly given to a much larger grouping of wasps. This was found to be paraphyletic, so most of the old subfamilies have been moved to the Crabronidae. Biology The biology of the Sphecidae, even under the restricted definition, is still fairly diverse; some sceliphrines even display rudimentary forms of sociality, and some sphecines rear multiple larvae in a single large brood cell. Many nest in pre-existing cavities, or dig simple burrows in the soil, but some species construct free-standing nests of mud and even (in one genus) resin. All are predatory and parasitoidal, but the type of prey ranges from spiders to various dictyopterans, orthopteroids and larvae of either Lepidoptera or other Hymenoptera; the vast majority practice mass provisioning, providing all the prey items prior to laying the egg. ...
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Angarosphecidae
Angarosphecidae is an extinct family of Mesozoic and early Cenozoic wasps in the superfamily Apoidea. Genera * †'' Angarosphex'' Rasnitsyn 1975 Weald Clay, United Kingdom, Hauterivian/Barremian La Pedrera de Rúbies Formation, Spain, Barremian Crato Formation, Brazil, Aptian Laiyang Formation, Yixian Formation, China, Aptian Gurvan-Eren Formation, Mongolia, Aptian Zaza Formation, Turga Formation, Russia, Aptian Sinuiju Formation, North Korea, Aptian *†'' Baissodes'' Rasnitsyn 1975 Weald Clay, United Kingdom, Hauterivian Yixian Formaiton, China, Aptian Gurvan-Eren Formation, Mongolia, Aptian Zaza Formation, Russia, Aptian * †'' Burmasphex'' Melo and Rosa, 2018, Burmese amber, Myanmar, Cenomanian *†'' Cretobestiola'' Pulawski and Rasnitsyn 2000 La Pedrera de Rúbies Formation, Spain, Barremian Crato Formation, Brazil, Aptian, Dzun-Bain Formation, Mongolia, Aptian Zaza Formation, Argun Formation, Russia, Aptian * †'' Cretosphecium'' Pulawski and Rasnitsyn 2000 Dz ...
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Pemphredoninae
The subfamily Pemphredoninae also known as the aphid wasps, is a large group in the wasp family Crabronidae, with over 1000 species. Historically, this subfamily has frequently been accorded family status. In some recent phylogenetic analyses, one of the subtribes within this group is the sister lineage to the superfamily Apoidea, and accorded family rank as Ammoplanidae along with Pemphredonidae and Psenidae so as to keep families monophyletic.Manuela Sann, Oliver Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn, and Michael Ohl (2018) Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. ''BMC Evolutionary Biology'' 18:71. doi:10.1186/s12862-018-1155-8 The subfamily consists of solitary wasps, each genus having its own distinct and consistent prey preferences. The adult females dig tunnels in the ground, or plant material, for nesting. As with all other sphecoid ...
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Sphecidae
The Sphecidae are a cosmopolitan family of wasps of the suborder Apocrita that includes sand wasps, mud daubers, and other thread-waisted wasps. The name Sphecidae was formerly given to a much larger grouping of wasps. This was found to be paraphyletic, so most of the old subfamilies have been moved to the Crabronidae. Biology The biology of the Sphecidae, even under the restricted definition, is still fairly diverse; some sceliphrines even display rudimentary forms of sociality, and some sphecines rear multiple larvae in a single large brood cell. Many nest in pre-existing cavities, or dig simple burrows in the soil, but some species construct free-standing nests of mud and even (in one genus) resin. All are predatory and parasitoidal, but the type of prey ranges from spiders to various dictyopterans, orthopteroids and larvae of either Lepidoptera or other Hymenoptera; the vast majority practice mass provisioning, providing all the prey items prior to laying the egg. ...
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Paraphyletic
In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia ( reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor, including all extant reptiles as well as the extinct synaps ...
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